Prodromus clypeatus Distant, 1904

Yeshwanth, H. M. & Konstantinov, Fedor V., 2021, Review of the plant bug tribe Eccritotarsini (Hemiptera: Heteroptera: Miridae) of India and Sri Lanka with description of two new genera and six new species, European Journal of Taxonomy 745, pp. 1-69: 29

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Prodromus clypeatus Distant, 1904


Prodromus clypeatus Distant, 1904  

Figs 9A–B View Fig , 27C View Fig

Prodromus clypeatus Distant, 1904b: 437   .

Prodromus cuneatus Distant, 1909: 453   .

Prodromus clypeatus   – Stonedahl 1988: 70–72 (redescr.). — Yasunaga & Ishikawa 2016: 30 (biology). — Anitha & Rajamony 1991: 439 (new record for S India   ).

Prodromus cuneatus   – Stonedahl 1988: 70 (syn. with P. clypeatus   ).


Recognized by the following combination of characters: body pale yellow, usually with brown to pale brown scutellum; eyes distinctly elevated, slightly less than anterior half of eye raised above vertex in frontal view; frons and clypeus weakly convex, with weak depression in between; antennal segment II 2.2–2.4× as long as first, narrowly reddish apically, rarely entire segment with reddish tinge; apex of cuneus well surpassing apex of cell ( Fig. 9A View Fig ), membranal vein weakly and gradually curved apically; sclerotized portion of aedeagus with long, curved subapical row of spicules, apical membranous lobe without any sclerotization.

Material examined


MYANMAR • ♀; Myitta , Tenasserim Valley; 14.1667° N, 98.5167° E; 183 m a.s.l.; Doherty leg.; NHM. GoogleMaps  


MYANMAR • 1 ♀; same collection data as for lectotype; NHM GoogleMaps   .

Other material

SRI LANKA • 1 ♂, lectotype of Prodromus cuneatus   ; Peradeniya ; 7.263° N, 80.603° E; 497 m a.s.l.; May 1909; E.E. Green leg.; AMNH _ PBI 00340350 View Materials ; NHM GoogleMaps   1 specimen, sex unknown, paralectotype of Prodromus cuneatus   ; same collection data as for lectotype; AMNH _ PBI 00340353 View Materials ; NHM GoogleMaps   .

INDIA   • 14 ♂♂, 6 ♀♀; Karnataka, Yellapura, Idagundi ; 14°55.064ˊ N, 64°37.801ˊ E; 446 m a.s.l.; 6 Dec. 2011; H.M. Yeshwanth leg; Ex: banana.; UASB GoogleMaps   1 ♂, 1 ♀; same collection data as for preceding; ZISP GoogleMaps   2 ♂♂, 2 ♀♀; Uttara Kannada, Dandeli ; 13°.13ˊ N, 77°.63ˊ E; 906 m a.s.l; 10 Nov. 2012; A.R.V. Kumar leg.; UASB   3 ♂♂; Tirthahalli, Hulgar ; 13.7° N, 75.23° E; A.R.V. Kumar. Leg.; UASB GoogleMaps   5 ♂♂, 1 ♂; Sirsi ; 14°44.023ˊ N, 74°46.711ˊ E; 506 m a.s.l; 25 May 2010; H.M. Yeshwanth leg.; UASB GoogleMaps   3 ♂♂, 1 ♀; Bangalore, Hessaraghatta ; 13°8.294ˊ N, 77°28.53ˊ E; 890 m a.s.l.; 5 Jan. 2014; H.M. Yeshwanth leg; UASB GoogleMaps   1 ♂; Tamil Nadu, Coimbatore ; 11°01ˊ83˝ N, 76°97ˊ25˝ E; 463 m a.s.l; 9 Jun. 2015; H.M. Yeshwanth leg.; UASB   2 ♂♂, 1 ♀; Kerala, Peechi ; 10.032° N, 76.021° E; 82m a.s.l., 21 Sep. 2008; S. Murthy leg. UASB GoogleMaps   3 ♂♂; Meghalaya, Ri-Bhoi ; 25°41ˊ N, 91°55ˊ E; 1031 m a.s.l; 6 Jun. 2019; H.M. Yeshwanth leg.; Ex: banana; UASB GoogleMaps   .


Widely distributed in India   and now documented from Kerala ( Anitha & Rajamony 1991), Karnataka, Tamil Nadu, and Megahalaya states. Also known from Sri Lanka, Burma, Thailand, Vietnam, southern China, Taiwan, Malaysia, Java, and Ambon ( Stonedahl 1988).


This widely distributed species is best distinguished from P. subflavus   by the shape of the membranal cell and structure of the aedeagus mentioned in the key. Additional features of P. subflavus   that vary from P. clypeatus   include comparatively short antennal segment II which is about twice as long as segment I (2.2–2.4× in P. clypeatus   ), and broadly subtriangular concave posterior margin of pronotum (broadly concave, without weakly angulate midpoint in P. clypeatus   ). According to Stonedahl (1988), P. clypeatus   further differs in having weakly elevated eyes and labium reaching only middle of mesosternum. However, the eyes in all studied specimens including the holotype are distinctly elevated, with slightly less than anterior half of eye raised above vertex in frontal view, while labium is reaching or almost reaching middle coxa.


United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]


India, Bangalore, Karnataka, University of Agricultural Sciences


Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute


American Museum of Natural History


University of Agricultural Sciences














Prodromus clypeatus Distant, 1904

Yeshwanth, H. M. & Konstantinov, Fedor V. 2021

Prodromus clypeatus

Yasunaga Y. & Ishikawa T. 2016: 30
Anitha N. & Rajamony L. 1991: 439
Stonedahl G. M. 1988: 70

Prodromus cuneatus

Stonedahl G. M. 1988: 70

Prodromus cuneatus

Distant W. L. 1909: 453

Prodromus clypeatus

Distant W. L. 1904: 437