Craspedophorus phenax Basilewsky, 1987

Häckel, Martin, 2017, A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae), Zootaxa 4330 (1), pp. 1-67 : 19-20

13. Craspedophorus phenax Basilewsky, 1987

( Plate 6 , Figs 52–55, Plate 13 , Fig. 108)

Craspedophorus congoanus Burgeon 1930b: 160 (ex parte). Burgeon 1935b: 180 (ex parte).

Craspdophorus phenax Basilewsky 1987: 196 (type locality “Zaïre: Tshuapa: Isekomboka sur la Maringa [=Democratic Republic of the Congo: Équateur Province]”). Lorenz 2005: 321, Häckel and Farkač 2012: 82.

Type material. Holotype (♂): “Genit. ♂ [printed]/ 87.042.4 [handwritten in black on yellow label]// Holotypus [printed on red label]// Musée du Congo [printed]/ Isekomboka / riv Maringa / Ghesquiére VIII–27 [handwritten on white label]// Craspedophorus / phenax n. sp. [handwritten in black]/ P. Basilewsky det., 19 [printed on white label]/ [DataMatrix] RMCA Ent / 0 0 0 0 20051 [printed in black on white label]” ( Plate 6 , Fig. 52, MRAC). Paratypes. 1♂: “Paratypus [printed in black on red label]// Kamerun / Musake [(= slopes of Mt. Cameroon, Southwest Region) printed in black]/ 6. X. [handwritten in black] 10 [printed in black]/ E. Hintz [printed in black on white label] Musée du Congo [printed in black]/ Musake (Kamerun) / E. Hintz / Don [handwritten in black] L. Burgeon [printed in black on white label]// Craspedophorus / phenax n. sp. [handwritten in black]/ P. Basilesky det. 19 [printed in black on white label]” (MRAC); 1♀ same data as previous except: “Sankuru: Komi [printed in black]/ 23–IV [handwritten in black]–1930 [printed in black]/ J. Ghesquière [printed in black on white label]” ( Plate 6 , Fig. 53, MRAC); 1 Ƌ same data as previous except: “Genit. ♂ [printed in black]/ 87.042.5 [handwritten in black on yellow label]// Coll. Mus. Congo / Lomami: Mwene-Ditu / Doutrelepont [printed in black on white label]”; 1 ♀ same data as previous except: “Musée du Congo [printed in black]/ Kwango / (R.P.H. Vanderyst) [handwritten in black on white label]” (MRAC).

Additional material examined. Cameroon: Centre. 1 Ƌ: “Prov . Centre. Region um Bafia, Umg. Ndjitam, 700m, N 05°22' 5 . O 11°19'1” (cAP). Southwest. 1♀: “Prov. Sud-Quest. Region um Eyumojok, Umg. Ekok, 100m, N 05°48' 5 . O 008°54' 1.” (cAP). West. 1♀: “Prov . Nord-Ouest. Str[asse] zw[ischen] Bafoussam u[nd] Foundam, Umg. Koutaba, 1210m, N 05°09' 5 . O 010°46'4.” (cAP). 1♀: “Prov. Quest. Region um Dschang, Umg. Bafou, 1500m, N 05°28' . O 010°26'.” ( Plate 6 , Fig. 55, cMH); 1 Ƌ: “Prov. Quest. Region um Dschang, Umg. Fontem, 450m, N 05°28' 3 . O 009°47'5.” ( Plate 6 , Fig. 54, Plate 13 , Fig. 108, cMH).

Note. This species is based on seven specimens, six collected in two provinces of the DR Congo, one specimen in Cameroon Mountains (1 specimen); holotype labeled “Isekomboka sur la Maringa”. The Maringa, and the Lopori River to the north, join near Basankusu to form the Lulonga River, a tributary of the Congo River. Description (in part, see Basilewsky 1987: 196). “Length 17–19 mm. Apterous, elytra strongly fused. Black, mandibles, palps and antennae with articles from IV to XI dark brown; each elytron with two large maculae, shiny orange, with not too serrate margins, somewhat transversal, almost rounded, humeral macula in basal third, reaching from V to VIII interval, although it could exceed to IX interval, never reaching to lateral margin, consists of 5 elongated macular spots, almost equal, except spot on IX interval, which shorter; preapical macula, located in apical fourth, reaching from IV to VIII interval. Body very weakly covered by setae, more markedly in the middle of pronotum and venter. Head elongated, anteriorly and posteriorly smooth, with vertex weakly, and frontal grooves grossly punctured; labrum with anterior margin quite deeply sinuate, and 4 pores, each with one seta. Eyes strongly convex. Labial dent large and wide, rectangular, with anterior margin almost sinuate. Terminal palpar article very long and kidney-shaped. Antennae long and fine, not dilated, reaching to pronotal base by somewhat more than 4 articles. Pronotum fairly transverse, 1.20–1.27 times wider than long, hardly more narrower anteriorly than posteriorly, base somewhat wider than anterior margin, which wider than neck, anterior angles with small lobes, fairly protruded anteriorly, weakly overreaching anterior margin; lateral margins rounded, but not too dilated at midlength, where maximum of width, from where margins strongly sinuate, narrowing almost perpendicularly towards base, each with an excavation, not well marked in front of posterior angle, which well marked, but only fairly protruded; base flat, without any peduncle; disk fairly convex, markedly separated from lateral rims, which widely flattened and strongly elevated posteriorly; basilar tips very long and deeply impressed; surface very finely punctured, not rugate on disk, with small dots, well separated, although more gross and closer near margins, microsculpture very finely transverse-reticular. Elytra very elongated, 1.63–1.7 times longer than wide, very convex, almost ovoid, maximal width at apical third; humeri fairly marked, margins not too distinctly sinuate in front of apex; intervals convex, finely punctured in 2–3 rows, dots well separated, not as well separated where elytral maculae located. Microsculpture net, reticular, almost isodiametrical; striae finely and deeply impressed, somewhat more distinctly punctured. Underside regularly punctured, with dots fine, shallow, and rarely spaced, venter more rugate near margins, with three setose points in the middle. Metepisterna anteriorly somewhat shorter than wide. Legs long. This species belongs to C. strachani species group and resembles C. congoanus Kolbe, 1889 , which was partially confused with it by Burgeon (1930: 160, partim). It differs [from C. congoanus ] in its statue, narrower and more elongated, in its elytra more convex, with more acuminated apex. Pronotum also more finely punctured in C. phenax , less convex, with lateral margins more regularly rounded anteriorly, lateral angles less distinct, posteriorly more sinuate, excavation in front of posterior angle shallower; elytral intervals more convex, more weakly and rarely punctured; elytral maculae more yellow and somewhat smaller; metepisterna longer and not transverse. Aedeagi quite similar (Fig 2c, d in Basilewsky 1987: 190 [here Plate 13 , Figs 108, 109). C. congoanus occurs in wide areal of central and western Zaïre, but absents in eastern part of the country (…) C. phenax seems to occur in areal less large.” [from French].

Distribution. Central, western, southwestern Cameroon; Democratic Congo: Bandundu, Équateur, Kasaï- Oriental Provinces.

Figures

PLATE 6. Habitus of Craspedophorus strachani group V (Scale bar: 10 mm). 44. C. ruficroides ruficroides n. sp., HT (male); 45. C. ruficroides ruficroides n. sp., PT female, Guinea; 46. C. ruficroides thomsoni n. ssp., PT male, Central Africa; 47 C. ruficroides thomsoni n. ssp., PT, female, Cameroon, Centre; 48. C. ruficroides thomsoni n. ssp., HT (male); 49. C. congoanus Kolbe, 1883, female, DR Congo: Équateur (comparative type Basilewsky 1954); 50. C. congoanus Kolbe, 1883, male, DR Congo: Équateur; 51. C. congoanus Kolbe, 1883, female, DR Congo: Orientale; 52. C. phenax Basilewsky, 1987, HT (male); 53. C. phenax Basilewsky, 1987, female, DR Congo: Kasaï-Oriental; 54. C. phenax Basilewsky, 1987, male, Camerooon, West; 55. C. phenax Basilewsky, 1987, male, Camerooon, West.

PLATE 13 Craspedophorus strachani group II, aedeagus in frontal and right lateral views: 101. C. chevalieri (Alluaud, 1915), Central Africa; 102. C. cuneatus cuneatus (Alluaud, 1915), Central Africa; 103. C. cuneatus paromius Basilewsky, 1987, Orientale, DR Congo; 104. C. cuneatus rotundatus new subspecies, HT; 105. C. glaber Bates, 1886, Cameroon, Southwestern; 106. C. ruficroides ruficroides new species, HT; 107. C. ruficroides thomsoni new subspecies, PT RCA; 108. C. phenax Basilewsky, 1987, PT Fontem, Cameroon, West; 109. C. congoanus Kolbe, 1889, Eala Bokatola-Bikoro, Équateur, DR Congo; 110. C. phenacoides new species, HT; 111. C. lemariei new species, HT.