Grishin, Zhang & Cong & Shen & Song & Grishin, 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Genomics-based taxonomic rearrangement of Achlyodini and Carcharodini (Lepidoptera: Hesperiidae: Pyrginae), Insecta Mundi 2023 (16), pp. 1-33 : 22-24

Quadralis Grishin , new subgenus

http://zoobank.org/ C859F739-AC31-4F2E-9A2A-14778E4FC67B

Type species. Pterygospidea extensa Mabille, 1891 .

Definition. Sister to the rest of the genus Gorgopas Godman and Salvin, 1894 (type species Achlyodes viridiceps Butler & H. Druce, 1872 , which is currently regarded as a junior subjective synonym of Pellicia chlorocephala Herrich-Schäffer, 1870 ) and sufficiently distinct from it ( Fig. 5 ), resembling in appearance some species of Polyctor Evans, 1953 (type species Pirgus [sic!] polyctor Prittwitz, 1868 ). Keys to E. 18.4 in Evans (1953) and distinguished from its relatives by nearly square hindwing due to a blunt tooth at the vein CuA 1 and hyaline spots in forewing cells between veins M 1 to 1A+2A. In DNA, a combination of the following nuclear genomic base pairs is diagnostic: aly770.25.10:A48C, aly451.25.1:G1816A, aly3686.9.6:G137C, aly 1656.12.3:G757C, aly5196.9.2:C832T.

Etymology. The name is a masculine noun (to agree in gender with the names of species in this subgenus) in the nominative singular, given for the square-shaped hindwing: in Latin, quadratum is square, and alis is wing.

Species included. Only the type species.

Parent taxon. Genus Gorgopas Godman and Salvin, 1894 .

Figures

Figure 5. The nuclear genome-based phylogeny of Carcharodini, middle segment (continues in 6, see Fig. 4 for the first segment) showing Pholisorina subtrib. n. Species formerly placed in Staphylus are labeled in red. Asterisk marks a species with a dedicated study (Lemes et al. 2023) that already assigned it to Bolla. See Fig. 1 for notations.