Grishin, Zhang & Cong & Shen & Song & Grishin, 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Genomics-based taxonomic rearrangement of Achlyodini and Carcharodini (Lepidoptera: Hesperiidae: Pyrginae), Insecta Mundi 2023 (16), pp. 1-33 : 21-22

Bezus Grishin , new subgenus

http://zoobank.org/ F70C2876-F6A2-4633-8938-B52917E75CF9

Type species. Pellicia bessus Möschler, 1877 .

Definition. The genus Nisoniades Hübner, 1819 (type species Papilio bromius Stoll, 1787 , a junior subjective synonym of Papilio mimas Cramer, 1775 ) splits into three prominent clades at the subgenus level ( Fig. 4 ). Because no available genus-group names have been associated with Nisoniades , two of three subgenera are new. This new subgenus keys to E.19.1c in Evans (1953) and is distinguished from its relatives by the following characters: vein RS on ventral hindwing in males does not run along vein M 1 but diverges from it at its origin, which is around the middle between the base and end of the discal cell, and is swollen until the end of cell, not much beyond, and harpe of right valva is not divided. In DNA, a combination of the following nuclear genomic base pairs is diagnostic: aly84.3.1:T55C, aly84.3.1:C101T, aly37338.21.1:A1627T, aly37338.21.1:G1861T, aly3486.1.4:T485A.

Etymology. The name is a masculine noun in the nominative singular formed from the type species name to keep it short and avoid a homonym.

Species included. Nisoniades godma Evans, 1953 , Nisoniades panama Evans, 1953 , Nisoniades suprapanama Steinhauser, 1989 , Nisoniades benda Evans, 1953 , Pellicia bessus Möschler, 1877 , Nisoniades brazia Evans, 1953 , Pellicia hecale Hayward, 1940 , Nisoniades cauca Evans, 1953 , Nisoniades remo Evans, 1953 , Pellicia guianae R. Williams and E. Bell, 1939 , Pellicia montana R. Williams and E. Bell, 1939 , Nisoniades coca Steinhauser, 1989 , and Nisoniades lata Steinhauser, 1989 .

Parent taxon. Genus Nisoniades Hübner, [1819] .

Figures

Figure 4. The nuclear genome-based phylogeny of Carcharodini, first segment (continues in Fig. 5, 6). See Fig.1 for notations.