Omalium corticalis, Lacordaire, 1835

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

corticalis Lacordaire, 1835 , see: clypeata P. W. J. Müller, 1821 .

elytrata Zerche, 1998: 42 ( Metopsia ; Type locality: Isl. Can.: Tenerife: Anaga , NO Chinobre , 800 m).

- DISTRIBUTION: Canary Islands.

feloi Zerche, 1998: 44 ( Metopsia ; Type locality: Tenerife: Mte. del Agua, Los Silos).

- DISTRIBUTION: Canary Islands.

fernandezi Jarrige, 1952 , see: cimicoides Wollaston, 1864 .

fernandezi Last, 1957 , see: tenoensis H. Franz, 1986 .

flava Stephens, 1834 , see: clypeata P. W. J. Müller, 1821 .

gallica C. Koch, 1938 , see: clypeata P. W. J. Müller, 1821 .

gomerensis H. Franz, 1986: 79 ( Phloeobium ; Type locality: El Cedro auf La Gomera).

— Fauvel, 1902b: 49 ( Metopsia ; [Note: Some specimens misidentified as cimicoides according to Zerche, 1998]; Canary Islands).

— Coiffait, 1954: 162 ( Metopsia ; [Note: Misidentified as cimicoides according to Zerche, 1998]; Canary Islands).

— Zerche, 1998: 51 ( Metopsia ; characters; Canary Islands).

— Assing, 1999a: 345 ( Metopsia ; Canary Islands).

- DISTRIBUTION: Canary Islands.

hispanica Scheerpeltz, 1960 , see: clypeata P. W. J. Müller, 1821 .

neglecta Zerche, 1998: 40 ( Metopsia ; Type locality: Isl. Can.: Tenerife: Cumbre Dorsal, Esperanza-Wald, 1250 m).

- DISTRIBUTION: Canary Islands.

palmensis H. Franz, 1986: 81 ( Phloeobium ; Type locality: O-Hang der Cumbre Nueva).

— Zerche, 1998: 49 ( Metopsia ; characters; Canary Islands).

— Assing and Wunderle, 1999: 434, 437 ( Metopsia ; collecting notes; Canary Islands).

- DISTRIBUTION: Canary Islands.

retusa Stephens, 1834 , see: clypeata P. W. J. Müller, 1821 .

similis Zerche, 1998: 16 ( Metopsia ; proposed for misidentification of clypeata by various workers [Note: References specified by Zerche as misidentifications of clypeata have been moved to similis ; some others still cited as subsequent references for clypeata may refer to similis ]; Type locality: Deutschland: Brandenburg, NSG “Pimpinellenberg”

bie Oderberg. Also cited from Austria, Czech Republic, Poland, Sweden, Denmark, Romania, Bulgaria, Slovenia, Kroatia, Bosnia-Herzegowina, Macedonia, Greece,

Italy, Switzerland, France, Ukraine, Belarus, Russia, Georgia, Azerbaijan, Turkey).

— Erichson, 1839a: 646 ( Megarthrus ; [Note: Misidentified as clypeatus according to Zerche, 1998]; characters; Germany).

— Erichson, 1840: 907 ( Phloeobium ; [Note: Some specimens misidentified as clypeatum according to Zerche, 1998]; characters; Germany; France).

— Redtenbacher, 1849: 757 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; characters; Austria).

— Kraatz, 1857d: 1033 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; characters; Germany).

— Redtenbacher, 1857: 258 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; characters; Austria).

— Hochhuth, 1862: 112 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; Russia).

— Fauvel, 1871a: 51 [= 1872: 25] ( Phloeobium ; [Note: Some specimens misidentified as clypeatum according to Zerche, 1998]; characters; France; Europe; Cyprus).

— Redtenbacher, 1874: 283 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; characters; Austria).

— Seidlitz, 1891: 336 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche,

— Seidlitz, 1891a: 359 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; characters; Europe).

— Ganglbauer, 1895: 764 ( Phloeobium ; [Note: Some specimens misidentified as clypeatum according to Zerche, 1998]; characters; middle and southern Europe).

— Sainte-Claire Deville, 1906: 66 ( Metopsia ; [Note: Misidentified as clypeatum according to Zerche, 1998]; Corsica).

— Sainte-Claire Deville, 1907: 14 ( Metopsia ; [Note: Some specimens misidentified as clypeata according to Zerche, 1998]; characters; France).

— Reitter, 1909: 198 ( Metopsia ; [Note: Misidentified as clypeata according to Zerche, 1998]; characters; Germany).

— Palm, 1948: 38 ( Metopsia ; [Note: Misidentified as clypeatum according to Zerche, 1998]; characters; Sweden; Denmark).

— Scheerpeltz, 1960a: 65 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; characters; Europe).

— Horion, 1963: 19 ( Phloeobium ; [Note: Misidentified as clypeatum according to Zerche, 1998]; Denmark; Sweden; Czechoslovakia; Poland; Hungary; Romania; Bosnia Hercegovina; Serbia; Croatia; Macedonia; Greece; Russia; Italy; Switzerland; Belgium; Holland; Britain; Germany; Austria).

— Tóth, 1982: 21 ( Metopsia ; [Note: Misidentified as clypeata according to Zerche, 1998]; characters; Hungary).

— Burakowski, Mroczkowski, and Stefańska, 2000: 39 ( Metopsia ; catalog; Poland).

— Assing, 2001: 75 ( Metopsia ; collected from xerothermous site; Germany).

- DISTRIBUTION: Europe, Russia, Georgia, Azerbaijan, Turkey.

tenoensis H. Franz, 1986: 80 ( Phloeobium ; Type locality: westlicher Teil des Mte. de Erjos unweit von El Palmar).

— Hernández, Outerelo, and Gamarra, 1994: 193 ( Metopsia synonym of fernandezi Last ).

— Zerche, 1998: 45 ( Metopsia ; characters; Canary Islands).

- DISTRIBUTION: Canary Islands.

fernandezi Last, 1957: 165 ( Metopsia ; [preoccupied]; Type locality: Canary Islands: Tenerife ).

— Hernández, Outerelo, and Gamarra, 1994: 193 ( Metopsia ; notes; Canary Island).

— Zerche, 1998: 45 ( Metopsia ; synonym of tenoensis ).

Proteinus

[36 species; Neotropical, Nearctic, Palaearctic, Ethiopian , and Oriental Regions] Proteinus Latreille, 1797: 9 (without included species). TYPE SPECIES: Dermestes brachypterus Fabricius , fixed by action of the Commission, Opinion 876 (ICZN, 1969).

— Latreille, 1806: 298 (characters).

— Latreille, 1810: 183, 427 (characters).

— Leach, 1815: 92 (characters).

— Samouelle, 1819: 175 (characters).

— Billberg, 1820: 15 (cited as Protinus ; characters; list of some species).

— Latreille, 1825: 245 (cited as Proteine; characters).

— Latreille, 1825a: 215 (characters).

— Berthold, 1827: 332 (characters).

— Curtis, 1829: 28, 245 (cited as Proteinnus in column 28 and corrected on column 245; catalog; British species).

— Stephens, 1829a: 24 (catalog; Britain).

— Stephens, 1829: 295 (catalog; Britain).

— Latreille, 1829: 440 (characters).

— Mannerheim, 1830: 10, 57 (key; characters).

— Mannerheim, 1831: 424, 471 (key; characters).

— Dejean, 1833: 69 (list of species).

— Gistel, 1834: 9 (list of species).

— Lacordaire, 1835: 490 (characters).

— Westwood, 1838a: 18 ( Proteinus ; type species).

— Erichson, 1839a: 642 (characters).

— Heer, 1839: 170 (characters).

— Stephens, 1839: 422 (characters).

— Erichson, 1840: 902 (characters).

— Laporte, 1840: 193 (characters).

— Redtenbacher, 1849: 755 (characters).

— Schaum, 1852: 31 (list of species; Europe).

— Lacordaire, 1854: 146 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 653 (characters).

— Gistel, 1856: 390 (cited as Protinus ; list of species).

— Jacquelin du Val, 1857: 78 (characters).

— Redtenbacher, 1857: 256 (characters; key to Austrian species).

— Kraatz, 1857d: 1022 (characters).

— Thomson, 1858: 38 (characters).

— G. Waterhouse, 1858: 32 (catalog of British species).

— Thomson, 1859: 52 (characters; type species: brachypterus ).

— Schaum, 1859: 32 (catalog; European species).

— LeConte, 1861a: 71 (characters).

— Thomson, 1861: 217 (characters).

— Gredler, 1863: 127 (list of species from Tirol).

— LeConte, 1863: 26 (list of North American species).

— Wencker and Silbermann, 1866: 37 (list of species; collecting notes; France).

— Fauvel, 1867a: 49 [= 1868: 54] (characters).

— Jacquelin du Val, 1868: 82 (catalog; European species).

— Fauvel, 1871a: 55 [= 1872: 29] (cited as Protinus ; characters; key to species).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 281 (characters; key to Austrian species).

— Fauvel, 1878d: 194 [= 1878e: 30] (key to North American species).

— Mulsant and Rey, 1878b: 218 (characters; key to French species).

— Heyden, 1880: 84 (cited as Protinus ; list of species of Siberian region).

— Duvivier, 1883: 201 (catalog; cited as Protinus ).

— Fowler, 1888: 427 (notes; key to British species).

— Seidlitz, 1891: 337 (key to some European species).

— Seidlitz, 1891a: 359 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 122 (cited as Protinus ; list of species of Europe and Caucasus).

— Heyden, 1893: 49 (cited as Protinus ; list of species of Siberian region).

— Ganglbauer, 1895: 757 (characters; key to middle European species).

— Heyden, 1896: 37 (cited as Protinus ; list of species of Siberian region).

— Fauvel, 1897d: 243 (catalog of species of Barbary).

— Heyden, Reitter, and Weise, 1906: 127 (list of species of Europe and Caucasus).

— Sainte-Claire Deville, 1907: 17 (key to species of Seine Basin, France).

— Reitter, 1909: 198 (characters; key to German species).

— Bernhauer and Schubert, 1910: 33 (world catalog; 14 species).

— Petri, 1912: 51 (list of species of Siebenbürgen).

— Johansen, 1914: 626 (characters; key to Danish species).

— Leng, 1920: 93 (catalog of North American species).

— Dodero, 1923: 45 (key to Italian species).

— Winkler, 1925: 325 (catalog for Palaearctic region).

— Cameron, 1925: 103 (catalog of Indian species).

— Porta, 1926: 9 (characters; key to species of Italy).

— Portevin, 1929: 457 (characters; key to French genera).

— Sjöberg, 1930: 188 (key to species).

— Roubal, 1930: 302 (catalog; Slovakia).

— Joy, 1932: 94 (key to British species).

— Scheerpeltz, 1933: 1026 (world catalog supplement).

— Normand, 1935: 356 (list of species of Tunisia).

— Scheerpeltz, 1937b: 186 (list of Bulgarian species).

— Blackwelder, 1944: 101 (checklist of species from Mexico, Central and South America , and the West Indies).

— Kloet and Hincks, 1945: 160 (list of British species).

— Scheerpeltz and Höfler, 1948: 151 (key to european species).

— Palm, 1948: 45 (characters; key to species of Sweden).

— Tottenham, 1949: 353, 405 (type species: brachypterus Fabricius ; notes; list of British species).

— Blackwelder, 1952: 413 (type species: pulicarius ).

— Tottenham, 1954: 14 (characters; key to British species).

— Hatch, 1957: 106 (characters; key to species of Pacific Northwest).

— Kocher, 1958: 92 (checklist of species; Morocco).

— Ferreira, 1962a: 9 (list of Portuguese species).

— Horion, 1963: 27 (list of central European species).

— Székessy, 1963: 11 (characters).

— Lohse, 1964: 24 (characters; key to central European species).

— Steel, 1966: 288 (larval characters).

— Kasule, 1966: 279 (larval characters).

— Jelinek and Steel, 1966 (application to Commission to designate as Dermestes brachypterus Fabricius as type species of genus and to place name on Official List of Generic Names in Zoology).

— Pototskaia, 1967: 19 (larval characters).

— Coiffait and Saiz, 1968: 456 (characters; Chilean species).

— Scheerpeltz, 1968a: 4 (catalog of Austrian species).

— ICZN, 1969 ( Proteinus placed on Official List of Generic Names in Zoology; type species designation) .

— Bordoni, 1973j: 655 (list of species from islands near Sicily).

— Tikhomirova, 1973a: 136 (checklist of species of USSR).

— Moore and Legner, 1974: 14 (characters; notes).

— Moore and Legner, 1975: 176 (catalog; North America north of Mexico).

— Shibata, 1976: 190 (checklist of species of Japan).

— Moore and Legner, 1976: 538 (includes seashore species; notes).

— Pope, 1977: 22 (list of British species).

— Topp, 1978: 307 (larval characters).

— Frank, 1979 (notes on habitat; generic nomenclature).

— Burakowski, Mroczkowski, and Stefańska, 1979: 13 (catalog; Poland).

— Moore and Legner, 1979: 158 (characters; notes).

— Muona, 1979: 17 (list of Scandinavian species).

— Uhlig, 1979: 242 (some species collected in Germany).

— Uhlig, Vogel, and Sieber, 1980: 240 (some species collected in Germany).

— Uhlig and Vogler, 1981: 83 (list of some species collected in Germany; habitat and collecting notes).

— Tóth, 1982: 23 (characters; habitus; key to species of Hungary).

— Frank and Thomas, 1984: 98 (notes on habitats; affinities of peckorum ).

— Ádám, 1985: 251 (list of species from southeast Hungary).

— Frank, 1986a: 365 (checklist of species of Florida).

— Muona and Viramo, 1986: 14 (list of species of northeastern Finland).

— Segers, 1986: 17 (checklist of species; Belgium).

— Hayashi, 1988: 22 (key to species of Japan).

— M. Hansen, Kristensen, Mahler, Pedersen, 1991: 106 (list of Danish species).

— Angelini, 1991: 193 (list of species from high elevations of Calabria, Italy). — Campbell and Davies, 1991: 87 (checklist of species; Alaska and Canada). — Silfverberg, 1992: 21 (list of species of Fennoscandia, Denmark, and the Baltic States). — Siitonen, 1993: 230 (list of Finnish species). — Welch, 1993: 227 (ovariole number and ovary structure). — Boháč, 1993: 40 (list of species of Czech Republic and Slovakia). — Newton and Thayer, 1995: 299 ( Proteinini ). — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species). — Zanetti, 1995: 5 (list of species of Italy). — Dauphin, 1995a: 101 (characters; habitat; notes; key to species of western Europe). — M. Hansen, 1996: 96 (list of Danish species). — M. Hansen, Mahler, Palm, and Pedersen, 1996: 240 (additions to list of Danish species). — Downie and Arnett, 1996: 429 (characters; key to species of Northeastern North America ). — Newton, Thayer, Ashe, and Chandler, 2000: 343 (10 Nearctic species; notes; characters in key).

Macropterum Gistel, 1834: 9 (species included: macropterum ). TYPE SPECIES: Omalium macropterum Gravenhorst , fixed by monotypy. — Gistel, 1856: 267, 390 (list of species). — Blackwelder, 1952: 229 (synonym of Megarthrus ; type species: macropterum ).

Protinus Gemminger and Harold, 1868: 671 (emendation of Proteinus ). TYPE SPECIES: Dermestes brachypterus Fabricius , fixed by objective synonymy with Proteinus .

Pteronius Blackwelder, 1952: 331 (Note: Proposed for the species of Staphylinidae included in Proteinus which, before action of the Commission, had a type species that would have placed it in the Nitidulidae ). TYPE SPECIES: Dermestes brachypterus Fabricius , fixed by original designation. — Muona, 1979: 17 (synonym of Proteinus ). — Ádám, 1987: 137 (list of three species with collecting notes; Hungary). — Boháč, 1993: 40 (synonym of Proteinus ). — Ádám, 1996: 237 (list of species collected in Bükk National Park, Hungary).

altaicus Reitter, 1905 a: 227 ( Proteinus ; Type locality: Altai, Ostsibirien; [Note: See lectotype designation by Muona, 1978]). — Poppius, 1909: 3 ( Proteinus ; Lena Valley, Siberia ). — Bernhauer and Schubert, 1910: 33 ( Proteinus ; catalog). — Scheerpeltz, 1933: 1026 ( Proteinus ; catalog). — Muona, 1978: 125 ( Proteinus ; lectotype designation; synonymic notes). - DISTRIBUTION: Finland, Russia.

apicidens Jansson and Sjöberg, 1929: 212 ( Proteinus ; Type locality: Hamra Naturschutzgebiet in Dalarne; Abisko im nördlichen Lappland). — Sjöberg, 1930: 181 ( Proteinus ; characters; notes). — Scheerpeltz, 1933: 1026 ( Proteinus ; catalog). — Palm, 1948: 48 ( Proteinus ; characters; Finland; Sweden; Norway; [Note: The records from Norway and Sweden many refer to hyperboreus ]). — Lohse, 1964: 24 ( Proteinus ; characters; Scandinavia). — Muona, 1978: 125 ( Proteinus ; synonym of altaicus ). — Lohse and Lucht, 1989: 122 ( Proteinus ; synonym of altaicus ).

angelinii Dauphin, 1999: 135 ( Proteinus ; Type locality: Basilicata, Accettura , bosco Montepiano). - DISTRIBUTION: Italy.

apicidens Jansson and Sjöberg, 1929 , see: altaicus Reitter, 1905 .

atomarius Erichson, 1840: 904 ( Proteinus ; Type locality: Germany). — Redtenbacher, 1849: 756 ( Proteinus ; characters; Austria). — Fairmaire and Laboulbène, 1856: 654 ( Proteinus ; characters; France). — Redtenbacher, 1857: 257 ( Proteinus ; characters; Austria). — Thomson, 1861: 218 ( Proteinus ; characters; Scandinavia).

— Pandellé, 1867: 169 ( Proteinus ; characters; France).

— Gemminger and Harold, 1868: 672 ( Proteinus ; synonym of clavicornis ).

— Fauvel, 1871a: 57 [= 1872: 31] ( Proteinus ; [Note: Bernhauer and Schubert, 1910: 33 cited this use by Fauvel of clavicornis as a synonym of atomarius ; Fauvel did not describe the species as new, he attributed it to Stephens. Furthermore, Fauvel cited atomarius as a junior synonym of clavicornis ]; synonym of clavicornis ).

— Fauvel, 1873b: 110 [= 1873c: 3] ( Protinus ; Austria; Sicily; Sardinia; Russia).

— Redtenbacher, 1874: 282 ( Proteinus ; characters; Austria).

— Seidlitz, 1875: 232 ( Proteinus ; characters; Baltic region).

— Fauvel, 1876a: 47 [= 1876: 232] ( Protinus ; France; Corsica; Malta).

— J. Sahlberg, 1876: 224 ( Proteinus ; synonym of clavicornis ).

— Fauvel, 1878: 87 [= 1878a: 7] ( Proteinus , cited as Protinus ; Algeria).

— Fauvel, 1878d: 196 [= 1878e: 32] ( Proteinus ; characters; Lake Superior; Michigan).

— Mulsant and Rey, 1878b: 230 ( Proteinus ; characters; France).

— Fauvel, 1886: 13 [= 1886a: 4] ( Proteinus , cited as Protinus ; Algeria; Morocco).

— Fowler, 1888: 428 ( Proteinus ; characters; Britain).

— Ganglbauer, 1895: 760 ( Proteinus ; characters; Europe; Mediterranean region; North America ).

— Fauvel, 1902b: 50 ( Proteinus ; Morocco; Algeria; Tunisia).

— Reitter, 1905a: 228 ( Proteinus ; characters; Europe; Mediterranean region; Caucasus; North America ).

— Sainte-Claire Deville, 1906: 67 ( Proteinus , cited as Protinus ; Corsica).

— Reitter, 1909: 199 ( Proteinus ; characters; Germany).

— Bernhauer and Schubert, 1910: 33 ( Proteinus ; catalog).

— Johansen, 1914: 628 ( Proteinus ; characters; Denmark).

— Portevin, 1929: 458 ( Proteinus ; characters; France).

— Sjöberg, 1930: 184 ( Proteinus ; characters; notes).

— Scheerpeltz, 1933: 1026 ( Proteinus ; catalog).

— Scheerpeltz and Höfler, 1948: 150 ( Proteinus ; fungus inhabitant).

— Palm, 1948: 52 ( Proteinus ; characters; Sweden; Denmark; Norway; Finland).

— Tottenham, 1949: 405 ( Proteinus ; checklist; Britain).

— Hatch, 1957: 106 ( Proteinus ; characters; British Columbia ).

— Kocher, 1958: 92 ( Proteinus ; Morocco).

— Ferreira, 1962: 9 ( Proteinus ; Portugal).

— Horion, 1963: 31 ( Proteinus ; Britain; Scotland; Norway; Sweden; Finland; Denmark; Portugal; Spain; Balearics; France; Corsica; Italy; Sicily; Dalmatia; Hercegovina; Greece; Germany; Austria).

— Lohse, 1964: 25 ( Proteinus ; characters; central Europe).

— Smetana, 1964d: 51 ( Proteinus ; Czechoslovakia).

— Scheerpeltz, 1968a: 5 ( Proteinus ; catalog; Austria).

— Smetana, 1970: 54 ( Proteinus ; distributional summary; Azores).

— Korge, 1971a: 5 ( Proteinus ; Turkey).

— Smetana 1975c: 154 ( Proteinus ; Mongolia).

— Coiffait, 1976b: 83 ( Proteinus ; Greece; Corfu; Rhodes).

— Pope, 1977: 22 ( Proteinus ; Britain).

— Frank, 1979: 330, 336 ( Proteinus ; notes on nomenclature and North American records; species may not be in North America ).

— Muona, 1979: 17 ( Proteinus ; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

— Burakowski, Mroczkowski, and Stefańska, 1979: 13 ( Proteinus ; catalog; Poland).

— Hammond, 1980: 135 ( Proteinus ; uncomfirmed in Ireland).

— Tóth, 1982: 28 ( Proteinus ; characters; Hungary).

— Frank and Thomas, 1984: 99, 100 ( Proteinus ; characters; Illinois).

— Muona and Viramo, 1986: 15 ( Proteinus ; Finland).

— Lucht, 1987: 85 ( Proteinus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Dettner and Reissenweber, 1991 ( Proteinus ; defensive secretions: chemistry and systematics).

— Angelini, 1991: 193 ( Proteinus ; Italy).

— Campbell and Davies, 1991: 87 ( Proteinus ; checklist; USA; Canada).

— Silfverberg, 1992: 21 ( Proteinus ; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

— Boháč, 1993: 40 ( Proteinus ; Czech Republic; Slovakia).

— Sparacio, 1995: 138 ( Proteinus ; characters; summary of distribution; notes; Sicily).

— Zanetti, 1995: 5 ( Proteinus ; Italy).

— Dauphin, 1995a: 108, 118 ( Proteinus ; characters; France).

— Downie and Arnett, 1996: 429 ( Proteinus ; characters; USA).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 240 ( Proteinus ; Denmark).

— M. Hansen, 1996: 96 ( Proteinus ; checklist; Denmark).

— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 42 ( Proteinus ; Latvia).

— Anderson, Nash, and O’Connor, 1997: 16 ( Proteinus ; Ireland).

— Ryabukhin, 1999: 11 ( Proteinus ; catalog; habitat; distribution; northeastern Russia).

— Owen, 2000: 250 ( Proteinus ; pitfall trap collections from underground at roots of trees; Britain).

— Herman, 2001: 24 ( Proteinus ; nomen protectum).

- DISTRIBUTION: Tunisia, Algeria, Morocco, Azores, Europe, Russia, Mongolia, Canada (?), USA (?).

clavicornis Stephens, 1834: 334 ( Proteinus ; [Note: This name is older than atomarius ]; Type locality: London; Suffolk; Oakhampton, Somerset).

— Stephens, 1839: 424 ( Proteinus ; characters; England).

— G. Waterhouse, 1858: 32 ( Proteinus ; synonym of atomarius ; [Note: Waterhouse cited the “ clavicornis , Kirby’s coll.” as a synonym of atomarius ; Stephens’ description of clavicornis cited the species as “ Pr. clavicornis . Kirby MSS”. I assume that Waterhouse’s use of clavicornis also referred to that of Stephens]).

— Gemminger and Harold, 1868: 672 ( Proteinus ; valid species).

— Fauvel, 1871a: 57 [= 1872: 31] ( Proteinus ; [Note: Fauvel cited this name as the senior synonym of atomarius ]; characters; France; Scandinavia; Britain; Germany; Algeria; North America ).

— J. Sahlberg, 1876: 224 ( Proteinus ; valid species; Finland).

— Mulsant and Rey, 1878b: 231 ( Proteinus ; Fauvel cited as author; synonym of atomarius ).

— Ganglbauer, 1895: 760 ( Proteinus ; Fauvel cited as author; synonym of atomarius ).

— Bernhauer and Schubert, 1910: 33 ( Proteinus ; cited with Fauvel as author; synonym of atomarius ).

— Herman, 2001: 24 ( Proteinus ; nomen oblitum).

olivierii Saulcy, 1866: 51 ( Proteinus ; [Note: The original spelling of this name was oliverii but the name is a patronym for the collector, M. Olivier so the correct spelling is olivierii as was used by Ganglbauer and others]; Type locality: Bône).

— Fauvel, 1871a: 57 [= 1872: 31] ( Proteinus ; synonym of clavicornis ).

— Ganglbauer, 1895: 760 ( Proteinus ; synonym of atomarius ).

— Fauvel, 1902b: 50 ( Proteinus ; synonym of atomarius ).

— Reitter, 1905a: 228 ( Proteinus ; synonym of atomarius ).

— Bernhauer and Schubert, 1910: 33 ( Proteinus ; synonym of atomarius ).

oblongus Petri, 1891: 22 ( Proteinus ; variety of atomarius ; Type locality: Umgebung Schässburg).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of limbatus ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of limbatus ).

— Scheerpeltz, 1933: 1027 ( Proteinus ; synonym of crenulatus ).

— Tóth, 1982: 28 ( Proteinus ; characters; variety of atomarius ).

basalis Mäklin, 1852: 324 ( Proteinus ; Type locality: insula Sitkha).

— Fauvel, 1878d: 195 [= 1878e: 31] ( Proteinus ; characters; Sitka Island).

— Bernhauer and Schubert, 1910: 33 ( Proteinus ; catalog).

— Scheerpeltz, 1933: 1026 ( Proteinus ; catalog).

— Hatch, 1957: 107 ( Proteinus ; characters; British Columbia ; Washington; Oregon; Idaho). — Campbell and Davies, 1991: 87 ( Proteinus ; checklist; USA; Canada).

- DISTRIBUTION: Canada, USA.

brachypterus Fabricius, 1792: 235 ( Dermestes ; Type locality: Germania).

— Panzer, 1795: 103 ( Dermestes ; characters).

— Panzer, 1796: 10 ( Dermestes ; characters; illustration).

— Gravenhorst, 1806: 215 ( Omalium ; synonym of ovatus Gravenhorst ).

— Latreille, 1806: 298 ( Proteinus ; characters; France).

— Gyllenhal, 1810: 207 ( Omalium ; characters).

— Olivier, 1811: 479 ( Omalium ; synonym of ovatus ).

— Leach, 1815: 92 ( Proteinus ; characters; France; England).

— Samouelle, 1819: 175 ( Proteinus ; characters; England; France).

— Latreille, 1825a: 215 ( Proteinus ; characters; France).

— Gyllenhal, 1827: 464 ( Omalium ).

— C. Sahlberg, 1827: 279 ( Omalium ; characters; Finland).

— Curtis, 1829: 28 ( Proteinus ; catalog; Britain).

— Mannerheim, 1830: 57 ( Proteinus ; Finland; Sweden).

— Mannerheim, 1831: 471 ( Proteinus ; Finland; Sweden).

— Stephens, 1834: 334 ( Proteinus ; characters; habitat).

— Runde, 1835: 24 ( Proteinus ; characters; Germany).

— Lacordaire, 1835: 491 ( Proteinus ; characters; France).

— Erichson, 1839a: 642 ( Proteinus ; characters; Germany).

— Heer, 1839: 170 ( Proteinus ; characters; Switzerland).

— Stephens, 1839: 424 ( Proteinus ; characters; England).

— Erichson, 1840: 903 ( Proteinus ; characters; Europe).

— Laporte, 1840: 193 ( Proteinus ; characters; France).

— Hochhuth, 1849: 205 ( Proteinus ; characters; Caucasus).

— Redtenbacher, 1849: 756 ( Proteinus ; characters; Austria).

— Hardy, 1851: 63 ( Proteinus ; notes; England).

— Fairmaire and Laboulbène, 1856: 653 ( Proteinus ; characters; France).

— Gistel, 1856: 267 ( Arpedium ).

— Redtenbacher, 1857: 257 ( Proteinus ; characters; Austria).

— Kraatz, 1857d: 1024 ( Proteinus ; characters; Germany).

— Thomson, 1859: 52 (type species of Proteinus ).

— Thomson, 1861: 217 ( Proteinus ; characters; Scandinavia).

— Pandellé, 1867: 169 ( Proteinus ; characters; France).

— Fauvel, 1871a: 57 [= 1872: 31] ( Proteinus ; characters; France; Europe; Algeria).

— Fauvel, 1873b: 110 [= 1873c: 3] ( Protinus ; Caucasus).

— Redtenbacher, 1874: 282 ( Proteinus ; characters; Austria).

— Seidlitz, 1875: 232 ( Proteinus ; characters; Baltic region).

— J. Sahlberg, 1876: 224 ( Proteinus ; Finland).

— Fauvel, 1878: 87 [= 1878a: 7] ( Protinus ; Algeria).

— Mulsant and Rey, 1878b: 223 ( Proteinus ; characters; France).

— Fauvel, 1886: 13 [= 1886a: 5] ( Protinus ; Algeria; Morocco).

— Fowler, 1888: 428 ( Proteinus ; characters; Britain).

— Ganglbauer, 1895: 759 ( Proteinus ; characters; Europe; Mediterranean region).

— Fauvel, 1902b: 50 ( Proteinus ; Morocco; Algeria; Tunisia).

— Reitter, 1905a: 228 ( Proteinus ; characters; Europe; Mediterranean region; Caucasus).

— Sainte-Claire Deville, 1906: 67 ( Protinus ; Corsica).

— Poppius, 1909: 3 ( Proteinus ; Lena Valley, Siberia ).

— Reitter, 1909: 199 ( Proteinus ; characters; Germany).

— Bernhauer and Schubert, 1910: 33 ( Proteinus ; catalog).

— Johansen, 1914: 627 ( Proteinus ; characters; Denmark).

— Eichelbaum, 1914: 25 ( Proteinus ; abdominal segmentation; apical segements).

— Gridelli, 1930: 61 ( Proteinus ; Libya).

— Sjöberg, 1930: 177 ( Proteinus ; characters; notes; Sweden; Finland; Russia; Germany; Transylvania; England).

— Scheerpeltz, 1933: 1026 ( Proteinus ; catalog).

— Paulian, 1941: 143 ( Proteinus ; larval characters; Gold Coast).

— Scheerpeltz and Höfler, 1948: 150 ( Proteinus ; fungus inhabitant).

— Palm, 1948: 48 ( Proteinus ; characters; Sweden; Denmark; Norway; Finland).

— Jeannel and Jarrige, 1949: 323 ( Proteinus ; cave species; Romania).

— Hatch, 1957: 106 ( Proteinus ; characters; no specimens examined from Pacific Northwest; British Columbia ).

— Scheerpeltz, 1963a: 414 ( Proteinus ; Greece).

— Horion, 1963: 30 ( Proteinus ; Lapland to Mediterranean region to eastern Siberia; Germany; Austria).

— Lohse, 1964: 24 ( Proteinus ; characters; central Europe).

— Smetana, 1964d: 51 ( Proteinus ; Czechoslovakia).

— Pototskaia, 1967: 19 ( Proteinus ; larval characters).

— Szujecki, 1968a: 710 ( Proteinus ; Poland).

— Pope, 1977: 22 ( Proteinus ; Britain).

— Jelinek and Steel, 1966 (application to Commission to designate brachypterus as type species of Proteinus and to place species on Official List of Specific Names in Zoology).

— ICZN, 1969 (name placed on Official List of Specific Names in Zoology; type species of Proteinus ) .

— Frank, 1979: 330 ( Proteinus ; notes on distribution in Canada [?] and USA [?]).

— Hammond, 1980: 135 ( Proteinus ; Ireland).

— Tóth, 1982: 24 ( Proteinus ; characters; Hungary).

— Frank and Thomas, 1984: 100 ( Proteinus ; characters).

— Lucht, 1987: 85 ( Proteinus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Dettner and Reissenweber, 1991 ( Proteinus ; defensive secretions: chemistry and systematics).

— Campbell and Davies, 1991: 87 ( Proteinus ; checklist; USA; Canada).

— Welch, 1993: 227 ( Proteinus ; ovariole number and ovary structure).

— Boháč, 1993: 40 ( Proteinus ; Czech Republic; Slovakia).

— Cuccodoro, 1995: 262 ( Proteinus ; water loading behavior).

— Øland, 1995: 23 ( Proteinus ; host mushroom; Norway).

— Muona and Viramo, 1995: 9 ( Proteinus ; collected from fungus and dung of Ursus arctos ; Finland).

— Sparacio, 1995: 138 ( Proteinus ; characters; notes; Sicily).

— Zanetti, 1995: 5 ( Proteinus ; Italy).

— Dauphin, 1995a: 106, 112, 118 ( Proteinus ; characters; France).

— Cuccodoro, 1996a: 108 ( Proteinus ; water loading behavior).

— Owen, 1999b: 61 ( Proteinus ; captured underground at roots of old trees; Britain).

— Ryabukhin, 1999: 12 ( Proteinus ; catalog; habitat; distribution; northeastern Russia).

— Kashcheev, 1999a: 57 ( Proteinus ; Kazakhstan).

— Owen, 2000: 250 ( Proteinus ; pitfall trap collections from underground at roots of trees; Britain).

— Assing, 2001: 75 ( Proteinus ; collected from xerothermous site; Germany).

- DISTRIBUTION: Europe, Russia, Libya, Tunisia, Algeria, Morocco, Ghana, Canada (?), USA (?).

ovatus Gravenhorst, 1806: 215 ( Omalium ; Type locality: Not cited).

— Gyllenhal, 1810: 207 ( Omalium ; synonym of brachypterum Fabricius ).

— Olivier, 1811: 479 ( Omalium ; characters; France; Germany; Sweden).

— Latreille, 1829: 440 ( Proteinus ; characters).

— Lacordaire, 1835: 491 ( Proteinus ; synonym of of brachypterus Fabricius ).

— Erichson, 1839a: 643 ( Proteinus ; synonym of brachypterus ).

— Erichson, 1840: 903 ( Proteinus ; synonym of brachypterus ).

— Kraatz, 1857d: 1025 ( Proteinus ; synonym of brachypterus ).

— Mulsant and Rey, 1878b: 223 ( Proteinus ; synonym of brachypterus ).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of brachypterus ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of brachypterus ).

nigricornis Stephens, 1834: 336 ( Anthobium ; Type locality: London ; Devonshire).

— Stephens, 1839: 424 ( Anthobium ; characters; England).

— G. Waterhouse, 1858: 32 ( Proteinus ; synonym of brachypterus ).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of brachypterus ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of brachypterus ).

nitidus Stephens, 1834: 337 ( Anthobium ; Type locality: London).

— Stephens, 1839: 425 ( Anthobium ; characters; England).

— G. Waterhouse, 1858: 32 ( Proteinus ; synonym of brachypterus ).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of brachypterus ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of brachypterus ).

laevicollis Heer, 1839: 180 ( Omalium ; Type locality: Zürich).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of brachypterus ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of brachypterus ).

lividipennis Reitter, 1905 a: 228 ( Proteinus ; variety of brachypterus ; Type locality: Kaukasus, Lenkoran).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; variety of brachypterus ).

brevicollis Erichson, 1840 , see: ovalis Stephens, 1834 .

clavicornis Stephens, 1834 , see: atomarius Erichson, 1840 .

collaris Hatch, 1957: 107 ( Proteinus ; Type locality: Washington: Seattle ).

— Frank, 1979: 336 ( Proteinus ; characters).

— Frank and Thomas, 1984: 100 ( Proteinus ; characters).

- DISTRIBUTION: USA.

crassicornis Sharp, 1874: 100 ( Proteinus ; Type locality: Nagasaki).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

— Scheerpeltz, 1933: 1027 ( Proteinus ; catalog).

— Nakane, 1963a: 82 ( Proteinus ; characters; habitus photograph; Japan).

— Yuh, Paik, Kwon, and Lee, 1985: 225 ( Proteinus ; Korea).

— Hayashi, 1986: 107 ( Proteinus ; characters; Japan).

— Hayashi, 1988: 22 ( Proteinus ; anatomy of mouthparts; key).

— J. Li, 1993: 52 ( Proteinus ; characters; China).

— Watanabe, 1996a: 8 ( Proteinus ; cave collection; Japan).

- DISTRIBUTION: Japan, Korea, China.

crenulatus Pandellé, 1867: 169 ( Proteinus ; Type locality: H.-Pyrénées, Louron).

— Fauvel, 1871a: 56 [= 1872: 30] ( Proteinus ; synonym of limbatus ).

— Fauvel, 1871a: 56 [= 1872: 30] ( Protinus ; cited as limbatus ; [Note: Misidentification: see Bernhauer, 1912i: 678; Horion, 1963: 28; Frank, 1979: 331]; characters; France; Italy; Corsica; Alaska [Note: This record refers to crenulatus ]).

— Mulsant and Rey, 1878b: 226 ( Proteinus ; synonym of limbatus ).

— Mulsant and Rey, 1878b: 225 ( Proteinus ; cited as limbatus ; [Note: Misidentification: see Bernhauer, 1912i: 678; Horion, 1963: 28; Frank, 1979: 331]; characters; France).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of limbatus ).

— Ganglbauer, 1895: 759 ( Proteinus ; cited as limbatus ; [Note: Misidentification: see Bernhauer, 1912i: 678; Horion, 1963: 28; Frank, 1979: 331]; characters; France; Piemont; Corsica; Siebenbürgen; Caucasus; North America ).

— Reitter, 1905a: 227 ( Proteinus ; cited as limbatus ; [Note: Misidentification: see Bernhauer, 1912i: 678; Horion, 1963: 28; Frank, 1979: 331]; characters; Italy; France; Corsica; Siebenbürgen; North America [Note: This record refers to limbatus ]).

— Sainte-Claire Deville, 1906: 67 ( Protinus ; cited as limbatus ; [Note: Misidentification: see Bern-

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of limbatus ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; cited as limbatus ; [Note: Misidentification:

see Bernhauer, 1912i: 678; Horion, 1963: 28; Frank, 1979: 331]; catalog).

— Portevin, 1929: 457 ( Proteinus ; cited as limbatus ; [Note: Misidentification: see Bernhauer, 1912i: 678; Horion, 1963: 28; Frank, 1979: 331]; characters; France).

— Sjöberg, 1930: 178 ( Proteinus ; characters; valid species; notes).

— Sjöberg, 1930: 185 ( Proteinus ; cited as limbatus ; [Note: Misidentification: see Bernhauer, 1912i: 678; Horion, 1963: 28; Frank, 1979: 331]; characters; notes).

— Scheerpeltz, 1933: 1027 ( Proteinus ; catalog).

— Palm, 1948: 48 ( Proteinus ; characters; Sweden; Norway).

— Smetana, 1960b: 258 ( Proteinus ; Czechoslovakia).

— Horion, 1963: 27 ( Proteinus ; Caucasus; western Siberia; Britain; Norway; Sweden; Finland; Russia; Holland; France; Portugal; Corsica; Italy; Romania; Czechoslovakia; Austria; Germany).

— Lohse, 1964: 25 ( Proteinus ; characters; central Europe).

— Pope, 1977: 22 ( Proteinus ; Britain).

— Hammond, 1980: 135 ( Proteinus ; unconfirmed in Ireland).

— Tóth, 1982: 26 ( Proteinus ; characters; Hungary).

— Lucht, 1987: 85 ( Proteinus ; checklist; Germany; Poland; Austria; Switzerland; France; Benelux; Sweden).

— Boháč, 1993: 40 ( Proteinus ; Czech Republic; Slovakia).

— Sparacio, 1995: 137 ( Proteinus ; characters; notes; Sicily).

— Zanetti, 1995: 5 ( Proteinus ; Italy).

— Dauphin, 1995a: 106, 110, 118 ( Proteinus ; characters; France).

— Tronquet, 1998a: 114 ( Proteinus ; collected from marmot burrow; France).

- DISTRIBUTION: Caucasus, Russia, Finland, Norway, Sweden, Britain, Ireland, Holland, France, Germany, Czechoslovakia, Austria, Hungary, Romania, Italy, Portugal.

densipennis Bernhauer, 1912 i: 678 ( Proteinus ; Type locality: Kalifornien: Sugar Pine ).

— Scheerpeltz, 1933: 1027 ( Proteinus ; catalog).

- DISTRIBUTION: USA.

egregius Redtenbacher, 1874 , see: Acrulia .

ezoensis Hayashi, 1988: 17 ( Proteinus ; Type locality: Japan: Hokkaido: Mt. Kurodake, Daisetsu Mts. ).

- DISTRIBUTION: Japan.

fallax Mulsant and Rey, 1878 , see: laevigatus Hochhuth, 1872 .

flavocaudatus Bierig, 1940: 374 ( Proteinus ; Type locality: Costa Rica: Vara Blanca , 1800 m, entre los volcanes Barba y Poás).

- DISTRIBUTION: Costa Rica.

fuscotestaceus Cameron, 1924b: 162 ( Proteinus ; Type locality: Dehra Dun, U.P.).

— Cameron, 1930: 125 ( Proteinus ; characters; India).

— Scheerpeltz, 1933: 1027 ( Proteinus ; catalog).

— Coiffait, 1977: 244 ( Proteinus ; Nepal).

- DISTRIBUTION: India, Nepal.

gotoi Hayashi, 1986: 111 ( Proteinus ; Type locality: Mt. Amaishi , Sasayama-Cho,

Hyogo Pref.).

— Hayashi, 1988: 23 ( Proteinus ; key).

- DISTRIBUTION: Japan.

hibernus Gistel, 1857: 63 ( Proteinus ; [Note: Nomen dubium]; Type locality: Bavaria).

- DISTRIBUTION: Germany.

hyperboreus Muona, 1978: 125 ( Proteinus ; Type locality: USSR: Lim : Porjeguba. Also cited from Finland, Sweden, and Norway).

— Sjöberg, 1930: 182 ( Proteinus ; [Note: Misidentified as altaicus according to Muona, 1978: — Palm, 1948: 49 ( Proteinus ; [Note: Misidentified as altaicus according to Muona, 1978: 125]; characters; Sweden; Norway).

— Muona and Viramo, 1995: 9 ( Proteinus ; collected from dung of Ursus arctos ; Finland).

- DISTRIBUTION: Russia, Finland, Norway, Sweden.

kashmiricus Cameron, 1941: 56 ( Proteinus ; Type locality: Kashmir: Gulmarg).

- DISTRIBUTION: India.

laevigatus Hochhuth, 1872: 174 ( Proteinus ; Type locality: Not cited, but presumably from Gouvernements Kiew and/or Volhynien).

— Reitter, 1905a: 226 ( Proteinus ; characters; Russia).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

— Scheerpeltz, 1933: 1027 ( Proteinus ; catalog).

— Gusarov, 1992a: 775 [= 1993a: 62] ( Proteinus ; synonymic notes).

— Dauphin, 1995a: 106, 108, 118 ( Proteinus ; characters; France).

— Assing, Frisch, Kahlen, et al., 1998: 121 ( Proteinus ; macropterus of Gyllenhal is laevigatus ).

- DISTRIBUTION: Europe, Russia, Turkmenistan.

fallax Mulsant and Rey, 1878b: 230 ( Proteinus ; [Note: This name is cited as a variety of macropterus Gyllenhal which was a misidentification of macropterus Gravenhorst so I assume fallax refers to laevigatus ]; Type locality: environs de Montpellier).

— Ganglbauer, 1895: 760 ( Proteinus ; variety of macropterus Gyllenhal ).

— Scheerpeltz, 1933: 1028 ( Proteinus ; aberration of macropterus Gyllenhal ).

serrifer Muona, 1977: 15 ( Proteinus ; proposed for macropterus Gyllenhal [Note: Gyllenhal did not propose a new species, he attributed it to Gravenhorst; Gyllenhal’s use was a misidentification]; Type locality: Finland).

— Gyllenhal, 1810: 209 ( Omalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 34, and most other authors have cited macropterus Gyllenhal as valid species but Gyllenhal did not describe the species as new, he attributed it to Gravenhorst, 1806]; characters; Finland).

— Heer, 1839: 171 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Switzerland).

— Erichson, 1840: 903 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Germany).

— Hochhuth, 1849: 205 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Caucasus).

— Redtenbacher, 1849: 756 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 654 ( Proteinus ; cited as macropterus Gyllenhal ; characters; France).

— Redtenbacher, 1857: 257 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Austria).

— Kraatz, 1857d: 1025 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Germany).

— Thomson, 1861: 217 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Scandinavia).

— Pandellé, 1867: 169 ( Proteinus ; cited as macropterus Gyllenhal ; characters; France).

— Fauvel, 1871a: 57 [= 1872: 31] ( Proteinus ; cited as macropterus Gyllenhal ; characters; France; Britain; Germany; Switzerland; Russia).

— Redtenbacher, 1874: 282 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Austria).

— Seidlitz, 1875: 232 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Baltic region).

— J. Sahlberg, 1876: 224 ( Proteinus ; cited as macropterus Gyllenhal ; Finland).

— Mulsant and Rey, 1878b: 227 ( Proteinus ; cited as macropterus Gyllenhal ; characters; France).

— Fowler, 1888: 428 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Britain).

— Eppelsheim, 1892: 346 ( Protinus ; cited as macropterus Gyllenhal ; Turkmenia).

— Ganglbauer, 1895: 760 ( Proteinus ; cited as macropterus Gyllenhal ; characters; north and middle Europe).

— Reitter, 1905a: 228 ( Proteinus ; cited as macropterus Gyllenhal ; characters; northern and central Europe; Caucasus).

— Reitter, 1909: 199 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Germany).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; cited as macropterus Gyllenhal ; catalog).

— Johansen, 1914: 628 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Denmark).

— Portevin, 1929: 458 ( Proteinus ; cited as macropterus Gyllenhal ; characters; France).

— Sjöberg, 1930: 183 ( Proteinus ; cited as macropterus Gyllenhal ; characters).

— Scheerpeltz, 1933: 1027 ( Proteinus ; cited as macropterus Gyllenhal ; catalog). — Scheerpeltz and Höfler, 1948: 150 ( Proteinus ; cited as macropterus Gyllenhal ; fungus inhabitant). — Palm, 1948: 49 ( Proteinus ; cited as macropterus ; characters Gyllenhal; Sweden; Denmark; Norway; Finland). — Horion, 1963: 30 ( Proteinus ; cited as macropterus Gyllenhal ; Siberia; Scotland; Lapland; Norway; Sweden; Finland; Portugal; Spain; Corsica; Italy; Dalmatia; Bosnia; Greece; Germany; Austria). — Lohse, 1964: 25 ( Proteinus ; cited as atomarius Gyllenhal ; characters; central Europe). — Smetana, 1964d: 51 ( Proteinus ; cited as macropterus Gyllenhal ; Czechoslovakia). — Szujecki, 1968a: 710 ( Proteinus ; cited as macropterus Gyllenhal ; Poland). — Muona, 1979: 17 ( Proteinus ; synonym of macropterus Gyllenhal ). — Tóth, 1982: 28 ( Proteinus ; cited as macropterus Gyllenhal ; characters; Hungary). — H. Franz, 1984: 59 ( Proteinus , cited as macropterus Gyllenhal ; Macedonia). — Dettner and Reissenweber, 1991 ( Proteinus ; cited as macropterus Gyllenhal ; defensive secretions: chemistry and systematics). — Gusarov, 1992a: 775 [= 1993a: 62] ( Proteinus ; macropterus Gyllenhal is synonym of laevigatus ). — Cuccodoro, 1995: 262 ( Proteinus ; cited as macropterus Gyllenhal ; water loading behavior). — Assing, Frisch, Kahlen, et al., 1998: 121 ( Proteinus ; synonym of laevigatus ).

limbatus Mäklin, 1852: 323 ( Proteinus ; Type locality: insula Sitkha). — Fauvel, 1871a: 56 [= 1872: 30] ( Protinus ; characters; Alaska). — Fauvel, 1878d: 194 [= 1878e: 30] ( Proteinus ; characters; Sitka Island). — Scheerpeltz, 1933: 1027 ( Proteinus ; catalog). — Notman, 1919a: 96 ( Proteinus ; Ontario). — Hatch, 1957: 106 ( Proteinus ; characters; British Columbia ; Washington; Oregon; Idaho). — Frank, 1979: 331 ( Proteinus ; notes on nomenclature and North American records; limbatus is not Holarctic). — Frank and Thomas, 1984: 100 ( Proteinus ; characters). — Downie and Arnett, 1996: 429 ( Proteinus ; characters; USA). - DISTRIBUTION: Canada, USA.

maeklini Fauvel, 1869: 494 ( Proteinus ; named for limbatus variety b of Mäklin; Type locality: Not cited). — Fauvel, 1873b: 110 [= 1873c: 3] ( Protinus ; synonym of limbatus ). — Fauvel, 1878d: 195 [= 1878e: 31] ( Proteinus ; synonym of limbatus ). — Mulsant and Rey, 1878b: 226 ( Proteinus ; synonym of limbatus ). — Ganglbauer, 1895: 759 ( Proteinus ; synonym of limbatus ). — Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of limbatus ). — Scheerpeltz, 1933: 1027 ( Proteinus ; aberration of limbatus ). — Herman, 2001: 13 ( Proteinus ; spelling correct as maeklini ).

laevicollis Heer, 1839 , see: brachypterus Fabricius, 1792 .

lividipennis Reitter, 1905 , see: brachypterus Fabricius, 1792 .

longicollis Gredler, 1874 , see: Acrulia .

longicornis Dodero, 1923: 44 ( Proteinus ; Type locality: Monte Rosa, Macugnaga ). — Scheerpeltz, 1933: 1027 ( Proteinus ; catalog). — Lohse, 1964: 25 ( Proteinus ; characters; central Europe). — Focarile, 1982: 543 ( Proteinus ; valid species; characters; Italy; Switzerland; France). — Tóth, 1982: 27 ( Proteinus ; characters). — Lucht, 1987: 85 ( Proteinus ; checklist; Germany; Austria). — Zanetti, 1995: 5 ( Proteinus ; Italy). — Dauphin, 1995a: 106, 110, 118 ( Proteinus ; characters; France). — Burakowski, Mroczkowski, and Stefańska, 2000: 40 ( Proteinus ; catalog; Poland). - DISTRIBUTION: Italy, Switzerland, France.

macropterus Gravenhorst, 1806: 215 ( Omalium ; [Note: The authors of most of the references for “ Megarthrus macropterus (Gravenhorst) ” listed the species as a junior synonym of Megarthrus depressus (Paykull) ; citations of M. macropterus are found under Megarthrus prosseni Schatzmayr because M. depressus has been misidentified by all authors according to Cuccodoro and Löbl, 1997. Proteinus macropterus (Gravenhorst) has been cited as a valid species by a several authors beginning in 1977. Cuccodoro and Löbl, 1997, cited Megarthrus macropterus Gravenhorst as a nomen dubium and stated that the type had been destroyed in the Second World War. However, since the name has been cited in Proteinus fairly recently, it is permitted to remain there for now]; Type locality: Not cited).

— Gyllenhal, 1810: 209 ( Omalium ; characters; Finland).

— Olivier, 1811: 479 ( Omalium ; characters; Germany).

— Gyllenhal, 1827: 464 ( Omalium ).

— C. Sahlberg, 1827: 280 ( Omalium ; characters; Finland).

— Laporte, 1840: 193 ( Proteinus ; characters; France).

— Pope, 1977: 22 ( Proteinus ; cited as valid species; Britain).

— Hammond, 1980: 135 ( Proteinus ; cited as valid species; Ireland).

— Lucht, 1987: 85 ( Proteinus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Boháč, 1993: 40 ( Proteinus ; Czech Republic; Slovakia).

— Muona and Viramo, 1995: 9 ( Proteinus ; collected from fungus and dung of Ursus arctos ; Finland).

— Sparacio, 1995: 138 ( Proteinus ; characters; notes; Sicily).

— Zanetti, 1995: 5 ( Proteinus ; Italy).

- DISTRIBUTION: Finland, Britain, Ireland, France, Germany.

maeklini Fauvel, 1869 , see: limbatus Mäklin, 1852 .

minutus Mannerheim, 1830: 58 ( Proteinus ; Type locality: Willnäs).

— Mannerheim, 1831: 472 ( Proteinus ; Finland).

- DISTRIBUTION: Finland.

meuseli Dauphin, 1995: 193 ( Proteinus ; Type locality: Kroatien: Stirovaca. Also cited from Romania).

- DISTRIBUTION: Croatia, Romania.

nigriceps Fauvel, 1867 , see: Alloproteinus .

nigricornis Stephens, 1834 , see: brachypterus Fabricius, 1792 .

nitidus Stephens, 1834 , see: brachypterus Fabricius, 1792 .

oblongus Petri, 1891 , see: atomarius Erichson, 1840 .

olivieri Saulcy, 1866 , see: atomarius Erichson, 1840 .

ovalis Stephens, 1834: 335 ( Proteinus ; Type locality: London; Suffolk; Devonshire; Raehills ).

— Curtis, 1829: 28 ( Proteinus ; nomen nudum; catalog; Britain).

— Stephens, 1839: 424 ( Proteinus ; characters; England).

— Hardy, 1851: 63 ( Proteinus ; synonym of brachypterus ).

— Fauvel, 1871a: 56 [= 1872: 30] ( Proteinus ; characters; France; Scandinavia; Britain; Germany; Austria; Italy; Algeria).

— Fauvel, 1873b: 110 [= 1873c: 3] ( Protinus ; Sicily; Russia).

— Fauvel, 1874: 320 [= 1874b: 26] ( Protinus ; Spain).

— Fauvel, 1878: 87 [= 1878a: 7] ( Protinus ; Algeria).

— Fauvel, 1886: 13 [= 1886a: 5] ( Protinus ; Algeria).

— Fowler, 1888: 428 ( Proteinus ; characters; Britain).

— Ganglbauer, 1895: 759 ( Proteinus ; characters; middle Europe; western Mediterranean region).

— Fauvel, 1902b: 50 ( Proteinus ; Algeria; Tunisia).

— Reitter, 1905a: 227 ( Proteinus ; characters; central Europe; Mediterranean region).

— Sainte-Claire Deville, 1906: 67 ( Protinus ; Corsica).

— Reitter, 1909: 199 ( Proteinus ; characters; Germany).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

— Sjöberg, 1930: 180 ( Proteinus ; characters; notes).

— Scheerpeltz, 1933: 1028 ( Proteinus ; catalog).

— Hinton, 1945: 52 ( Proteinus ; characters).

— Scheerpeltz and Höfler, 1948: 150 ( Proteinus ; fungus inhabitant).

— Jeannel and Jarrige, 1949: 323 ( Proteinus ; cave species; Italy).

— Scheerpeltz, 1963a: 414 ( Proteinus ; Greece).

— Horion, 1963: 29 ( Proteinus ; Ireland; Britain; Holland; Belgium; France; Spain; Balearics; Corsica; Sicily; Italy; Croatia; Bosnia Hercegovina; Macedonia; Bulgaria; Romania; Slovakia; Czech Republic; Poland; Germany; Austria).

— Lohse, 1964: 24 ( Proteinus ; characters; central Europe).

— Pope, 1977: 22 ( Proteinus ; Britain).

— Coiffait, 1973a: 269 ( Proteinus ; Morocco).

— Osella and Zanetti, 1975: 83 ( Proteinus ; collected from nest of Talpa europaea ; notes; Italy).

— Hammond, 1980: 135 ( Proteinus ; Ireland).

— Tóth, 1982: 25 ( Proteinus ; characters; Hungary).

— Lucht, 1987: 85 ( Proteinus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Welch, 1993: 227 ( Proteinus ; ovariole number and ovary structure).

— Boháč, 1993: 40 ( Proteinus ; Czech Republic; Slovakia).

— Sparacio, 1995: 137 ( Proteinus ; characters; notes; Sicily).

— Zanetti, 1995: 5 ( Proteinus ; Italy).

— Dauphin, 1995a: 106, 110, 118 ( Proteinus ; characters; France).

— Tronquet, 1998a: 114 ( Proteinus ; collected from marmot burrow; France).

— Outerelo, Gamarra, and Salgado, 1998: 128 ( Proteinus ; collected from cave; Spain).

— Owen, 1999b: 61 ( Proteinus ; captured underground at roots of old trees; Britain).

— Owen, 2000: 251 ( Proteinus ; pitfall trap collections from underground at roots of trees; Britain).

— Assing, 2001: 75 ( Proteinus ; collected from xerothermous site; Germany).

- DISTRIBUTION: Europe, Russia.

subsulcatus Stephens, 1834: 336 ( Anthobium ; Type locality: London; Devonshire; Somersetshire ).

— Stephens, 1839: 424 ( Anthobium ; characters; England).

— G. Waterhouse, 1858: 32 ( Proteinus ; synonym of ovalis ).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of ovalis ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of ovalis ).

brevicollis Erichson, 1840: 903 ( Proteinus ; Type locality: Italia; Lutetiae ).

— Fairmaire and Laboulbène, 1856: 653 ( Proteinus ; characters; France).

— Redtenbacher, 1857: 995 ( Proteinus ; characters; Germany).

— Kraatz, 1857d: 1024 ( Proteinus ; characters; Germany).

— G. Waterhouse, 1858: 32 ( Proteinus ; synonym of ovalis ).

— Pandellé, 1867: 168 ( Proteinus ; characters; France).

— Fauvel, 1871a: 56 [= 1872: 30] ( Proteinus ; synonym of ovalis ).

— Redtenbacher, 1874: 282 ( Proteinus ; characters).

— Mulsant and Rey, 1878b: 220 ( Proteinus ; characters; France).

— Ganglbauer, 1895: 759 ( Proteinus ; synonym of ovalis ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; synonym of ovalis ).

— Scheerpeltz, 1933: 1028 ( Proteinus ; synonym of ovalis ).

ovatus Gravenhorst, 1806 , see: brachypterus Fabricius, 1792 .

parvulus LeConte, 1863 a: 58 ( Proteinus ; Type locality: Lake Superior ).

— Fauvel, 1871a: 57 [= 1872: 31] ( Proteinus ; synonym of clavicornis ).

— Fauvel, 1878d: 196 [= 1878e: 32] ( Proteinus ; synonym of atomarius ).

— Ganglbauer, 1895: 760 ( Proteinus ; synonym of atomarius ).

— Reitter, 1905a: 228 ( Proteinus ; synonym of atomarius ).

— Bernhauer and Schubert, 1910: 33 ( Proteinus ; synonym of atomarius ).

— Frank, 1979: 331, 335 ( Proteinus ; valid species; characters).

— Frank and Thomas, 1984: 100 ( Proteinus ; characters).

— Campbell and Davies, 1991: 87 ( Proteinus ; synonym of atomarius ; [Note: I assume the authors overlooked the elevation of this name]).

- DISTRIBUTION: USA.

peckorum Frank, 1984 a: 98 ( Proteinus ; Type locality: Jamaica: Clarendon Jackson Bay Cave ).

— Bordoni and Oromi, 1998: 1157 ( Proteinus ; facultative cave species; Jamaica).

- DISTRIBUTION: Jamaica.

planicollis Reitter, 1905: 201 ( Proteinus ; Type locality: Russisch-Armenien: Araxestal bei Ordubad).

— Reitter, 1905a: 226 ( Proteinus ; characters; Araxes Valley).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

— Scheerpeltz, 1933: 1028 ( Proteinus ; catalog).

- DISTRIBUTION: Azerbaijan.

reflexicollis Reitter, 1905 a: 226 ( Proteinus ; Type locality: Araxestal bei Ordubad, Russisch-Armenien ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

— Scheerpeltz, 1933: 1028 ( Proteinus ; catalog).

- DISTRIBUTION: Azerbaijan.

ruficollis Cameron, 1924b: 162 ( Proteinus ; Type locality: Narkanda, 9000 feet; Fagu,

8000 feet; Theog, 7600 feet, in the Simla Hills; Mussorie, 7000 feet).

— Cameron, 1930: 124 ( Proteinus ; characters; India).

— Scheerpeltz, 1933: 1028 ( Proteinus ; catalog).

— Coiffait, 1982: 40 ( Proteinus ; India).

- DISTRIBUTION: India.

salebrosus Casey, 1885 a: 323 ( Proteinus ; cited as Protinus ; Type locality: California:

Santa Cruz).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

— Scheerpeltz, 1933: 1028 ( Proteinus ; catalog).

- DISTRIBUTION: USA.

sawadai Hayashi, 1988: 18 ( Proteinus ; Type locality: Japan: Nagano Pref.: Mt. Kisokoma , about 2000 m.).

- DISTRIBUTION: Japan.

serrifer Muona, 1977 , see: laevigatus Hochhuth, 1872 .

shibatai Hayashi, 1986: 109 ( Proteinus ; Type locality: Mt. Ifuri , Echizen-Ohno).

— Hayashi, 1988: 22 ( Proteinus ; key).

- DISTRIBUTION: Japan.

siculus Dodero, 1923: 43 ( Proteinus ; Type locality: Sicilia: Castelbuono ).

— Scheerpeltz, 1933: 1028 ( Proteinus ; catalog).

— Sparacio, 1995: 137 ( Proteinus ; characters; notes; Sicily).

— Zanetti, 1995: 5 ( Proteinus ; Italy).

— Dauphin, 1995a: 104, 108 ( Proteinus ; characters).

- DISTRIBUTION: Italy.

subsulcatus Stephens, 1834 , see: ovalis Stephens, 1834 .

sulcatus Fauvel, 1878d: 195 [= 1878e: 31] ( Proteinus ; Type locality: Californie, Mariposa ).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

- DISTRIBUTION: USA.

tateoitoi Hayashi, 1988: 20 ( Proteinus ; Type locality: Japan: Japan: Kyoto Pref.: Hanase ).

- DISTRIBUTION: Japan.

thomasi Frank, 1979: 333 ( Proteinus ; Type locality: U.S.A.: Florida : Marion Co., Ocala. Also cited from New Jersey).

— Frank and Thomas, 1984: 100 ( Proteinus ; characters).

— Lundgren, 1998: 38 ( Proteinus ; Florida; New Jersey).

- DISTRIBUTION: USA.

zanettii Dauphin, 1999: 138 ( Proteinus ; cited as “ zanetti ” at head of description, but as “ zanettii ” in abstract; Type locality: Grecia: Polludroso).

- DISTRIBUTION: Greece.

Tribe Silphotelini

Silphotelini Newton and Thayer, 1995: 247–286 , 297 (description; phylogenetic relationships; includes Silphotelus and Alloproteinus ; discussion).

— Klimaszewski, Newton, and Thayer, 1996: 146 (list of New Zealand genera; number of New Zealand species).

Alloproteinus

[1 species; Neotropical Region]

Alloproteinus Steel, 1966: 300 (species included: nigriceps ). TYPE SPECIES: Proteinus nigriceps Fauvel , fixed by original designation and monotypy.

— Newton and Thayer, 1995: 297 ( Silphotelini ).

nigriceps Fauvel, 1867 a: 50 [= 1868: 55] ( Proteinus ; Type locality: Santiago).

— Bernhauer and Schubert, 1910: 34 ( Proteinus ; catalog).

— Steel, 1966: 302 ( Alloproteinus ; [Note: Although the species has remained in Proteinus in the works of other authors, I retain it in Alloproteinus because I assume that Steel’s work was overlooked and that the genera have not been synonymized]; characters; Chile).

— Coiffait and Saiz, 1968: 457 ( Proteinus ; characters; Chile).

— Scheerpeltz, 1972: 8 ( Proteinus ; Argentina).

- DISTRIBUTION: Argentina, Chile.

Silphotelus

[2 species; Australian Region]

Silphotelus Broun, 1895: 83 (species included: nitidus ). TYPE SPECIES: Silphotelus nitidus

Broun, fixed by monotypy.

— Steel, 1966: 288, 303 (adult characters; larval characters, cited as Silhpotelus; key to species). — Newton and Thayer, 1995: 297 ( Silphotelini ).

— Klimaszewski, Newton, and Thayer, 1996: 146 (endemic to New Zealand and nearby islands). nitidus Broun, 1895: 84 ( Silphotelus ; Type locality: Hunua Range).

— Kuschel, 1990: 25 ( Silphotelus ; endemic; winged; habitat; New Zealand).

— Steel, 1966: 304 ( Silphotelus ; characters; New Zealand).

— Klimaszewski and Watt, 1997: 170 ( Silphotelus ; characters; collecting notes; New Zealand).

- DISTRIBUTION: New Zealand.

obliquus Broun, 1912: 416 ( Silphotelus ; Type locality: Greymouth).

— Steel, 1966: 306 ( Silphotelus ; characters; New Zealand).

- DISTRIBUTION: New Zealand.