Philorinum nitidulum, Kraatz, 1857 d: 966

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.1

persistent identifier

https://treatment.plazi.org/id/BF684949-423C-B5F7-00DC-2DB4FB577CB2

treatment provided by

Felipe

scientific name

Philorinum nitidulum
status

 

nitidulum Kraatz, 1857d: 966 View in CoL , note ( Philorinum ; Type locality: Griechenland).

— Kraatz, 1858b: 128 ( Philorinum ; characters; Greece).

— Mulsant and Rey, 1880: 158 ( Philorinum ; synonym of humile ).

— Ganglbauer, 1895: 727 ( Philorinum ; synonym of sordidum ).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; synonym of sordidum ).

— Scheerpeltz, 1933: 1052 ( Philorinum ; synonym of sordidum ).

ruficolle Schaufuss, 1861: 47 ( Philorinum ; cited as rufieolle ; Type locality: Hisp. mer.). — Fauvel, 1876a: 51 [= 1876: 236] ( Philorinum ; synonym of sordidum ).

— Mulsant and Rey, 1880: 158 ( Philorinum ; synonym of humile ).

— Ganglbauer, 1895: 727 ( Philorinum ; synonym of sordidum ).

— Fauvel, 1902b: 55 ( Philorinum ; synonym of sordidum ).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; synonym of sordidum ).

cadomense Fauvel, 1863 a: 42 ( Philorinum ; Type locality: Calvados, Ille-et-Vilaine).

— Mulsant and Rey, 1880: 156 ( Philorinum ; synonym of humile ).

— Ganglbauer, 1895: 727 ( Philorinum ; synonym of sordidum ).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; synonym of sordidum ).

brevipenne Luze, 1910: 240 ( Anthobium ; Type locality: Andalusien).

— Scheerpeltz, 1933: 1033 ( Anthobium ; catalog).

— C. Koch, 1940: 372 ( Anthobium ; characters; Spain).

— Zanetti, 1986: 95 ( Philorinum ; synonym of sordidum ).

marocanum Coiffait, 1978 r: 271 ( Eusphalerum ; Type locality: Maroc: Sidi Bettache). — Zanetti, 1991a: 29 ( Philorinum ; synonym of sordidum ).

subpubescens Stephens, 1834 , see: sordidum Stephens, 1834 .

Philhydrodema

[1 species; Palaearctic Region]

Philhydrodema Scheerpeltz, 1976: 11 (species included: longelytratum ). TYPE SPECIES: Philhydrodema longelytratum Scheerpeltz , fixed by original designation.

longelytratum Scheerpeltz, 1976: 12 ( Philhydrodema ; Type locality: Nepal: Nare-Glacier

[= Ama-Dablam-Gletscher] oberhalb der Green Hut der Hillary-Makalu-Expedition, 5570 m).

- DISTRIBUTION: Nepal.

Philydrodes

[26 species; Palaearctic Region]

Philydrodes Bernhauer, 1929 g: 179 (species included: aquatilis ). TYPE SPECIES: Anthophagus aquatilis Sharp , fixed by original designation and monotypy.

— Scheerpeltz, 1933: 1067 (world catalog supplement).

— Blackwelder, 1952: 303 (type species: aquatilis ).

— Nakane and Sawada, 1956a: 179 (characters; key to Japanese species; type species: aquatilis ).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Shibata, 1976: 119 (checklist of species of Japan).

— Watanabe, 1990: 309 (characters; key to species of Japan).

— Kashcheev, 1999: 146 (characters).

Minyphilydrodes Watanabe, 1990: 334 (subgenus of Philydrodes ; species included: watanabei , asamanis , parallelus , maculatus , ishidai , biguttatus , flavidus , shimomurai , chichibunis , niger ). TYPE SPECIES: Philydrodes watanabei R. Dvořák , fixed by original designation.

aquatilis Sharp, 1889: 469 ( Anthophagus ; Type locality: Hakone).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog).

— Bernhauer, 1929g: 180 ( Philydrodes ).

— Scheerpeltz, 1933: 1067 ( Philydrodes ; catalog).

— Nakane, 1963a: 83 ( Philydrodes ; characters; habitus photograph; Japan).

— Watanabe, 1990: 314 ( Philydrodes ; characters; Japan).

— Watanabe, 1996a: 9 ( Philydrodes ; cave collection; Japan).

- DISTRIBUTION: Japan.

japonicus Yoshida and Nomura, 1952: 6 ( Coryphium ; Type locality: Aoiwa L. S. C., Kanto-

Mountainland, Japan).

— Shibata, 1970: 57 (suggested transfer from Coryphium to Philydrodes ).

— Watanabe, 1990: 314 ( Philydrodes ; synonym of aquatilis ).

— Zerche, 1990: 18 ( Philydrodes ). asamanis Watanabe, 1990: 337 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan:

Honshu: Gunma Pref.: near Nidoage).

- DISTRIBUTION: Japan. biguttatus Watanabe, 1990: 343 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality:

Japan: Honshu: Nagano Pref.: Hakuba-mura).

- DISTRIBUTION: Japan. chichibunis Watanabe, 1990: 347 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality:

Japan: Honshu: Saitama Pref.: Kawamata, Chichibu).

- DISTRIBUTION: Japan. dewanus Watanabe, 1990: 325 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan:

Honshu: Yamagata Pref.: Tachiyazawa, Higashitagawa).

- DISTRIBUTION: Japan. flavidus Watanabe, 1990: 344 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan:

Honshu: Yamanashi Pref.: Mt. Kitazawa-tôge, S. Jpn. Alps).

- DISTRIBUTION: Japan. gigas Watanabe, 1990: 317 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu:

Shizuoka Pref.: Umegashima Spa).

- DISTRIBUTION: Japan. grandis Nakane and Sawada, 1956 a: 181 ( Philydrodes ; Type locality: Japan: Kyushu:

Chinoike-jigoku, Beppu).

— Watanabe, 1990: 320 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan. hikosanensis Nakane and Sawada, 1956 a: 180 ( Philydrodes ; Type locality: Japan: Kyushu:

Hikosan).

— Nakane, 1963a: 83 ( Philydrodes ; characters; habitus photograph; Japan).

— Watanabe, 1990: 318 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan. horioi Nakane and Sawada, 1956 a: 180 ( Philydrodes ; Type locality: Japan: Sampuku Pass,

Nagano).

— Watanabe, 1990: 316 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan. ishidai Nakane and Sawada, 1956 a: 182 ( Philydrodes ; Type locality: Kamikochi, Nagano).

— Nakane, 1963a: 83 ( Philydrodes ; characters; habitus photograph; Japan).

— Watanabe, 1990: 342 ( Philydrodes ; subgenus Minyphilydrodes ; characters; Japan).

- DISTRIBUTION: Japan. japonicus Yoshida and Nomura, 1952 , see: aquatilis Sharp, 1889 . kawaseorum Hayashi, 1994 a: 133 ( Philydrodes , cited as Phylidrodes; Type locality: Japan:

Mie: Jirakura).

- DISTRIBUTION: Japan. maculatus Watanabe, 1990: 340 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality:

Japan: Honshu: Yamanashi Pref.: Hirogawara, foot of Mt. Kitadake).

michikoae Watanabe, 1990: 323 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu: Yamagata Pref.: Mt. Atsumidake, Nishitagawa).

- DISTRIBUTION: Japan.

minipunctatus Watanabe, 1990: 321 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu: Nagano Pref.: Kamikôchi).

- DISTRIBUTION: Japan.

niger Watanabe, 1990: 349 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan: Honshu: Yamanashi Pref.: Hirogawara, foot of Mt. Kitadake).

- DISTRIBUTION: Japan.

parallelus Watanabe, 1990: 338 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan: Honshu: Nagano Pref.: Kamikôchi).

- DISTRIBUTION: Japan.

pulla Nakane and Sawada, 1956 , see: Liophilydrodes .

puncticollis Nakane and Sawada, 1956 , see: Liophilydrodes .

rubidus Nakane and Sawada, 1956 a: 180 ( Philydrodes ; variety of aquatilis ; Type locality: Japan: Lake Marunuma, Nikko).

— Watanabe, 1990: 331 ( Philydrodes ; valid species; characters; Japan).

- DISTRIBUTION: Japan.

rufescens Watanabe, 1990: 327 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Shikoku: Ehime Pref.: western ravine of Mt. Shiratsue).

- DISTRIBUTION: Japan.

shimomurai Watanabe, 1990: 346 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan: Honshu: Mt. Sanpuku-tôge, S. Jpn. Alps).

- DISTRIBUTION: Japan.

similis Nakane and Sawada, 1956 a: 181 ( Philydrodes ; subgenus Philydrodes ; variety of horioi ; Type locality: Japan: Nagano, Sampuku Pass).

— Watanabe, 1990: 330 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

takaoensis Nakane and Sawada, 1956 a: 182 ( Philydrodes ; Type locality: Japan: Takaoyama , Tokyo).

— Watanabe, 1990: 334 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

troglophila Nakane and Sawada, 1956 , see: Liophilydrodes .

tsurugisanus Watanabe, 1990: 326 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Shikoku: Tokushima Pref.: Ryôtsurugi-dans on Mt. Tsurugi-san).

- DISTRIBUTION: Japan.

yoshidai Watanabe, 1990: 328 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Shikoku: Tokushima Pref.: Ryôtsurugidani on Mt. Tsurugi-san).

- DISTRIBUTION: Japan.

watanabei R. Dvořák, 1957: 9 ( Philydrodes ; Type locality: Itsukaichi , Nishitama, Tokyo, Japon).

— Watanabe, 1990: 335 ( Philydrodes ; subgenus Minyphilydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

zipanguensis Watanabe, 1990: 333 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu: Nagano Pref.: Kamikôchi).

- DISTRIBUTION: Japan.

Phlaeopterus

[14 species; Nearctic Region]

Phlaeopterus Motschulsky, 1853 a: 78 (species included: fusconiger ). TYPE SPECIES:

— Kraatz, 1857d: 930 (synonym of Lesteva ).

— Gemminger and Harold, 1868: 659 (synonym of Lesteva ).

— Fauvel, 1871a: 127 [= 1872: 101] (synonym of Lesteva ).

— Fauvel, 1875a:VII [= 1875b: 209] (synonym of Lesteva ).

— Bernhauer and Schubert, 1910: 74 (synonym of Tilea ).

— Scheerpeltz, 1933: 1065 (world catalog supplement).

— Blackwelder, 1952: 303 (type species: fusconiger ).

— Hatch, 1957: 49, 57 (characters; key to species of Pacific Northwest).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1975: 208 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 192 (characters; notes).

— Campbell and Davies, 1991: 90 (checklist of species; Alaska and Canada).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (14 Nearctic species; notes; characters in key). Tilea Fauvel, 1878d: 246 [= 1878e: 82] (species included: cavicollis ). TYPE SPECIES: Tilea cavicollis Fauvel , fixed by monotypy.

— Duvivier, 1883: 195 (catalog).

— Casey, 1894: 402 (notes; key to North American species).

— Bernhauer and Schubert, 1910: 74 (world catalog; 7 species).

— Leng, 1920: 95 (catalog of North American species).

— Scheerpeltz, 1933: 1065 (synonym of Phlaeopterus ).

— Blackwelder, 1952: 390 (type species: cavicollis ; synonym of Phlaeopterus ).

brevipennis Casey, 1894: 404 ( Tilea ; Type locality: Wyoming).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 60 ( Phlaeopterus ; characters; British Columbia ; Washington; Oregon; Idaho; Montana).

- DISTRIBUTION: Canada, USA.

cascadiensis Hatch, 1957: 59 ( Phlaeopterus ; Type locality: Washington: Mt. Rainier ,

Tipsoo L.; British Columbia ; Oregon).

- DISTRIBUTION: Canada, USA.

castaneus Casey, 1894: 404 ( Tilea ; Type locality: Colorado).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

- DISTRIBUTION: USA.

cavicollis Fauvel, 1878d: 246 [= 1878e: 82] ( Tilea ; Type locality: Colombie britannique).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 60 ( Phlaeopterus ; characters; British Columbia ; Washington; Idaho).

- DISTRIBUTION: Canada, USA.

filicornis Casey, 1886: 234 ( Phlaeopterus , cited as Phloeopterus ; Type locality: California: Placer Co.).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

- DISTRIBUTION: USA.

frosti Hatch, 1957: 59 ( Phlaeopterus ; Type locality: Washington: Mt. Rainier, below Sluskin Falls. Also cited from British Columbia and Oregon).

- DISTRIBUTION: Canada, USA.

fusconiger Motschulsky, 1853 a: 78 ( Phlaeopterus ; Type locality: Ounalaschka).

— Mäklin, 1853: 193 ( Lesteva ; characters; Unalaska).

— LeConte, 1866: 375 ( Lesteva ; California).

— Fauvel, 1878d: 248 [= 1878e: 84] ( Lesteva ; characters; Alaska).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

- DISTRIBUTION: USA.

houkae Hatch, 1957: 58 ( Phlaeopterus ; Type locality: Washington: Mt. Baker Lodge. Also cited from British Columbia ).

- DISTRIBUTION: Canada, USA.

kootenayensis Hatch, 1957: 58 ( Phlaeopterus ; Type locality: British Columbia : Bull River,

6 mi. from mouth).

- DISTRIBUTION: Canada.

lagrandeuri Hatch, 1957: 58 ( Phlaeopterus ; Type locality: Seattle, Carkeek Park, Washington. Also cited from Oregon and British Columbia ).

- DISTRIBUTION: Canada, USA.

loganensis Hatch, 1957: 60 ( Phlaeopterus ; Type locality: G[lacier] N[ational] P[ark]: Logan Pass, Montana. Also cited from British Columbia and Idaho).

- DISTRIBUTION: Canada, USA.

longipalpus Casey, 1885 a: 318 ( Phlaeopterus , cited as Phloeopterus ; Type locality: California: Middle Sierras).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 60 ( Phlaeopterus ; characters; British Columbia ; Washington; Oregon; Idaho; Utah).

- DISTRIBUTION: Canada, USA.

rufitarsis Casey, 1894: 403 ( Tilea ; Type locality: California: Siskiyou Co.).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 59 ( Phlaeopterus ; characters; British Columbia ; California; Montana).

- DISTRIBUTION: Canada, USA.

stacesmithi Hatch, 1957: 60 ( Phlaeopterus ; Type locality: British Columbia : Barkerville, Roundtop Mt., 6000 ft.).

- DISTRIBUTION: Canada.

Phyllodrepoidea

[1 species; Palaearctic and Nearctic Regions]

Phyllodrepoidea Ganglbauer, 1895: 724 (species included: crenata ). TYPE SPECIES: Phyllodrepoidea crenata Ganglbauer , fixed by monotypy (see Herman, 2001).

— Luze, 1905c: 247 (characters).

— Heyden, Reitter, and Weise, 1906: 133 (list of species of Europe and Caucasus).

— Reitter, 1909: 188 (characters).

— Bernhauer and Schubert, 1910: 63 (world catalog; 1 species).

— Petri, 1912: 54 (list of species of Siebenbürgen).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Porta, 1926: 24 (characters; species of Italy).

— Roubal, 1930: 312 (catalog; Slovakia).

— Joy, 1932: 102 (key to British species).

— Scheerpeltz, 1933: 1054 (world catalog supplement).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 96 (characters; species of Sweden).

— Tottenham, 1949: 356, 407 (type species: crenata Gravenhorst ; list of British species).

— Blackwelder, 1952: 307 (type species: crenata ).

— Tottenham, 1954: 28 (characters; British species).

— Horion, 1963: 101 (list of central European species).

— Székessy, 1963: 18 (characters).

— Lohse, 1964: 51 (characters; key to central European species).

— Kasule, 1968: 133, 134 (larval characters).

— Scheerpeltz, 1968a: 13 (catalog of Austrian species).

— Steel, 1970a: 17 (larval characters; biological notes).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Pope, 1977: 22 (list of British species).

— Burakowski, Mroczkowski, and Stefańska, 1979: 52 (catalog; Poland).

— Muona, 1979: 18 (list of Scandinavian species).

— Tóth, 1982: 74 (characters; habitus; species of Hungary).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Boháč, 1993: 41 (in list of species of Czech Republic and Slovakia).

— Zanetti, 1995: 8 (list of species of Italy).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (1 undescribed Nearctic species; notes; characters in key).

— Herman, 2001: 8 (type species; discussion).

creatoris Gozis, 1886 , see: crenata Gravenhorst, 1802 .

crenata Ganglbauer, 1895: 724 ( Phyllodrepoidea ; Type locality: Mittel- und Südeuropa).

— Gravenhorst, 1802: 114 ( Omalium ; [Note: Misidentification: Gravenhorst attributed this name to Paykull who attributed it to Fabricius whose species is cited in this catalog in Acidota . All subsequent authors have attributed the species to Gravenhorst]).

— Latreille, 1804: 371 ( Omalium ; characters).

— Gravenhorst, 1806: 207 ( Omalium ; characters).

— C. Sahlberg, 1830: 288 ( Omalium ; characters; Finland).

— Erichson, 1840: 873 ( Deliphrum ; characters; Germany).

— Redtenbacher, 1849: 829 ( Deliphrum ; characters).

— Fairmaire and Laboulbène, 1856: 638 ( Deliphrum ; characters; France).

— Redtenbacher, 1857: 249 ( Deliphrum ; characters; Germany).

— Kraatz, 1857d: 954 ( Deliphrum ; characters; Germany).

— Thomson, 1871: 377 ( Deliphrum ; characters; Sweden).

— Fauvel, 1871a: 122 [= 1872: 96] ( Deliphrum ; characters; France; Britain; Germany; Switzerland; Italy; Austria; Corsica; Sicily).

— Redtenbacher, 1874: 270 ( Deliphrum ; characters).

— Mulsant and Rey, 1880: 112 ( Deliphrum ; characters; France).

— Fowler, 1888: 406 ( Deliphrum ; characters; Britain).

— Ganglbauer, 1895: 724 ( Phyllodrepoidea ; characters; middle and southern Europe).

— Luze, 1905c: 249 ( Phyllodrepoidea ; characters; Hungary; Croatia; Bosnia; France, Italy, Greece; Germany; Switzerland; Corsica; Sicily).

— Sainte-Claire Deville, 1906: 71 ( Phyllodrepoidea ; Corsica).

— Sainte-Claire Deville, 1907: 42 ( Phyllodrepoidea ; characters; France).

— Reitter, 1909: 188 ( Phyllodrepoidea ; characters; Germany).

— Bernhauer and Schubert, 1910: 63 ( Phyllodrepoidea ; catalog).

— Portevin, 1929: 442 ( Phyllodrepoidea ; characters; France).

— Scheerpeltz, 1933: 1054 ( Phyllodrepoidea ; catalog).

— Palm, 1948: 97 ( Phyllodrepoidea ; characters; Sweden; Norway).

— Blackwelder, 1952: 307 ( Phyllodrepoidea ; type species).

— Tottenham, 1954: 28 ( Phyllodrepoidea ; characters; Scotland).

— Székessy, 1963: 17 ( Phyllodrepoidea ; illustration).

— Horion, 1963: 101 ( Phyllodrepoidea ; Scotland; Norway; Sweden; France; Belgium; Switzerland; Corsica; Sicily; Italy; Greece; Croatia; Bosnia Hercegovina; Hungary; Romania; Slovakia; Germany; Austria).

— Lohse, 1964: 51 ( Phyllodrepoidea ; characters; central Europe).

— Kasule, 1968: 134 ( Phyllodrepoidea ; larval characters).

— Scheerpeltz, 1968a: 13 ( Phyllodrepoidea ; catalog; Austria).

— Steel, 1970a: 17 ( Phyllodrepoidea ; larval characters; biological notes).

— Tikhomirova, 1973a: 139 ( Phyllodrepoidea ; checklist; Europe).

— Pope, 1977: 22 ( Phyllodrepoidea ; Britain).

— Muona, 1979: 18 ( Phyllodrepoidea ; Sweden; Norway).

— Burakowski, Mroczkowski, and Stefańska, 1979: 52 ( Phyllodrepoidea ; notes; Poland).

— Hammond, 1980: 137 ( Phyllodrepoidea ; notes; Ireland).

— Tóth, 1982: 74 ( Phyllodrepoidea ; characters; Hungary).

— Crowson, 1982: 125 ( Phyllodrepoidea ; natural history observations; Britain).

— Terlutter, 1984: 26, 41 ( Phyllodrepoidea ; habitat notes; Germany).

— Segers, 1986: 21 ( Phyllodrepoidea ; Belgium).

— Lucht, 1987: 88 ( Phyllodrepoidea ; Germany; Poland; Austria; Switzerland; France; Benelux; Sweden).

— Silfverberg, 1992: 22 ( Phyllodrepoidea ; Sweden; Norway).

— Boháč, 1993: 41 ( Phyllodrepoidea ; Slovakia; Czech Republic).

— Allen, 1994: 247 ( Phyllodrepoidea ; Britain).

— Zanetti, 1995: 8 ( Phyllodrepoidea ; Italy).

— Zanetti and Sabella, 1998: 31 ( Phyllodrepoidea ; notes; Sicily).

— M. Hansen, Pedersen, and Pritzl, 1999: 77 ( Phyllodrepoidea ; characters; notes; Denmark).

— Fowles, Alexander, and Key, 1999: 129 ( Phyllodrepoidea ; saproxylic species; status as endangered or threatened species; Britain).

— Herman, 2001: 8, 21 ( Phyllodrepoidea ; type species of genus; Ganglbauer author species not Gravenhorst who attributed it to Fabricius; nomen protectum).

- DISTRIBUTION: Europe.

creatoris Gozis, 1886: 15 ( Deliphrum ; proposed for Gravenhorst’s misidentification of crenatum ; Type locality: Not cited).

— Bernhauer and Schubert, 1910: 63 ( Phyllodrepoidea ; synonym of crenata ).

— Herman, 2001: 21 ( Phyllodrepoidea ; synonym of crenata ; nomen oblitum).

Porrhodites

[2 species; Nearctic and Palaearctic Regions]

Porrhodites Kraatz, 1857d: 961 (species included: brevicollis ). TYPE SPECIES: Arpedium brevicolle Mäklin , fixed by monotypy.

— Redtenbacher, 1857: 992 (characters).

— Thomson, 1859: 50 (characters; type species: fenestralis ).

— Schaum, 1859: 32 (catalog; European species).

— LeConte, 1861a: 70 (characters).

— Thomson, 1861: 195 (characters).

— LeConte, 1863: 26 (list of North American species).

— Gemminger and Harold, 1868: 663 (catalog).

— Fauvel, 1871a: 111 [= 1872: 85] (characters).

— Redtenbacher, 1874: 271 (characters).

— Fauvel, 1875a:VI [= 1875b: 208] (catalog).

— Heyden, 1880: 82 (list of species of Siberian region).

— Heyden, Reitter, and Weise, 1891: 119 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 726 (characters).

— Heyden, Reitter, and Weise, 1906: 133 (list of species of Europe and Caucasus).

— Reitter, 1909: 184 (characters).

— Bernhauer and Schubert, 1910: 62 (world catalog; 62 species).

— Leng, 1920: 94 (catalog of North American species).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Porta, 1926: 24 (characters; species of Italy).

— Scheerpeltz, 1933: 1053 (world catalog supplement).

— Palm, 1948: 92 (characters; species of Sweden).

— Blackwelder, 1952: 320 (type species: brevicollis ).

— Hatch, 1957: 50, 69 (characters; species of Pacific Northwest).

— Horion, 1963: 98 (list of central European species).

— Székessy, 1963: 16 (characters).

— Lohse, 1964: 49 (characters; key to central European species).

— Moore, 1966: 48, 51 (characters; notes).

— Scheerpeltz, 1968a: 12 (catalog of Austrian species).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Moore and Legner, 1975: 210 (catalog; North America north of Mexico).

— Burakowski, Mroczkowski, and Stefańska, 1979: 52 (catalog; Poland).

— Moore and Legner, 1979: 193 (characters; notes).

— Tóth, 1982: 69 (characters; habitus; key to species of Hungary).

— Campbell, 1984a: 1228 (characters; review of and key to North American species).

— Zanetti, 1987: 267 (characters; habitus; species of Italy).

— Campbell and Davies, 1991: 91 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Downie and Arnett, 1996: 433 (characters; key to species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (2 Nearctic species; notes; characters in key).

brevicollis Mäklin, 1852 , see: fenestralis Zetterstedt, 1828 .

fenestralis Zetterstedt, 1828: 50 ( Omalium ; Type locality: Lapponia Tornensis).

— Zetterstedt, 1838: 50 ( Omalium ; characters; habitat; Lapland).

— Erichson, 1840: 888 ( Omalium ; characters; Lapland).

— Kraatz, 1857d: 963 ( Porrhodites ; synonym of brevicollis ).

— Thomson, 1859: 50 (type species of Porrhodites ).

— Thomson, 1861: 196 ( Porrhodites ; characters; Scandinavia).

— Fauvel, 1871a: 111 [= 1872: 85] ( Porrhodites ; characters; Switzerland; Lapland; Alaska).

— Fauvel, 1874: 323 [= 1874b: 29] ( Porrhodites ; Finland).

— J. Sahlberg, 1876: 214 ( Porrhodites ; Finland).

— Fauvel, 1878d: 223 [= 1878e: 59] ( Porrhodites ; characters; Alaska; British Columbia ; Lake Superior).

— Seidlitz, 1891: 344 ( Porrhodites ; characters; Europe).

— Seidlitz, 1891a: 366 ( Porrhodites ; characters; Europe).

— Ganglbauer, 1895: 726 ( Porrhodites ; characters; Lapland; Finland; Siberia; North America ).

— Poppius, 1909: 5 ( Porrhodites ; Lena Valley, Siberia ).

— Reitter, 1909: 184 ( Porrhodites ; characters; Germany).

— Bernhauer and Schubert, 1910: 62 ( Porrhodites ; catalog).

— Scheerpeltz, 1933: 1053 ( Porrhodites ; catalog).

— Palm, 1948: 93 ( Porrhodites ; characters; Sweden; Norway; Finland).

— Hatch, 1957: 69 ( Porrhodites ; characters; British Columbia ).

— Horion, 1963: 98 ( Porrhodites ; Norway; Sweden; Finland; Russia; Poland; Germany; Czechoslovakia; Austria; Corsica).

— Lohse, 1964: 49 ( Porrhodites ; characters; central Europe).

— Tóth, 1982: 69 ( Porrhodites ; characters).

— Campbell, 1984a: 1230 ( Porrhodites ; characters; Alberta; British Columbia ; Manitoba; Newfoundland; Labrador; Northwest Territories; Quebec; Yukon Territory; Alaska; Montana; Oregon; Wyoming).

— Zanetti, 1987: 267 ( Porrhodites ; characters; Italy).

— Lucht, 1987: 88 ( Porrhodites ; checklist; Austria).

— Boháč, 1993: 41 ( Porrhodites ; Czech Republic).

— Downie and Arnett, 1996: 433 ( Porrhodites ; characters; Canada; USA).

— Ryabukhin, 1999: 20 ( Porrhodites ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Norway, Sweden, Finland, Russia, Poland, Germany, Czechoslovakia, Austria, Italy, Switzerland, Corsica, Canada, USA.

pubescens Zetterstedt, 1828: 52 ( Omalium ; Type locality: Lapponia: Turtola; Raresuando).

— Zetterstedt, 1838: 52 ( Omalium ; characters; habitat; Lapland).

— Erichson, 1840: 889 ( Omalium ; characters; Lapland).

— Fauvel, 1871a: 111 [= 1872: 85] ( Porrhodites ; synonym of fenestralis ).

— Seidlitz, 1875: 239 ( Porrhodites ; characters; Baltic region).

— Ganglbauer, 1895: 726 ( Porrhodites ; synonym of fenestralis ).

— Bernhauer and Schubert, 1910: 62 ( Porrhodites ; synonym of fenestralis ).

— Campbell, 1984a: 1230 ( Porrhodites ; synonym of fenestralis ).

brevicollis Mäklin, 1852: 320 ( Arpedium ; Type locality: insula Sitkha).

— Mäklin, 1853: 196 ( Deliphrum ; characters; Kenai Peninsula).

— Redtenbacher, 1857: 992 ( Porrhodites ; characters; Austria).

— Kraatz, 1857d: 963 ( Porrhodites ; characters; Germany). — Redtenbacher, 1874: 271 ( Porrhodites ; characters).

— J. Sahlberg, 1876: 214 ( Porrhodites ; synonym of fenestralis ).

— Fauvel, 1878d: 223 [= 1878e: 59] ( Porrhodites ; synonym of fenestralis ).

— Ganglbauer, 1895: 726 ( Porrhodites ; synonym of fenestralis ).

— Bernhauer and Schubert, 1910: 62 ( Porrhodites ; synonym of fenestralis ).

— Scheerpeltz, 1933: 1053 ( Porrhodites ; synonym of fenestralis ).

— Campbell, 1984a: 1230 ( Porrhodites ; synonym of fenestralis ).

— Zanetti, 1987: 267 ( Porrhodites ; synonym of fenestralis ).

inflatus Hatch, 1957: 68 ( Paradeliphrum ; Type locality: B. C. [ British Columbia ]: Copper Mt.).

— Campbell, 1984a: 1232 ( Porrhodites ; characters; Alberta; British Columbia ; Ontario; Quebec; California; Colorado; New Hampshire; New York; Oregon).

— Levesque and Levesque, 1996: 290 ( Porrhodites ; seasonal abundance; Quebec).

— Downie and Arnett, 1996: 434 ( Porrhodites ; characters; Canada; USA).

- DISTRIBUTION: Canada, USA.

pubescens Zetterstedt, 1828 , see: fenestralis Zetterstedt, 1828 .

Tanyrhinus

[1 species; Nearctic Region]

Tanyrhinus Mannerheim, 1852: 349 (species included: singularis ). TYPE SPECIES: Tanyrhinus singularis Mannerheim , fixed by monotypy.

— Gemminger and Harold, 1870: 2062 (cited as Tanyrrhinus ; catalog).

— Fauvel, 1878d: 230 [= 1878e: 66] (cited as Tanyrrhinus ; characters).

— Duvivier, 1883: 197 (catalog; cited as Tanyrrhinus ).

— Bernhauer and Schubert, 1910: 37 (world catalog; 1 species).

— Leng, 1920: 93 (catalog of North American species).

— Scheerpeltz, 1933: 1032 (world catalog supplement).

— Blackwelder, 1952: 374 (type species: singularis ).

— Hatch, 1957: 50, 68 (characters; species of Pacific Northwest).

— Moore, 1966: 48, 56 (cited as Tanyrrhinus ; characters; notes).

— Moore and Legner, 1975: 216 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 195 (characters; notes).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (1 Nearctic species; notes; characters in key). singularis Mannerheim, 1852: 350 ( Tanyrhinus ; Type locality: insula Sitkha).

— Gemminger and Harold, 1870: 2062 ( Tanyrhinus ; cited as Tanyrrhinus ; catalog).

— Fauvel, 1878d: 230 [= 1878e: 66] ( Tanyrhinus ; cited as Tanyrrhinus ; characters; Alaska).

— Bernhauer and Schubert, 1910: 37 ( Tanyrhinus ; catalog).

— Scheerpeltz, 1933: 1032 ( Tanyrhinus ; catalog).

— Hatch, 1957: 69 ( Tanyrhinus ; characters; British Columbia ; Washington; Oregon).

— Campbell and Davies, 1991: 91 ( Tanyrhinus ; checklist of species; Alaska and Canada).

- DISTRIBUTION: Canada, USA.

Trichodromeus

[7 species; Palaearctic Region]

Trichodromeus Luze, 1903: 116 (subgenus of Geodromicus ; species included: penicillatus , tumidus ). TYPE SPECIES: Hygrogeus penicillatus Reitter , fixed by subsequent designation by Blackwelder, 1952: 305.

— Bernhauer and Schubert, 1910: 75 (subgenus of Geodromicus ).

— Tikhomirova, 1973a: 141 (subgenus of Geodromicus ; checklist of species of USSR).

— Bordoni, 1984a: 57 (valid genus).

— Bordoni, 1990a: 100 (valid genus; characters).

— Zerche, 1992a: 126, 134 (characters; valid genus; key to some species; phylogeny of species; character polarity; phylogenetic relationship of genus).

Neogeodromicus Bordoni, 1985: 373 (species included: ketmeniensis ). TYPE SPECIES: Neogeodromicus ketmeniensis Bordoni , fixed by monotypy.

— Zerche, 1992a: 126 (synonym of Trichodromeus ).

armatus Cameron, 1941: 144 ( Hygrogeus , cited as Hygrogoeus ; Type locality: Kashmir: Aphar Wat, altitude 13,000 feet).

— Zerche, 1992a: 134 ( Trichodromeus ; Pakistan).

- DISTRIBUTION: India, Pakistan.

brevisulcus Champion, 1925 , see: kumaonensis Champion, 1925 .

flavipes Motschulsky, 1860: 551 ( Geodromus ; Type locality: Alpes du Caucase en Touchétie; [Note: See lectotype designation by Bordoni, 1993]).

— Gemminger and Harold, 1868: 658 ( Anthophagus ; synonym of globulicollis ).

— Fauvel, 1869: 493 ( Anthophagus ; synonym of globulicollis ).

— Fauvel, 1871a: 135 [= 1872: 109] ( Geodromicus ; synonym of globulicollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ).

— Bordoni, 1993: 158 ( Neogeodromicus ; lectotype designation; cited as valid species; characters).

- DISTRIBUTION: Georgia.

ketmeniensis Bordoni, 1985: 374 ( Neogeodromicus ; Type locality: Turkestan: Ketmen-tjube, Sussamir Tan).

— Zerche, 1992a: 128 ( Trichodromeus ; characters; Kirghizia; Turkestan; Kazakhstan; Turkmenia; Transcaspian region).

- DISTRIBUTION: Kyrgyzstan, Kazakhstan, Turkmenistan.

kumaonensis Champion, 1925: 105 ( Hygrogeus ; Type locality: Shelshel, alt. 15,750 ft.).

— Cameron, 1930: 166 ( Hygrogeus ; characters; India).

— Scheerpeltz, 1933: 1067 ( Hygrogeus ; catalog).

— Coiffait, 1983: 275 ( Hygrogeus ; Kashmir).

— Zerche, 1992a: 133 ( Trichodromeus ; Pakistan).

- DISTRIBUTION: India, Pakistan.

brevisulcus Champion, 1925: 106 ( Hygrogeus ; variety of kumaonensis ; Type locality: N. Kumaon: Laptel, alt. 15,000 ft.).

— Cameron, 1930: 166 ( Hygrogeus ; variety of kumaonensis ; characters; India).

— Scheerpeltz, 1933: 1067 ( Hygrogeus ; variety of kumaonensis ).

penicillatus Reitter, 1900c: 49 ( Hygrogeus , cited as Hygrogaeus ; Type locality: Buchara: Karatak; [Note: See lectotype designation by Zerche, 1992a]).

— Luze, 1903: 116 ( Geodromicus ; subgenus Trichodromeus ; characters; Buchara; Turkestan).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Tikhomirova, 1973a: 141 ( Geodromicus ; subgenus Trichodromeus ; checklist; Central Asia).

— Bordoni, 1984a: 57 ( Trichodromeus ).

— Bordoni, 1990a: 100 ( Trichodromeus ; characters; notes; Turkestan).

— Zerche, 1992a: 131 ( Trichodromeus ; lectotype designation; characters).

- DISTRIBUTION: Uzbekistan.

tumidus Luze, 1903: 117 ( Geodromicus ; subgenus Trichodromeus ; Type locality: Russisches Zentralasien: Buchara: Karatagh).

— Luze, 1904: 78 ( Geodromicus ; Turkestan).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Tikhomirova, 1973a: 141 ( Geodromicus ; subgenus Trichodromeus ; checklist; Central Asia).

— Bordoni, 1984a: 57 ( Trichodromeus ).

— Bordoni, 1990a: 102 ( Trichodromeus ; characters; notes).

— Zerche, 1992a: 131 ( Trichodromeus ; synonym of penicillatus ).

sogdianus Bordoni, 1990 a: 104 ( Trichodromeus ; Type locality: Bukara: “Mts. Karateghin, Baldschuan, 924 m).

— Zerche, 1992a: 131 ( Trichodromeus ; synonym of penicillatus ).

schmidti Zerche, 1992 a: 130 ( Trichodromeus ; Type locality: Asia centr., Tadsch., Dschirgatalski Raj., Sukran-Schlucht, 2590 m / Burs-Gletscher-Fuss).

- DISTRIBUTION: Tajikistan.

sogdianus Bordoni, 1990 , see: penicillatus Reitter, 1900 .

transversus Motschulsky, 1860: 550 ( Geodromus ; Type locality: Alpes du Caucase en Touschéte; [Note: See lectotype designation by Bordoni, 1993]).

— Fauvel, 1871a: 136 [= 1872: 110] ( Geodromicus ; synonym of globulicollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ).

— Bordoni, 1993: 158 ( Neogeodromicus ; lectotype designation; cited as valid species; characters). - DISTRIBUTION: Caucasus.

tumidus Luze, 1903 , see: penicillatus Reitter, 1900 .

Trigonodemus

[11 species; Nearctic and Palaearctic Regions]

Trigonodemus LeConte, 1863 a: 56 (species included: striatus ). TYPE SPECIES: Trigonodemus striatus LeConte , fixed by monotypy.

— LeConte, 1863: 26 ([nomen nudum]; list of North American species).

— Gemminger and Harold, 1868: 663 (catalog; cited as Trigonodesmus ).

— Fauvel, 1878d: 231 [= 1878e: 67] (characters).

— Bernhauer and Schubert, 1910: 37 (world catalog; 1 species).

— Leng, 1920: 93 (catalog of North American species).

— Scheerpeltz, 1933: 1032 (world catalog supplement).

— Blackwelder, 1952: 396 (type species: striatus ).

— Hatch, 1957: 50, 68 (characters; species of Pacific Northwest).

— Moore, 1966: 48, 56 (characters; notes).

— Tikhomirova, 1973a: 137 (checklist of species of USSR).

— Moore and Legner, 1975: 217 (catalog; North America north of Mexico).

— Shibata, 1976: 97 (checklist of species of Japan).

— Moore and Legner, 1979: 196 (characters; notes).

— Watanabe, 1990: 250 (characters; species of Japan).

— Campbell and Davies, 1991: 91 (checklist of species; Alaska and Canada).

— Smetana, 1996d: 2 (review; characters; notes; key to species).

— Smetana, 1996h: 245 (key to species).

— Downie and Arnett, 1996: 434 (characters; species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (2 Nearctic species; notes; characters in key).

Arimimelus Kraatz, 1877: 104 (species included: lebioides ). TYPE SPECIES: Arimimelus lebioides Kraatz , fixed by monotypy.

— Sharp, 1889: 473 (synonym of Trigonodemus by citation of lebioides in Trigonodemus ).

— Bernhauer and Schubert, 1910: 37 (synonym of Trigonodemus ).

— Blackwelder, 1952: 61 (type species: lebioides ).

— Watanabe, 1990: 250 (synonym of Trigonodemus ).

— Smetana, 1996d: 2 (synonym of Trigonodemus ).

— Smetana, 1996h: 245 (key to species).

— Smetana, 2000: 301 (key to species).

Klapperichianellia Hlisnikovsky´, 1962: 455, 456 (species included: mirabilis ). TYPE SPECIES: Klapperichianellia mirabilis Hlisnikovsky´, fixed by original designation and monotypy.

— Thayer, 1987: 389 (may be synonym of Trigonodemus ).

— Smetana, 1996d: 2 (synonym of Trigonodemus ).

audax Smetana, 1996d: 11 ( Trigonodemus ; Type locality: Taiwan: Nantou Hsien, Yushan N.P., S.W. slope blw. Yushan Mn. Pk. 3650 m).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

fasciatus Leech, 1939: 258 ( Trigonodemus ; Type locality: Vancouver, B. C. [ British Columbia ]).

— Hatch, 1957: 68 ( Trigonodemus ; characters; British Columbia ; Oregon).

— Smetana, 1996d: 6 ( Trigonodemus ; characters; British Columbia ).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 301 ( Trigonodemus ; characters in key).

- DISTRIBUTION: Canada, USA.

fungicola Smetana, 1996d: 9 ( Trigonodemus ; Type locality: China: Sichuan, Gongga Shan, above Camp 3, 3050 m).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

lebioides Kraatz, 1877: 105 ( Arimimelus ; Type locality: Hagi).

— Sharp, 1889: 473 ( Trigonodemus ; notes; Japan).

— Scheerpeltz, 1933: 1032 ( Trigonodemus ; catalog).

— Nakane, 1963a: 82 ( Trigonodemus ; characters; habitus photograph; Japan).

— Watanabe, 1990: 253 ( Trigonodemus ; characters; Japan).

— J. Li, 1993: 18 ( Trigonodemus ; China).

— Smetana, 1996d: 8 ( Trigonodemus ; characters; Japan).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: Japan, China.

modestus Smetana, 2000: 300 ( Trigonodemus ; Type locality: China: W. Sichuan: Ganzi Tibet Aut. Pref., Kangding Co., Daxue Shan, Mu Ge Cuo , ob. See, 15 km NW Kangding, 3700 m, 30°09N, 101°52E).

- DISTRIBUTION: China.

montanus Smetana, 1996 h: 241 ( Trigonodemus ; Type locality: China: N. Yunnan, Xue Shan nr. Zhongdian , 4000–4100 m).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

mirabilis Hlisnikovsky´, 1962: 458 ( Klapperichianellia ; Type locality: China: Fukien , Kuatun, 2300 m, 27.40 nördl. Breite, 114.40 östl Länge) .

— Schawaller, 1978: 176, 181, 182, 183 ( Klapperichianellia ; notes; habitus outline; type locality). — Smetana, 1996d: 13 ( Trigonodemus ; characters; China).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

pictus Smetana, 2000: 295 ( Trigonodemus ; Type locality: China: NW Yunnan, Bai Ma Xue Shan, 35 km S. Dêqên, 4300–4800).

- DISTRIBUTION: China.

puncticollis Smetana, 2000: 298 ( Trigonodemus ; Type locality: China: Sichuan, sud Barkam ). - DISTRIBUTION: China.

schuelkei Smetana, 1996 h: 244 ( Trigonodemus ; Type locality: China: Shaanxi, Qin Ling Shan, 107.56E, 33.45 N, Autoroute   GoogleMaps km 93 S of Zhouzhi, 108 km SW Xian Mountain Forest, 1650 m).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

striatus LeConte, 1863 a: 57 ( Trigonodemus ; Type locality: Southern States).

— Fauvel, 1878d: 232 [= 1878e: 68] ( Trigonodemus ; characters; Canada; West Virginia).

— Bernhauer and Schubert, 1910: 37 ( Trigonodemus ; catalog).

— Smetana, 1996d: 5 ( Trigonodemus ; characters; Nova Scotia ; Ontario; Quebec; New Hampshire; New York).

— Downie and Arnett, 1996: 434 ( Trigonodemus ; characters; USA).

— Smetana, 2000: 301 ( Trigonodemus ; characters in key).

- DISTRIBUTION: Canada, USA.

Unamis

[5 species; Nearctic Region]

Unamis Casey, 1894: 398 , 400 (species included: truncata ). TYPE SPECIES: Lesteva truncata Casey , fixed by monotypy.

— Bernhauer and Schubert, 1910: 73 (world catalog; 1 species).

— Leng, 1920: 95 (catalog of North American species).

— Blackwelder, 1952: 402 (type species: truncata ).

— Hatch, 1957: 49, 56 (characters; key to species of Pacific Northwest).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1972: 21 (characters; key to species).

— Moore and Legner, 1975: 218 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 198 (characters; notes).

— Campbell and Davies, 1991: 90 (checklist of species; Alaska and Canada).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (5 Nearctic species; notes; characters in key). bjorkmanae Hatch, 1957: 57 ( Unamis ; Type locality: Crater L., Oregon).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: USA.

columbiensis Hatch, 1957: 56 ( Unamis ; Type locality: Sanca, B. C. [ British Columbia ]).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: Canada, USA.

fulvipes Fall, 1922: 12 ( Unamis ; Type locality: California: Humboldt County: Korbel ).

— Hatch, 1957: 57 ( Unamis ; characters; British Columbia ).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: Canada, USA.

giulianii Moore and Legner, 1972: 21 ( Unamis ; Type locality: California: Inyo County, near Finger Lake , 11,300 feet).

- DISTRIBUTION: USA.

truncata Casey, 1885 a: 322 ( Lesteva ; Type locality: California: Middle Sierras ).

— Casey, 1894: 400 ( Unamis ).

— Bernhauer and Schubert, 1910: 73 ( Unamis ; catalog).

— Fall, 1922: 13 ( Unamis ; Colorado specimen provisionally referred to genus).

— Hatch, 1957: 56 ( Unamis ; characters; Idaho; Washington; Oregon).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: USA.

Uenohadesina

[1 species; Palaearctic Region]

Uenohadesina Smetana, 2000 a: 285 (species included: styx ). TYPE SPECIES: Uenohadesina styx Smetana , fixed by original designation and monotypy.

styx Smetana, 2000 a: 291 ( Uenohadesina ; Type locality: South Korea: Yong’yeon-gul Cave nr. Hwangji , Changseong-eub, Kangweon-do).

- DISTRIBUTION: South Korea.

Vellica

[1 species; Nearctic Region]

Vellica Casey, 1885 a: 321 (species included: longipennis ). TYPE SPECIES: Vellica longipennis Casey , fixed by monotypy.

— Casey, 1894: 401 (notes).

— Leng, 1920: 95 (catalog of North American species).

— Blackwelder, 1952: 403 (type species: longipennis ).

— Hatch, 1957: 49, 61 (characters; species of Pacific Northwest).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1975: 219 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 199 (characters; notes).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (1 Nearctic species; notes; characters in key). longipennis Casey, 1885 a: 321 ( Vellica ; Type locality: California: Middle Sierras).

— Bernhauer and Schubert, 1910: 74 ( Vellica ; catalog).

— Hatch, 1957: 61 ( Vellica ; characters; British Columbia ).

— Campbell and Davies, 1991: 90 ( Vellica ; checklist of species; Canada).

- DISTRIBUTION: Canada, USA.

Xenicopoda

[1 species; Nearctic Region]

Xenicopoda Moore and Legner, 1971: 51 (species included: helenae ). TYPE SPECIES: Xenicopoda helenae Moore and Legner , fixed by original designation and monotypy.

— Moore and Legner, 1975: 220 (catalog; North America north of Mexico).

— Thayer, 1978: 146 (characters; notes; discussion).

— Moore and Legner, 1979: 200 (characters; notes).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (1 Nearctic species; notes; characters in key). helenae Moore and Legner, 1971: 53 ( Xenicopoda ; Type locality: Mt. Wilson, Calif.).

— Thayer, 1978: 143 ( Xenicopoda ; characters; notes; California).

- DISTRIBUTION: USA.

Tribe Aphaenostemmini

Aphaenostemmini Peyerimhoff, 1914: 248 (genera included: Aphaenostemmus ). TYPE GENUS: Aphaenostemmus (see Newton and Thayer, 1992: 55).

— Scheerpeltz, 1933: 1031 (world catalog supplement).

— Peyerimhoff, 1939: 212 (characters in key).

— Scheerpeltz, 1940: 14 (key to Palaearctic genera).

— Newton and Thayer, 1995: 247–286, 295 (phylogenetic relationships; characters; discussion; tribe of Omaliinae ; includes Aphaenostemmus , Giulianium ).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (characters; notes).

Aphaenostemminae Peyerimhoff, 1914.

— Newton and Thayer, 1992: 55 (summary of current classification; notes).

Aphaenostemmus

[5 species; Oriental and Palaearctic Regions]

Aphaenostemmus Peyerimhoff, 1914: 245 (species included: bordei ). TYPE SPECIES: Aphaenostemmus bordei Peyerimhoff , fixed by monotypy.

— Winkler, 1925: 326 (catalog for Palaearctic region).

— Scheerpeltz, 1933: 1031 (world catalog supplement).

— Normand, 1935: 357 (list of species of Tunisia).

— C. Koch, 1936c: 332 (key to species).

— Peyerimhoff, 1939: 211 (characters; key to North African species).

— Blackwelder, 1952: 58 (type species: bordei ).

— Kocher, 1958: 92 (checklist of species; Morocco).

— Newton and Thayer, 1995: 296 (list of synonyms).

Torretassoella C. Koch, 1936b: 126 (species included: testacea ). TYPE SPECIES: Torretassoella testacea C. Koch , fixed by monotypy.

— C. Koch, 1936c: 332 (synonym of Aphaenostemmus ).

— Peyerimhoff, 1939: 211 (synonym of Aphaenostemmus ).

Paratriga Cameron, 1940c: 209 (species included: indica , associata ). TYPE SPECIES: Paratriga indica Cameron , fixed by original designation.

— Blackwelder, 1952: 294 (type species: indica ).

— Newton and Thayer, 1995: 296 (synonym of Aphaenostemmus ).

associatus Cameron, 1940c: 209 ( Paratriga ; Type locality: United Provinces: Dehra Dun).

— Newton and Thayer, 1995: 296 ( Aphaenostemmus ).

- DISTRIBUTION: India.

bordei Peyerimhoff, 1914: 248 ( Aphaenostemmus ; Type locality: Biskra).

— Scheerpeltz, 1933: 1031 ( Aphaenostemmus ; catalog).

— C. Koch, 1936c: 332 ( Aphaenostemmus ; characters; Algeria).

— Peyerimhoff, 1939: 213 ( Aphaenostemmus ; characters in key).

- DISTRIBUTION: Algeria.

himalayicus Bernhauer, 1943 a: 170 ( Aphaenostemmus ; Type locality: Dehra-Dun: Bhagwantung, Himalaya).

- DISTRIBUTION: India.

indicus Cameron, 1940c: 209 ( Paratriga ; Type locality: United Provinces: Dehra Dun).

— Newton and Thayer, 1995: 296 ( Aphaenostemmus ).

- DISTRIBUTION: India.

testaceus C. Koch, 1936b: 126 View in CoL , 133 (Torretassoella; Type locality: Aegypten: Wadi Digla. Sinai: Wadi Isla; Wadi Luotaie ).

— C. Koch, 1936c: 332 ( Aphaenostemmus View in CoL ; characters; Egypt).

— Peyerimhoff, 1939: 213 ( Aphaenostemmus , transferred from Torretassoella; characters in key).

— Coiffait, 1979a: 163, 179 ( Aphaenostemmus ; illustrations of forebody and aedeagus; Saudi Arabia).

— Gusarov, 1997: 278 ( Aphaenostemmus ; Oman).

- DISTRIBUTION: Egypt, Saudia Arabia, Oman.

Giulianium

[3 species; Nearctic and Palaearctic Regions]

Giulianium Moore, 1976: 56 (species included: campbelli ). TYPE SPECIES: Giulianium campbelli Moore , fixed by original designation and monotypy.

— Moore and Legner, 1979: 256 (characters; notes).

— Smetana and Campbell, 1980: 1069 (characters).

— Newton and Thayer, 1995: 295 ( Aphaenostemmini ).

— Ahn and Ashe, 1999: 159 (characters; key to species).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (3 Nearctic species; notes; characters in key).

alaskanum Ahn and Ashe, 1999: 162 ( Giulianium ; Type locality: USA: Alaska: Seward. Also cited from Japan).

- DISTRIBUTION: USA, Japan.

campbelli Moore, 1976: 57 ( Giulianium ; Type locality: California: Humboldt County: Bear Harbor ).

— Ahn and Ashe, 1999: 162 ( Giulianium ; characters; USA).

- DISTRIBUTION: USA.

newtoni Ahn and Ashe, 1999: 163 ( Giulianium ; Type locality: USA: California: San Mateo Co., Moss Beach ).

- DISTRIBUTION: USA.

Tribe Corneolabiini

Corneolabiini Steel, 1950: 54 (including key to genera; genera included: Corneolabium , Paracorneolabium , Metacorneolabium ). TYPE GENUS: Corneolabium , by original designation.

— Thayer, 1985: 115 (characters; notes).

— Newton and Thayer, 1992: 58 (tribe of Omaliinae ).

— Newton and Thayer, 1995: 247–286, 295 (phylogenetic discussion; characters; discussion; includes Corneolabium , Paracorneolabium ; Metacorneolabium ).

— Klimaszewski, Newton, and Thayer, 1996: 146 (list of New Zealand genera; number of New Zealand species).

Corneolabium

[1 species; Australian Region]

Corneolabium Steel, 1950: 54 (species included: mandibulare ). TYPE SPECIES: Corneolabium mandibulare Steel , fixed by original designation and monotypy.

— Thayer, 1985: 115, 163 (characters; phylogenetic relationships).

mandibulare Steel, 1950: 56 ( Corneolabium ; Type locality: New Zealand: West Plains ).

— Klimaszewski, Newton, and Thayer, 1996: 146 ( Corneolabium ; endemic to New Zealand).

- DISTRIBUTION: New Zealand.

Metacorneolabium

[22 species; Neotropical and Australian Regions]

Metacorneolabium Steel, 1950: 62 (species included: minus , cited as minor ). TYPE SPECIES: Metacorneolabium minus Steel , fixed by original designation and monotypy.

— Thayer, 1985: 116, 122, 135, 144, 163 (characters; revision of and key to species; habitats; phylogeny; biogeography).

Metomalium Coiffait and Saiz, 1968: 449 (species included: chilensis ). TYPE SPECIES: Metomalium chilense Coiffait and Saiz , fixed by original designation and monotypy. — Thayer, 1985: 116 (synonym of Metacorneolabium ).

alulatum Thayer, 1985: 137 ( Metacorneolabium ; Type locality: Australia: Tasmania: W side Lake St. Clair , c. 750 m).

- DISTRIBUTION: Australia.

batyle Thayer, 1985: 155 ( Metacorneolabium ; Type locality: Chile: Concepción Prov. : Ramuncho).

- DISTRIBUTION: Chile.

bipertica Thayer, 1985: 159 ( Metacorneolabium ; Type locality: Chile: Osorno Prov. : Parque Nacional Puyehue, Antillanca road, 965 m).

- DISTRIBUTION: Chile.

chilense Coiffait and Saiz, 1968: 450 ( Metomalium ; cited as chilensis ; Type locality: Chili: Nahuelbuta, Malleco).

— Thayer, 1985: 148 ( Metacorneolabium ; emended to chilense ; characters; Chile).

- DISTRIBUTION: Chile.

convexum Thayer, 1985: 123 ( Metacorneolabium ; Type locality: New Zealand: NC: 8.0 km S [town of] Arthur’s Pass , 670 m).

- DISTRIBUTION: New Zealand.

dactylotum Thayer, 1985: 154 ( Metacorneolabium ; Type locality: Chile: Osorno Prov. : Parque Nacional Puyehue, Antillanca road, 965 m).

- DISTRIBUTION: Chile.

darlingtoni Thayer, 1985: 138 ( Metacorneolabium ; Type locality: Australia: Tasmania: Lower Gordon River   GoogleMaps , 42.31S, 145.45E – 42.32S, 145.47E 7).

- DISTRIBUTION: Australia.

diffusum Thayer, 1985: 156 ( Metacorneolabium ; Type locality: Chile: Llanquihue Prov. : Lago Chapo, 13. 5 km E Correntoso, 310 m).

- DISTRIBUTION: Chile.

echidna Thayer, 1985: 142 ( Metacorneolabium ; Type locality: Australia: Victoria: Mt. Buf-

falo, 500 m).

exochlum Thayer, 1985: 158 (Metacorneolabium; Type locality: Chile: Cautín Prov.: Volcán Villarrica, 1250 m).

- DISTRIBUTION: Chile.

exuberatum Thayer, 1985: 145 (Metacorneolabium; Type locality: Chile: Cautín Prov.: Volcán Villarrica, 1250 m).

- DISTRIBUTION: Chile.

gigas Thayer, 1985: 124 (Metacorneolabium; Type locality: New Zealand: BR: Mt. Murchison, 1420 m).

- DISTRIBUTION: New Zealand.

hokitika Thayer, 1985: 132 (Metacorneolabium; Type locality: New Zealand: WD: Hokitika, Glow Worm Dell, 5 m).

- DISTRIBUTION: New Zealand.

logor Thayer GoogleMaps , 1985: 143 ( Metacorneolabium   GoogleMaps ; Type locality: Australia   GoogleMaps : Tasmania: Lower Gordon River, 42.42S, 145.53E- 42.41.5S, 145.53E).

- DISTRIBUTION: Australia.

minus Steel, 1950: 64 (Metacorneolabium; cited as minor; Type locality: New Zealand: Mount Hutt).

— Thayer, 1985: 130 (Metacorneolabium; emended from minor to minus; characters; New Zealand).

— Emberson, 1998: 32 (Metacorneolabium; collecting notes; Chatham Islands).

- DISTRIBUTION: New Zealand, Chatham Islands.

pilatum Thayer, 1985: 153 (Metacorneolabium; Type locality: Chile: Osorno Prov.: Parque Nacional Puyehue, Volcán Casa Blanca, 1370 m).

- DISTRIBUTION: Chile.

pyriforme Thayer, 1985: 125 (Metacorneolabium; Type locality: New Zealand: BR: Nelson Lakes N. P., Lake Rotoiti, 600 m).

- DISTRIBUTION: New Zealand.

rangipo Thayer, 1985: 133 (Metacorneolabium; Type locality: New Zealand: TO: Rangipo Intake, 36 road km S Turangi, 900 m).

- DISTRIBUTION: New Zealand.

rivalis Thayer, 1985: 147 (Metacorneolabium; Type locality: Chile: Osorno Prov.: 7. 7 km NE Termas de Puyehue, 200 m).

- DISTRIBUTION: Chile.

soror Thayer, 1985: 161 (Metacorneolabium; Type locality: Chile: Malleco Prov.: 12 km E Malalcahuello, 1350 m).

- DISTRIBUTION: Chile.

victas Thayer, 1985: 135 (Metacorneolabium; Type locality: Australia: Tasmania: Rufus Canal, 13. 5 km WNW Derwent Bridge, 800 m).

- DISTRIBUTION: Australia.

zanotium Thayer, 1985: 129 (Metacorneolabium; Type locality: New Zealand: SL: Owaka, Glenomaru Reserve).

- DISTRIBUTION: New Zealand.

Paracorneolabium

[1 species; Australian Region]

Paracorneolabium Steel, 1950: 59 (species included: brouni ). TYPE SPECIES: Paracorneolabium brouni Steel , fixed by original designation and monotypy.

— Thayer, 1985: 115-116, 162-171 (characters; phylogenetic relationships).

brouni Steel, 1950: 60 ( Paracorneolabium ; Type locality: New Zealand: Hollyd.; according to Steel this is probably an abbreviation for Hollyford).

Tribe Coryphiini

Coryphiini Jakobson, 1908 .

— Portevin, 1929: 430 (key to French genera).

— Joy, 1932: 97 (key to British genera).

— Hatch, 1957: 49 (characters; key to genera of Pacific Northwest).

— Arnett, 1963: 241, 259 (key to North American genera; characters; list of North American genera).

— Campbell, 1978a: 22 (characters; key to genera and species; Eudectus , Haida , Pseudohaida excluded; biological notes).

— Moore and Legner, 1979: 161 (characters; tabular key to genera of North America north of Mexico).

— Muona, 1979: 18 (list of Scandinavian species).

— Outerelo and Gamarra, 1985: 82 (key to genera of Iberian Peninsula).

— Watanabe, 1990: 264, 377 (characters; key to genera of Japan; relationships of genera).

— Zerche, 1989: 27 (supplement to Lohse, 1964).

— Zerche, 1990 (monograph; phylogeny; characters; checklist of and keys to taxa of Palaearctic region; natural history notes; systematic position and taxonomic status of some Nearctic taxa changed).

— Newton and Thayer, 1992: 58 (tribe of Omaliinae ).

— Zerche, 1993: 352, 354–357 ( Geodromicus, Liophilydrodes and Philydrodes placed in Hygrogeus group of Anthophagini ; bionomics; Zoogeography).

— Newton and Thayer, 1995: 247–286, 294 (phylogenetic relationships; characters; discussion).

— M. Hansen, 1996: 95 (list of Danish species).

— Newton, Thayer, Ashe, and Chandler, 2000: 285 (key to Nearctic genera).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (characters; notes).

Subtribe Boreaphilina

Boreaphilina Zerche, 1990: 22 , 158 (genera included: Archaeoboreaphilus , Caloboreaphilus , Boreaphilus , Niphetodes, Hypsonothrus ). TYPE GENUS: Boreaphilus (see Newton and Thayer, 1992: 58).

— Newton and Thayer, 1992: 58 (subtribe of Coryphiini ).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (characters; notes).

Boréaphilaires Mulsant and Rey, 1880: 391 (genera included: Coryphium , Eudectus , Boreaphilus ). TYPE GENUS: Boreaphilus (see Newton and Thayer, 1992: 58).

— Newton and Thayer, 1992: 58 (synonym of Boreaphilina ).

Archaeoboreaphilus

[6 species; Palaearctic Region]

Archaeoboreaphilus Zerche, 1990: 158 (species included: nikkoensis , miyamorii , tateoi , fuscus , iwakisanensis , yasutoshii ). TYPE SPECIES: Boreaphilus nikkoensis Sawada , fixed by original designation.

fuscus Watanabe, 1988: 53 ( Coryphium ; Type locality: Japan: Honshu: Yamaguchi Pref.: Azami-ga-take, Kanochô ).

— Watanabe, 1990: 362 ( Coryphium ; characters; Japan).

— Zerche, 1990: 164 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

iwakisanensis Watanabe, 1988: 54 ( Coryphium ; Type locality: Japan: Honshu: Aomori Pref.: Mt. Iwakisan ).

— Watanabe, 1990: 363 ( Coryphium ; characters; Japan).

— Zerche, 1990: 165 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

miyamorii Watanabe, 1988: 49 ( Coryphium ; Type locality: Japan: Hokkaido: Horobetsukôzan, Iburi ).

— Watanabe, 1990: 359 ( Coryphium ; characters; Japan).

nikkoensis Sawada, 1964: 36 ( Boreaphilus ; Type locality: Japan: Honshu: Pref. Tochigi: Chuzenji near Nikko ).

— Watanabe, 1988: 56 ( Coryphium ; characters; Japan).

— Watanabe, 1990: 364 ( Coryphium ; characters; Japan).

— Zerche, 1990: 159 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

tateoi Watanabe, 1988: 52 ( Coryphium ; Type locality: Japan: Honshu: Nara Pref.: Tônomine, Yamato ).

— Watanabe, 1990: 361 ( Coryphium ; characters; Japan).

— Zerche, 1990: 164 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

yasutoshii Watanabe, 1988: 58 ( Coryphium ; Type locality: Japan: Honshu: Gunma Pref.: near Edozawa, Mt. Jôshû-Hotaka ).

— Watanabe, 1990: 366 ( Coryphium ; characters; Japan).

— Zerche, 1990: 166 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

Boreaphilus

[23 species; Nearctic and Palaearctic Regions]

Boreaphilus C. Sahlberg, 1832: 433 (species included: henningianus ). TYPE SPECIES: Boreaphilus henningianus C. Sahlberg , fixed by monotypy.

— Zetterstedt, 1838: 72 (characters).

— Erichson, 1840: 899 (characters).

— Haliday, 1841: 186 (characters).

— Schaum, 1852: 31 (list of species; Europe).

— Lacordaire, 1854: 137 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 632 (characters).

— Kraatz, 1857: 39 (characters).

— Jacquelin du Val, 1857: 67 (characters).

— Redtenbacher, 1857: 245 (characters).

— Thomson, 1858: 37 (characters).

— Thomson, 1859: 49 (characters; type species: henningianus ).

— Schaum, 1859: 32 (catalog; European species).

— Thomson, 1861: 183 (characters).

— Gemminger and Harold, 1868: 664 (catalog).

— Jacquelin du Val, 1868: 79 (catalog; European species).

— Fauvel, 1871a: 106 [= 1872: 80] (characters).

— Redtenbacher, 1874: 273 (characters).

— Fauvel, 1875a:V [= 1875b: 207] (catalog).

— Mulsant and Rey, 1880: 401 (characters).

— Heyden, 1880: 82 (list of species of Siberian region).

— Duvivier, 1883: 197 (catalog).

— Seidlitz, 1891: 349 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 118 (list of species of Europe and Caucasus).

— Heyden, 1893: 48 (list of species of Siberian region).

— Ganglbauer, 1895: 699 (characters; key to middle European species).

— Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 84 (world catalog; 8 species).

— Notman, 1918 (characters; notes).

— Leng, 1920: 95 (catalog of North American species).

— Winkler, 1925: 337 (catalog for Palaearctic region).

— Porta, 1926: 33 (characters; species of Italy).

— Scheerpeltz, 1933: 1072 (world catalog supplement).

— Normand, 1935: 358 (list of species of Tunisia).

— Blackwelder, 1952: 83 (type species: henningianus ).

— Smetana, 1959a: 196 (list of species collected in Albania).

— Horion, 1963: 154 (list of central European species).

— Székessy, 1963: 24 (characters).

— Lohse, 1964: 69 (characters; key to central European species).

— Scheerpeltz, 1968a: 19 (catalog of Austrian species).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Moore and Legner, 1975: 186 (catalog; North America north of Mexico).

— Shibata, 1976: 125 (checklist of species of Japan).

— Campbell, 1978a: 24, 58 (key to species; characters; notes).

— Moore and Legner, 1979: 162 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Tóth, 1982: 107 (characters; habitus; species of Hungary).

— Muona and Viramo, 1986: 16 (list of species of northeastern Finland).

— Zanetti, 1987: 434 (characters; habitus; key to species of Italy).

— Zerche, 1989: 123, 129 (characters; notes; distributional dates discussed).

— Watanabe, 1990: 370 (characters; key to species of Japan).

— Zerche, 1990: 169, 286, 306 (characters; key to species of Palaearctic region; phylogeny; distribution).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Zerche, 1993: 344, 350, 355 (modification of key; phylogeny; distribution; distribution in Nearctic and Palaearctic regions compared).

— Siitonen, 1993: 230 (list of Finnish species).

— Zanetti, 1995: 10 (list of species of Italy).

— Downie and Arnett, 1996: 436 (characters; key to species of Northeastern North America ).

— Kashcheev, 1999: 144 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (1 Nearctic species; notes; characters in key).

Chevrieria Heer, 1839: 188 (species included: velox ). TYPE SPECIES: Chevrieria velox Heer , fixed by monotypy.

— Heer, 1841: 571 (notes).

— Schaum, 1852: 30 (list of species; Europe).

— Lacordaire, 1854: 135 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 631 (characters).

— Jacquelin du Val, 1857: 67 (synonym of Boreaphilus ).

— Redtenbacher, 1857: 241 (characters).

— Redtenbacher, 1857: 992 (synonym of Boreaphilus ).

— Schaum, 1859: 32 (synonym of Boreaphilus ).

— Gemminger and Harold, 1868: 664 (synonym of Boreaphilus ).

— Fauvel, 1871a: 106 [= 1872: 80] (synonym of Boreaphilus ).

— Ganglbauer, 1895: 699 (synonym of Boreaphilus ).

— Bernhauer and Schubert, 1910: 84 (synonym of Boreaphilus ).

— Scheerpeltz, 1933: 1072 (synonym of Boreaphilus ).

— Blackwelder, 1952: 99 (type species: velox ).

— Campbell, 1978a: 59 (synonym of Boreaphilus ).

— Zanetti, 1987: 434 (synonym of Boreaphilus ).

— Zerche, 1990: 169 (synonym of Boreaphilus ).

Boreophilus Agassiz, 1847: 49 (emendation of Boreaphilus ). TYPE SPECIES: Boreaphilus henningianus C. Sahlberg , fixed by objective synonymy with Boreaphilus .

— Blackwelder, 1952: 84 (type species).

— Campbell, 1978a: 59 (synonym of Boreaphilus ).

Catocopa Gistel, 1856: 29 . TYPE SPECIES: Chevreiria velox Heer , fixed by monotypy.

— Blackwelder, 1952: 96 (synonym of Boreaphilus ; type species: velox ).

— Zerche, 1990: 169 (synonym of Boreaphilus ).

albanicus Zerche, 1990: 197 ( Boreaphilus ; Type locality: Nordalban.: Alpen Casa Valbones).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Philorinum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Loc

Philorinum nitidulum

Herman, Lee H. 2001
2001
Loc

testaceus

C. Koch 1936: 126
1936
Loc

Aphaenostemmus

Peyerimhoff 1914
1914
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