Philorinum nitidulum, Kraatz, 1857 d: 966

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.1

persistent identifier

https://treatment.plazi.org/id/BF684949-423C-B5F7-00DC-2DB4FB577CB2

treatment provided by

Felipe

scientific name

Philorinum nitidulum
status

 

nitidulum Kraatz, 1857d: 966 View in CoL , note ( Philorinum ; Type locality: Griechenland).

— Kraatz, 1858b: 128 ( Philorinum ; characters; Greece).

— Mulsant and Rey, 1880: 158 ( Philorinum ; synonym of humile ).

— Ganglbauer, 1895: 727 ( Philorinum ; synonym of sordidum ).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; synonym of sordidum ).

— Scheerpeltz, 1933: 1052 ( Philorinum ; synonym of sordidum ).

ruficolle Schaufuss, 1861: 47 ( Philorinum ; cited as rufieolle ; Type locality: Hisp. mer.). — Fauvel, 1876a: 51 [= 1876: 236] ( Philorinum ; synonym of sordidum ).

— Mulsant and Rey, 1880: 158 ( Philorinum ; synonym of humile ).

— Ganglbauer, 1895: 727 ( Philorinum ; synonym of sordidum ).

— Fauvel, 1902b: 55 ( Philorinum ; synonym of sordidum ).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; synonym of sordidum ).

cadomense Fauvel, 1863 a: 42 ( Philorinum ; Type locality: Calvados, Ille-et-Vilaine).

— Mulsant and Rey, 1880: 156 ( Philorinum ; synonym of humile ).

— Ganglbauer, 1895: 727 ( Philorinum ; synonym of sordidum ).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; synonym of sordidum ).

brevipenne Luze, 1910: 240 ( Anthobium ; Type locality: Andalusien).

— Scheerpeltz, 1933: 1033 ( Anthobium ; catalog).

— C. Koch, 1940: 372 ( Anthobium ; characters; Spain).

— Zanetti, 1986: 95 ( Philorinum ; synonym of sordidum ).

marocanum Coiffait, 1978 r: 271 ( Eusphalerum ; Type locality: Maroc: Sidi Bettache). — Zanetti, 1991a: 29 ( Philorinum ; synonym of sordidum ).

subpubescens Stephens, 1834 , see: sordidum Stephens, 1834 .

Philhydrodema

[1 species; Palaearctic Region]

Philhydrodema Scheerpeltz, 1976: 11 (species included: longelytratum ). TYPE SPECIES: Philhydrodema longelytratum Scheerpeltz , fixed by original designation.

longelytratum Scheerpeltz, 1976: 12 ( Philhydrodema ; Type locality: Nepal: Nare-Glacier

[= Ama-Dablam-Gletscher] oberhalb der Green Hut der Hillary-Makalu-Expedition, 5570 m).

- DISTRIBUTION: Nepal.

Philydrodes

[26 species; Palaearctic Region]

Philydrodes Bernhauer, 1929 g: 179 (species included: aquatilis ). TYPE SPECIES: Anthophagus aquatilis Sharp , fixed by original designation and monotypy.

— Scheerpeltz, 1933: 1067 (world catalog supplement).

— Blackwelder, 1952: 303 (type species: aquatilis ).

— Nakane and Sawada, 1956a: 179 (characters; key to Japanese species; type species: aquatilis ).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Shibata, 1976: 119 (checklist of species of Japan).

— Watanabe, 1990: 309 (characters; key to species of Japan).

— Kashcheev, 1999: 146 (characters).

Minyphilydrodes Watanabe, 1990: 334 (subgenus of Philydrodes ; species included: watanabei , asamanis , parallelus , maculatus , ishidai , biguttatus , flavidus , shimomurai , chichibunis , niger ). TYPE SPECIES: Philydrodes watanabei R. Dvořák , fixed by original designation.

aquatilis Sharp, 1889: 469 ( Anthophagus ; Type locality: Hakone).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog).

— Bernhauer, 1929g: 180 ( Philydrodes ).

— Scheerpeltz, 1933: 1067 ( Philydrodes ; catalog).

— Nakane, 1963a: 83 ( Philydrodes ; characters; habitus photograph; Japan).

— Watanabe, 1990: 314 ( Philydrodes ; characters; Japan).

— Watanabe, 1996a: 9 ( Philydrodes ; cave collection; Japan).

- DISTRIBUTION: Japan.

japonicus Yoshida and Nomura, 1952: 6 ( Coryphium ; Type locality: Aoiwa L. S. C., Kanto-

Mountainland, Japan).

— Shibata, 1970: 57 (suggested transfer from Coryphium to Philydrodes ).

— Watanabe, 1990: 314 ( Philydrodes ; synonym of aquatilis ).

— Zerche, 1990: 18 ( Philydrodes ). asamanis Watanabe, 1990: 337 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan:

Honshu: Gunma Pref.: near Nidoage).

- DISTRIBUTION: Japan. biguttatus Watanabe, 1990: 343 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality:

Japan: Honshu: Nagano Pref.: Hakuba-mura).

- DISTRIBUTION: Japan. chichibunis Watanabe, 1990: 347 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality:

Japan: Honshu: Saitama Pref.: Kawamata, Chichibu).

- DISTRIBUTION: Japan. dewanus Watanabe, 1990: 325 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan:

Honshu: Yamagata Pref.: Tachiyazawa, Higashitagawa).

- DISTRIBUTION: Japan. flavidus Watanabe, 1990: 344 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan:

Honshu: Yamanashi Pref.: Mt. Kitazawa-tôge, S. Jpn. Alps).

- DISTRIBUTION: Japan. gigas Watanabe, 1990: 317 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu:

Shizuoka Pref.: Umegashima Spa).

- DISTRIBUTION: Japan. grandis Nakane and Sawada, 1956 a: 181 ( Philydrodes ; Type locality: Japan: Kyushu:

Chinoike-jigoku, Beppu).

— Watanabe, 1990: 320 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan. hikosanensis Nakane and Sawada, 1956 a: 180 ( Philydrodes ; Type locality: Japan: Kyushu:

Hikosan).

— Nakane, 1963a: 83 ( Philydrodes ; characters; habitus photograph; Japan).

— Watanabe, 1990: 318 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan. horioi Nakane and Sawada, 1956 a: 180 ( Philydrodes ; Type locality: Japan: Sampuku Pass,

Nagano).

— Watanabe, 1990: 316 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan. ishidai Nakane and Sawada, 1956 a: 182 ( Philydrodes ; Type locality: Kamikochi, Nagano).

— Nakane, 1963a: 83 ( Philydrodes ; characters; habitus photograph; Japan).

— Watanabe, 1990: 342 ( Philydrodes ; subgenus Minyphilydrodes ; characters; Japan).

- DISTRIBUTION: Japan. japonicus Yoshida and Nomura, 1952 , see: aquatilis Sharp, 1889 . kawaseorum Hayashi, 1994 a: 133 ( Philydrodes , cited as Phylidrodes; Type locality: Japan:

Mie: Jirakura).

- DISTRIBUTION: Japan. maculatus Watanabe, 1990: 340 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality:

Japan: Honshu: Yamanashi Pref.: Hirogawara, foot of Mt. Kitadake).

michikoae Watanabe, 1990: 323 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu: Yamagata Pref.: Mt. Atsumidake, Nishitagawa).

- DISTRIBUTION: Japan.

minipunctatus Watanabe, 1990: 321 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu: Nagano Pref.: Kamikôchi).

- DISTRIBUTION: Japan.

niger Watanabe, 1990: 349 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan: Honshu: Yamanashi Pref.: Hirogawara, foot of Mt. Kitadake).

- DISTRIBUTION: Japan.

parallelus Watanabe, 1990: 338 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan: Honshu: Nagano Pref.: Kamikôchi).

- DISTRIBUTION: Japan.

pulla Nakane and Sawada, 1956 , see: Liophilydrodes .

puncticollis Nakane and Sawada, 1956 , see: Liophilydrodes .

rubidus Nakane and Sawada, 1956 a: 180 ( Philydrodes ; variety of aquatilis ; Type locality: Japan: Lake Marunuma, Nikko).

— Watanabe, 1990: 331 ( Philydrodes ; valid species; characters; Japan).

- DISTRIBUTION: Japan.

rufescens Watanabe, 1990: 327 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Shikoku: Ehime Pref.: western ravine of Mt. Shiratsue).

- DISTRIBUTION: Japan.

shimomurai Watanabe, 1990: 346 ( Philydrodes ; subgenus Minyphilydrodes ; Type locality: Japan: Honshu: Mt. Sanpuku-tôge, S. Jpn. Alps).

- DISTRIBUTION: Japan.

similis Nakane and Sawada, 1956 a: 181 ( Philydrodes ; subgenus Philydrodes ; variety of horioi ; Type locality: Japan: Nagano, Sampuku Pass).

— Watanabe, 1990: 330 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

takaoensis Nakane and Sawada, 1956 a: 182 ( Philydrodes ; Type locality: Japan: Takaoyama , Tokyo).

— Watanabe, 1990: 334 ( Philydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

troglophila Nakane and Sawada, 1956 , see: Liophilydrodes .

tsurugisanus Watanabe, 1990: 326 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Shikoku: Tokushima Pref.: Ryôtsurugi-dans on Mt. Tsurugi-san).

- DISTRIBUTION: Japan.

yoshidai Watanabe, 1990: 328 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Shikoku: Tokushima Pref.: Ryôtsurugidani on Mt. Tsurugi-san).

- DISTRIBUTION: Japan.

watanabei R. Dvořák, 1957: 9 ( Philydrodes ; Type locality: Itsukaichi , Nishitama, Tokyo, Japon).

— Watanabe, 1990: 335 ( Philydrodes ; subgenus Minyphilydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

zipanguensis Watanabe, 1990: 333 ( Philydrodes ; subgenus Philydrodes ; Type locality: Japan: Honshu: Nagano Pref.: Kamikôchi).

- DISTRIBUTION: Japan.

Phlaeopterus

[14 species; Nearctic Region]

Phlaeopterus Motschulsky, 1853 a: 78 (species included: fusconiger ). TYPE SPECIES:

— Kraatz, 1857d: 930 (synonym of Lesteva ).

— Gemminger and Harold, 1868: 659 (synonym of Lesteva ).

— Fauvel, 1871a: 127 [= 1872: 101] (synonym of Lesteva ).

— Fauvel, 1875a:VII [= 1875b: 209] (synonym of Lesteva ).

— Bernhauer and Schubert, 1910: 74 (synonym of Tilea ).

— Scheerpeltz, 1933: 1065 (world catalog supplement).

— Blackwelder, 1952: 303 (type species: fusconiger ).

— Hatch, 1957: 49, 57 (characters; key to species of Pacific Northwest).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1975: 208 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 192 (characters; notes).

— Campbell and Davies, 1991: 90 (checklist of species; Alaska and Canada).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (14 Nearctic species; notes; characters in key). Tilea Fauvel, 1878d: 246 [= 1878e: 82] (species included: cavicollis ). TYPE SPECIES: Tilea cavicollis Fauvel , fixed by monotypy.

— Duvivier, 1883: 195 (catalog).

— Casey, 1894: 402 (notes; key to North American species).

— Bernhauer and Schubert, 1910: 74 (world catalog; 7 species).

— Leng, 1920: 95 (catalog of North American species).

— Scheerpeltz, 1933: 1065 (synonym of Phlaeopterus ).

— Blackwelder, 1952: 390 (type species: cavicollis ; synonym of Phlaeopterus ).

brevipennis Casey, 1894: 404 ( Tilea ; Type locality: Wyoming).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 60 ( Phlaeopterus ; characters; British Columbia ; Washington; Oregon; Idaho; Montana).

- DISTRIBUTION: Canada, USA.

cascadiensis Hatch, 1957: 59 ( Phlaeopterus ; Type locality: Washington: Mt. Rainier ,

Tipsoo L.; British Columbia ; Oregon).

- DISTRIBUTION: Canada, USA.

castaneus Casey, 1894: 404 ( Tilea ; Type locality: Colorado).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

- DISTRIBUTION: USA.

cavicollis Fauvel, 1878d: 246 [= 1878e: 82] ( Tilea ; Type locality: Colombie britannique).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 60 ( Phlaeopterus ; characters; British Columbia ; Washington; Idaho).

- DISTRIBUTION: Canada, USA.

filicornis Casey, 1886: 234 ( Phlaeopterus , cited as Phloeopterus ; Type locality: California: Placer Co.).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

- DISTRIBUTION: USA.

frosti Hatch, 1957: 59 ( Phlaeopterus ; Type locality: Washington: Mt. Rainier, below Sluskin Falls. Also cited from British Columbia and Oregon).

- DISTRIBUTION: Canada, USA.

fusconiger Motschulsky, 1853 a: 78 ( Phlaeopterus ; Type locality: Ounalaschka).

— Mäklin, 1853: 193 ( Lesteva ; characters; Unalaska).

— LeConte, 1866: 375 ( Lesteva ; California).

— Fauvel, 1878d: 248 [= 1878e: 84] ( Lesteva ; characters; Alaska).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

- DISTRIBUTION: USA.

houkae Hatch, 1957: 58 ( Phlaeopterus ; Type locality: Washington: Mt. Baker Lodge. Also cited from British Columbia ).

- DISTRIBUTION: Canada, USA.

kootenayensis Hatch, 1957: 58 ( Phlaeopterus ; Type locality: British Columbia : Bull River,

6 mi. from mouth).

- DISTRIBUTION: Canada.

lagrandeuri Hatch, 1957: 58 ( Phlaeopterus ; Type locality: Seattle, Carkeek Park, Washington. Also cited from Oregon and British Columbia ).

- DISTRIBUTION: Canada, USA.

loganensis Hatch, 1957: 60 ( Phlaeopterus ; Type locality: G[lacier] N[ational] P[ark]: Logan Pass, Montana. Also cited from British Columbia and Idaho).

- DISTRIBUTION: Canada, USA.

longipalpus Casey, 1885 a: 318 ( Phlaeopterus , cited as Phloeopterus ; Type locality: California: Middle Sierras).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 60 ( Phlaeopterus ; characters; British Columbia ; Washington; Oregon; Idaho; Utah).

- DISTRIBUTION: Canada, USA.

rufitarsis Casey, 1894: 403 ( Tilea ; Type locality: California: Siskiyou Co.).

— Bernhauer and Schubert, 1910: 74 ( Tilea ; catalog).

— Hatch, 1957: 59 ( Phlaeopterus ; characters; British Columbia ; California; Montana).

- DISTRIBUTION: Canada, USA.

stacesmithi Hatch, 1957: 60 ( Phlaeopterus ; Type locality: British Columbia : Barkerville, Roundtop Mt., 6000 ft.).

- DISTRIBUTION: Canada.

Phyllodrepoidea

[1 species; Palaearctic and Nearctic Regions]

Phyllodrepoidea Ganglbauer, 1895: 724 (species included: crenata ). TYPE SPECIES: Phyllodrepoidea crenata Ganglbauer , fixed by monotypy (see Herman, 2001).

— Luze, 1905c: 247 (characters).

— Heyden, Reitter, and Weise, 1906: 133 (list of species of Europe and Caucasus).

— Reitter, 1909: 188 (characters).

— Bernhauer and Schubert, 1910: 63 (world catalog; 1 species).

— Petri, 1912: 54 (list of species of Siebenbürgen).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Porta, 1926: 24 (characters; species of Italy).

— Roubal, 1930: 312 (catalog; Slovakia).

— Joy, 1932: 102 (key to British species).

— Scheerpeltz, 1933: 1054 (world catalog supplement).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 96 (characters; species of Sweden).

— Tottenham, 1949: 356, 407 (type species: crenata Gravenhorst ; list of British species).

— Blackwelder, 1952: 307 (type species: crenata ).

— Tottenham, 1954: 28 (characters; British species).

— Horion, 1963: 101 (list of central European species).

— Székessy, 1963: 18 (characters).

— Lohse, 1964: 51 (characters; key to central European species).

— Kasule, 1968: 133, 134 (larval characters).

— Scheerpeltz, 1968a: 13 (catalog of Austrian species).

— Steel, 1970a: 17 (larval characters; biological notes).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Pope, 1977: 22 (list of British species).

— Burakowski, Mroczkowski, and Stefańska, 1979: 52 (catalog; Poland).

— Muona, 1979: 18 (list of Scandinavian species).

— Tóth, 1982: 74 (characters; habitus; species of Hungary).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Boháč, 1993: 41 (in list of species of Czech Republic and Slovakia).

— Zanetti, 1995: 8 (list of species of Italy).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (1 undescribed Nearctic species; notes; characters in key).

— Herman, 2001: 8 (type species; discussion).

creatoris Gozis, 1886 , see: crenata Gravenhorst, 1802 .

crenata Ganglbauer, 1895: 724 ( Phyllodrepoidea ; Type locality: Mittel- und Südeuropa).

— Gravenhorst, 1802: 114 ( Omalium ; [Note: Misidentification: Gravenhorst attributed this name to Paykull who attributed it to Fabricius whose species is cited in this catalog in Acidota . All subsequent authors have attributed the species to Gravenhorst]).

— Latreille, 1804: 371 ( Omalium ; characters).

— Gravenhorst, 1806: 207 ( Omalium ; characters).

— C. Sahlberg, 1830: 288 ( Omalium ; characters; Finland).

— Erichson, 1840: 873 ( Deliphrum ; characters; Germany).

— Redtenbacher, 1849: 829 ( Deliphrum ; characters).

— Fairmaire and Laboulbène, 1856: 638 ( Deliphrum ; characters; France).

— Redtenbacher, 1857: 249 ( Deliphrum ; characters; Germany).

— Kraatz, 1857d: 954 ( Deliphrum ; characters; Germany).

— Thomson, 1871: 377 ( Deliphrum ; characters; Sweden).

— Fauvel, 1871a: 122 [= 1872: 96] ( Deliphrum ; characters; France; Britain; Germany; Switzerland; Italy; Austria; Corsica; Sicily).

— Redtenbacher, 1874: 270 ( Deliphrum ; characters).

— Mulsant and Rey, 1880: 112 ( Deliphrum ; characters; France).

— Fowler, 1888: 406 ( Deliphrum ; characters; Britain).

— Ganglbauer, 1895: 724 ( Phyllodrepoidea ; characters; middle and southern Europe).

— Luze, 1905c: 249 ( Phyllodrepoidea ; characters; Hungary; Croatia; Bosnia; France, Italy, Greece; Germany; Switzerland; Corsica; Sicily).

— Sainte-Claire Deville, 1906: 71 ( Phyllodrepoidea ; Corsica).

— Sainte-Claire Deville, 1907: 42 ( Phyllodrepoidea ; characters; France).

— Reitter, 1909: 188 ( Phyllodrepoidea ; characters; Germany).

— Bernhauer and Schubert, 1910: 63 ( Phyllodrepoidea ; catalog).

— Portevin, 1929: 442 ( Phyllodrepoidea ; characters; France).

— Scheerpeltz, 1933: 1054 ( Phyllodrepoidea ; catalog).

— Palm, 1948: 97 ( Phyllodrepoidea ; characters; Sweden; Norway).

— Blackwelder, 1952: 307 ( Phyllodrepoidea ; type species).

— Tottenham, 1954: 28 ( Phyllodrepoidea ; characters; Scotland).

— Székessy, 1963: 17 ( Phyllodrepoidea ; illustration).

— Horion, 1963: 101 ( Phyllodrepoidea ; Scotland; Norway; Sweden; France; Belgium; Switzerland; Corsica; Sicily; Italy; Greece; Croatia; Bosnia Hercegovina; Hungary; Romania; Slovakia; Germany; Austria).

— Lohse, 1964: 51 ( Phyllodrepoidea ; characters; central Europe).

— Kasule, 1968: 134 ( Phyllodrepoidea ; larval characters).

— Scheerpeltz, 1968a: 13 ( Phyllodrepoidea ; catalog; Austria).

— Steel, 1970a: 17 ( Phyllodrepoidea ; larval characters; biological notes).

— Tikhomirova, 1973a: 139 ( Phyllodrepoidea ; checklist; Europe).

— Pope, 1977: 22 ( Phyllodrepoidea ; Britain).

— Muona, 1979: 18 ( Phyllodrepoidea ; Sweden; Norway).

— Burakowski, Mroczkowski, and Stefańska, 1979: 52 ( Phyllodrepoidea ; notes; Poland).

— Hammond, 1980: 137 ( Phyllodrepoidea ; notes; Ireland).

— Tóth, 1982: 74 ( Phyllodrepoidea ; characters; Hungary).

— Crowson, 1982: 125 ( Phyllodrepoidea ; natural history observations; Britain).

— Terlutter, 1984: 26, 41 ( Phyllodrepoidea ; habitat notes; Germany).

— Segers, 1986: 21 ( Phyllodrepoidea ; Belgium).

— Lucht, 1987: 88 ( Phyllodrepoidea ; Germany; Poland; Austria; Switzerland; France; Benelux; Sweden).

— Silfverberg, 1992: 22 ( Phyllodrepoidea ; Sweden; Norway).

— Boháč, 1993: 41 ( Phyllodrepoidea ; Slovakia; Czech Republic).

— Allen, 1994: 247 ( Phyllodrepoidea ; Britain).

— Zanetti, 1995: 8 ( Phyllodrepoidea ; Italy).

— Zanetti and Sabella, 1998: 31 ( Phyllodrepoidea ; notes; Sicily).

— M. Hansen, Pedersen, and Pritzl, 1999: 77 ( Phyllodrepoidea ; characters; notes; Denmark).

— Fowles, Alexander, and Key, 1999: 129 ( Phyllodrepoidea ; saproxylic species; status as endangered or threatened species; Britain).

— Herman, 2001: 8, 21 ( Phyllodrepoidea ; type species of genus; Ganglbauer author species not Gravenhorst who attributed it to Fabricius; nomen protectum).

- DISTRIBUTION: Europe.

creatoris Gozis, 1886: 15 ( Deliphrum ; proposed for Gravenhorst’s misidentification of crenatum ; Type locality: Not cited).

— Bernhauer and Schubert, 1910: 63 ( Phyllodrepoidea ; synonym of crenata ).

— Herman, 2001: 21 ( Phyllodrepoidea ; synonym of crenata ; nomen oblitum).

Porrhodites

[2 species; Nearctic and Palaearctic Regions]

Porrhodites Kraatz, 1857d: 961 (species included: brevicollis ). TYPE SPECIES: Arpedium brevicolle Mäklin , fixed by monotypy.

— Redtenbacher, 1857: 992 (characters).

— Thomson, 1859: 50 (characters; type species: fenestralis ).

— Schaum, 1859: 32 (catalog; European species).

— LeConte, 1861a: 70 (characters).

— Thomson, 1861: 195 (characters).

— LeConte, 1863: 26 (list of North American species).

— Gemminger and Harold, 1868: 663 (catalog).

— Fauvel, 1871a: 111 [= 1872: 85] (characters).

— Redtenbacher, 1874: 271 (characters).

— Fauvel, 1875a:VI [= 1875b: 208] (catalog).

— Heyden, 1880: 82 (list of species of Siberian region).

— Heyden, Reitter, and Weise, 1891: 119 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 726 (characters).

— Heyden, Reitter, and Weise, 1906: 133 (list of species of Europe and Caucasus).

— Reitter, 1909: 184 (characters).

— Bernhauer and Schubert, 1910: 62 (world catalog; 62 species).

— Leng, 1920: 94 (catalog of North American species).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Porta, 1926: 24 (characters; species of Italy).

— Scheerpeltz, 1933: 1053 (world catalog supplement).

— Palm, 1948: 92 (characters; species of Sweden).

— Blackwelder, 1952: 320 (type species: brevicollis ).

— Hatch, 1957: 50, 69 (characters; species of Pacific Northwest).

— Horion, 1963: 98 (list of central European species).

— Székessy, 1963: 16 (characters).

— Lohse, 1964: 49 (characters; key to central European species).

— Moore, 1966: 48, 51 (characters; notes).

— Scheerpeltz, 1968a: 12 (catalog of Austrian species).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Moore and Legner, 1975: 210 (catalog; North America north of Mexico).

— Burakowski, Mroczkowski, and Stefańska, 1979: 52 (catalog; Poland).

— Moore and Legner, 1979: 193 (characters; notes).

— Tóth, 1982: 69 (characters; habitus; key to species of Hungary).

— Campbell, 1984a: 1228 (characters; review of and key to North American species).

— Zanetti, 1987: 267 (characters; habitus; species of Italy).

— Campbell and Davies, 1991: 91 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Downie and Arnett, 1996: 433 (characters; key to species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (2 Nearctic species; notes; characters in key).

brevicollis Mäklin, 1852 , see: fenestralis Zetterstedt, 1828 .

fenestralis Zetterstedt, 1828: 50 ( Omalium ; Type locality: Lapponia Tornensis).

— Zetterstedt, 1838: 50 ( Omalium ; characters; habitat; Lapland).

— Erichson, 1840: 888 ( Omalium ; characters; Lapland).

— Kraatz, 1857d: 963 ( Porrhodites ; synonym of brevicollis ).

— Thomson, 1859: 50 (type species of Porrhodites ).

— Thomson, 1861: 196 ( Porrhodites ; characters; Scandinavia).

— Fauvel, 1871a: 111 [= 1872: 85] ( Porrhodites ; characters; Switzerland; Lapland; Alaska).

— Fauvel, 1874: 323 [= 1874b: 29] ( Porrhodites ; Finland).

— J. Sahlberg, 1876: 214 ( Porrhodites ; Finland).

— Fauvel, 1878d: 223 [= 1878e: 59] ( Porrhodites ; characters; Alaska; British Columbia ; Lake Superior).

— Seidlitz, 1891: 344 ( Porrhodites ; characters; Europe).

— Seidlitz, 1891a: 366 ( Porrhodites ; characters; Europe).

— Ganglbauer, 1895: 726 ( Porrhodites ; characters; Lapland; Finland; Siberia; North America ).

— Poppius, 1909: 5 ( Porrhodites ; Lena Valley, Siberia ).

— Reitter, 1909: 184 ( Porrhodites ; characters; Germany).

— Bernhauer and Schubert, 1910: 62 ( Porrhodites ; catalog).

— Scheerpeltz, 1933: 1053 ( Porrhodites ; catalog).

— Palm, 1948: 93 ( Porrhodites ; characters; Sweden; Norway; Finland).

— Hatch, 1957: 69 ( Porrhodites ; characters; British Columbia ).

— Horion, 1963: 98 ( Porrhodites ; Norway; Sweden; Finland; Russia; Poland; Germany; Czechoslovakia; Austria; Corsica).

— Lohse, 1964: 49 ( Porrhodites ; characters; central Europe).

— Tóth, 1982: 69 ( Porrhodites ; characters).

— Campbell, 1984a: 1230 ( Porrhodites ; characters; Alberta; British Columbia ; Manitoba; Newfoundland; Labrador; Northwest Territories; Quebec; Yukon Territory; Alaska; Montana; Oregon; Wyoming).

— Zanetti, 1987: 267 ( Porrhodites ; characters; Italy).

— Lucht, 1987: 88 ( Porrhodites ; checklist; Austria).

— Boháč, 1993: 41 ( Porrhodites ; Czech Republic).

— Downie and Arnett, 1996: 433 ( Porrhodites ; characters; Canada; USA).

— Ryabukhin, 1999: 20 ( Porrhodites ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Norway, Sweden, Finland, Russia, Poland, Germany, Czechoslovakia, Austria, Italy, Switzerland, Corsica, Canada, USA.

pubescens Zetterstedt, 1828: 52 ( Omalium ; Type locality: Lapponia: Turtola; Raresuando).

— Zetterstedt, 1838: 52 ( Omalium ; characters; habitat; Lapland).

— Erichson, 1840: 889 ( Omalium ; characters; Lapland).

— Fauvel, 1871a: 111 [= 1872: 85] ( Porrhodites ; synonym of fenestralis ).

— Seidlitz, 1875: 239 ( Porrhodites ; characters; Baltic region).

— Ganglbauer, 1895: 726 ( Porrhodites ; synonym of fenestralis ).

— Bernhauer and Schubert, 1910: 62 ( Porrhodites ; synonym of fenestralis ).

— Campbell, 1984a: 1230 ( Porrhodites ; synonym of fenestralis ).

brevicollis Mäklin, 1852: 320 ( Arpedium ; Type locality: insula Sitkha).

— Mäklin, 1853: 196 ( Deliphrum ; characters; Kenai Peninsula).

— Redtenbacher, 1857: 992 ( Porrhodites ; characters; Austria).

— Kraatz, 1857d: 963 ( Porrhodites ; characters; Germany). — Redtenbacher, 1874: 271 ( Porrhodites ; characters).

— J. Sahlberg, 1876: 214 ( Porrhodites ; synonym of fenestralis ).

— Fauvel, 1878d: 223 [= 1878e: 59] ( Porrhodites ; synonym of fenestralis ).

— Ganglbauer, 1895: 726 ( Porrhodites ; synonym of fenestralis ).

— Bernhauer and Schubert, 1910: 62 ( Porrhodites ; synonym of fenestralis ).

— Scheerpeltz, 1933: 1053 ( Porrhodites ; synonym of fenestralis ).

— Campbell, 1984a: 1230 ( Porrhodites ; synonym of fenestralis ).

— Zanetti, 1987: 267 ( Porrhodites ; synonym of fenestralis ).

inflatus Hatch, 1957: 68 ( Paradeliphrum ; Type locality: B. C. [ British Columbia ]: Copper Mt.).

— Campbell, 1984a: 1232 ( Porrhodites ; characters; Alberta; British Columbia ; Ontario; Quebec; California; Colorado; New Hampshire; New York; Oregon).

— Levesque and Levesque, 1996: 290 ( Porrhodites ; seasonal abundance; Quebec).

— Downie and Arnett, 1996: 434 ( Porrhodites ; characters; Canada; USA).

- DISTRIBUTION: Canada, USA.

pubescens Zetterstedt, 1828 , see: fenestralis Zetterstedt, 1828 .

Tanyrhinus

[1 species; Nearctic Region]

Tanyrhinus Mannerheim, 1852: 349 (species included: singularis ). TYPE SPECIES: Tanyrhinus singularis Mannerheim , fixed by monotypy.

— Gemminger and Harold, 1870: 2062 (cited as Tanyrrhinus ; catalog).

— Fauvel, 1878d: 230 [= 1878e: 66] (cited as Tanyrrhinus ; characters).

— Duvivier, 1883: 197 (catalog; cited as Tanyrrhinus ).

— Bernhauer and Schubert, 1910: 37 (world catalog; 1 species).

— Leng, 1920: 93 (catalog of North American species).

— Scheerpeltz, 1933: 1032 (world catalog supplement).

— Blackwelder, 1952: 374 (type species: singularis ).

— Hatch, 1957: 50, 68 (characters; species of Pacific Northwest).

— Moore, 1966: 48, 56 (cited as Tanyrrhinus ; characters; notes).

— Moore and Legner, 1975: 216 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 195 (characters; notes).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (1 Nearctic species; notes; characters in key). singularis Mannerheim, 1852: 350 ( Tanyrhinus ; Type locality: insula Sitkha).

— Gemminger and Harold, 1870: 2062 ( Tanyrhinus ; cited as Tanyrrhinus ; catalog).

— Fauvel, 1878d: 230 [= 1878e: 66] ( Tanyrhinus ; cited as Tanyrrhinus ; characters; Alaska).

— Bernhauer and Schubert, 1910: 37 ( Tanyrhinus ; catalog).

— Scheerpeltz, 1933: 1032 ( Tanyrhinus ; catalog).

— Hatch, 1957: 69 ( Tanyrhinus ; characters; British Columbia ; Washington; Oregon).

— Campbell and Davies, 1991: 91 ( Tanyrhinus ; checklist of species; Alaska and Canada).

- DISTRIBUTION: Canada, USA.

Trichodromeus

[7 species; Palaearctic Region]

Trichodromeus Luze, 1903: 116 (subgenus of Geodromicus ; species included: penicillatus , tumidus ). TYPE SPECIES: Hygrogeus penicillatus Reitter , fixed by subsequent designation by Blackwelder, 1952: 305.

— Bernhauer and Schubert, 1910: 75 (subgenus of Geodromicus ).

— Tikhomirova, 1973a: 141 (subgenus of Geodromicus ; checklist of species of USSR).

— Bordoni, 1984a: 57 (valid genus).

— Bordoni, 1990a: 100 (valid genus; characters).

— Zerche, 1992a: 126, 134 (characters; valid genus; key to some species; phylogeny of species; character polarity; phylogenetic relationship of genus).

Neogeodromicus Bordoni, 1985: 373 (species included: ketmeniensis ). TYPE SPECIES: Neogeodromicus ketmeniensis Bordoni , fixed by monotypy.

— Zerche, 1992a: 126 (synonym of Trichodromeus ).

armatus Cameron, 1941: 144 ( Hygrogeus , cited as Hygrogoeus ; Type locality: Kashmir: Aphar Wat, altitude 13,000 feet).

— Zerche, 1992a: 134 ( Trichodromeus ; Pakistan).

- DISTRIBUTION: India, Pakistan.

brevisulcus Champion, 1925 , see: kumaonensis Champion, 1925 .

flavipes Motschulsky, 1860: 551 ( Geodromus ; Type locality: Alpes du Caucase en Touchétie; [Note: See lectotype designation by Bordoni, 1993]).

— Gemminger and Harold, 1868: 658 ( Anthophagus ; synonym of globulicollis ).

— Fauvel, 1869: 493 ( Anthophagus ; synonym of globulicollis ).

— Fauvel, 1871a: 135 [= 1872: 109] ( Geodromicus ; synonym of globulicollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ).

— Bordoni, 1993: 158 ( Neogeodromicus ; lectotype designation; cited as valid species; characters).

- DISTRIBUTION: Georgia.

ketmeniensis Bordoni, 1985: 374 ( Neogeodromicus ; Type locality: Turkestan: Ketmen-tjube, Sussamir Tan).

— Zerche, 1992a: 128 ( Trichodromeus ; characters; Kirghizia; Turkestan; Kazakhstan; Turkmenia; Transcaspian region).

- DISTRIBUTION: Kyrgyzstan, Kazakhstan, Turkmenistan.

kumaonensis Champion, 1925: 105 ( Hygrogeus ; Type locality: Shelshel, alt. 15,750 ft.).

— Cameron, 1930: 166 ( Hygrogeus ; characters; India).

— Scheerpeltz, 1933: 1067 ( Hygrogeus ; catalog).

— Coiffait, 1983: 275 ( Hygrogeus ; Kashmir).

— Zerche, 1992a: 133 ( Trichodromeus ; Pakistan).

- DISTRIBUTION: India, Pakistan.

brevisulcus Champion, 1925: 106 ( Hygrogeus ; variety of kumaonensis ; Type locality: N. Kumaon: Laptel, alt. 15,000 ft.).

— Cameron, 1930: 166 ( Hygrogeus ; variety of kumaonensis ; characters; India).

— Scheerpeltz, 1933: 1067 ( Hygrogeus ; variety of kumaonensis ).

penicillatus Reitter, 1900c: 49 ( Hygrogeus , cited as Hygrogaeus ; Type locality: Buchara: Karatak; [Note: See lectotype designation by Zerche, 1992a]).

— Luze, 1903: 116 ( Geodromicus ; subgenus Trichodromeus ; characters; Buchara; Turkestan).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Tikhomirova, 1973a: 141 ( Geodromicus ; subgenus Trichodromeus ; checklist; Central Asia).

— Bordoni, 1984a: 57 ( Trichodromeus ).

— Bordoni, 1990a: 100 ( Trichodromeus ; characters; notes; Turkestan).

— Zerche, 1992a: 131 ( Trichodromeus ; lectotype designation; characters).

- DISTRIBUTION: Uzbekistan.

tumidus Luze, 1903: 117 ( Geodromicus ; subgenus Trichodromeus ; Type locality: Russisches Zentralasien: Buchara: Karatagh).

— Luze, 1904: 78 ( Geodromicus ; Turkestan).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Tikhomirova, 1973a: 141 ( Geodromicus ; subgenus Trichodromeus ; checklist; Central Asia).

— Bordoni, 1984a: 57 ( Trichodromeus ).

— Bordoni, 1990a: 102 ( Trichodromeus ; characters; notes).

— Zerche, 1992a: 131 ( Trichodromeus ; synonym of penicillatus ).

sogdianus Bordoni, 1990 a: 104 ( Trichodromeus ; Type locality: Bukara: “Mts. Karateghin, Baldschuan, 924 m).

— Zerche, 1992a: 131 ( Trichodromeus ; synonym of penicillatus ).

schmidti Zerche, 1992 a: 130 ( Trichodromeus ; Type locality: Asia centr., Tadsch., Dschirgatalski Raj., Sukran-Schlucht, 2590 m / Burs-Gletscher-Fuss).

- DISTRIBUTION: Tajikistan.

sogdianus Bordoni, 1990 , see: penicillatus Reitter, 1900 .

transversus Motschulsky, 1860: 550 ( Geodromus ; Type locality: Alpes du Caucase en Touschéte; [Note: See lectotype designation by Bordoni, 1993]).

— Fauvel, 1871a: 136 [= 1872: 110] ( Geodromicus ; synonym of globulicollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ).

— Bordoni, 1993: 158 ( Neogeodromicus ; lectotype designation; cited as valid species; characters). - DISTRIBUTION: Caucasus.

tumidus Luze, 1903 , see: penicillatus Reitter, 1900 .

Trigonodemus

[11 species; Nearctic and Palaearctic Regions]

Trigonodemus LeConte, 1863 a: 56 (species included: striatus ). TYPE SPECIES: Trigonodemus striatus LeConte , fixed by monotypy.

— LeConte, 1863: 26 ([nomen nudum]; list of North American species).

— Gemminger and Harold, 1868: 663 (catalog; cited as Trigonodesmus ).

— Fauvel, 1878d: 231 [= 1878e: 67] (characters).

— Bernhauer and Schubert, 1910: 37 (world catalog; 1 species).

— Leng, 1920: 93 (catalog of North American species).

— Scheerpeltz, 1933: 1032 (world catalog supplement).

— Blackwelder, 1952: 396 (type species: striatus ).

— Hatch, 1957: 50, 68 (characters; species of Pacific Northwest).

— Moore, 1966: 48, 56 (characters; notes).

— Tikhomirova, 1973a: 137 (checklist of species of USSR).

— Moore and Legner, 1975: 217 (catalog; North America north of Mexico).

— Shibata, 1976: 97 (checklist of species of Japan).

— Moore and Legner, 1979: 196 (characters; notes).

— Watanabe, 1990: 250 (characters; species of Japan).

— Campbell and Davies, 1991: 91 (checklist of species; Alaska and Canada).

— Smetana, 1996d: 2 (review; characters; notes; key to species).

— Smetana, 1996h: 245 (key to species).

— Downie and Arnett, 1996: 434 (characters; species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (2 Nearctic species; notes; characters in key).

Arimimelus Kraatz, 1877: 104 (species included: lebioides ). TYPE SPECIES: Arimimelus lebioides Kraatz , fixed by monotypy.

— Sharp, 1889: 473 (synonym of Trigonodemus by citation of lebioides in Trigonodemus ).

— Bernhauer and Schubert, 1910: 37 (synonym of Trigonodemus ).

— Blackwelder, 1952: 61 (type species: lebioides ).

— Watanabe, 1990: 250 (synonym of Trigonodemus ).

— Smetana, 1996d: 2 (synonym of Trigonodemus ).

— Smetana, 1996h: 245 (key to species).

— Smetana, 2000: 301 (key to species).

Klapperichianellia Hlisnikovsky´, 1962: 455, 456 (species included: mirabilis ). TYPE SPECIES: Klapperichianellia mirabilis Hlisnikovsky´, fixed by original designation and monotypy.

— Thayer, 1987: 389 (may be synonym of Trigonodemus ).

— Smetana, 1996d: 2 (synonym of Trigonodemus ).

audax Smetana, 1996d: 11 ( Trigonodemus ; Type locality: Taiwan: Nantou Hsien, Yushan N.P., S.W. slope blw. Yushan Mn. Pk. 3650 m).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

fasciatus Leech, 1939: 258 ( Trigonodemus ; Type locality: Vancouver, B. C. [ British Columbia ]).

— Hatch, 1957: 68 ( Trigonodemus ; characters; British Columbia ; Oregon).

— Smetana, 1996d: 6 ( Trigonodemus ; characters; British Columbia ).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 301 ( Trigonodemus ; characters in key).

- DISTRIBUTION: Canada, USA.

fungicola Smetana, 1996d: 9 ( Trigonodemus ; Type locality: China: Sichuan, Gongga Shan, above Camp 3, 3050 m).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

lebioides Kraatz, 1877: 105 ( Arimimelus ; Type locality: Hagi).

— Sharp, 1889: 473 ( Trigonodemus ; notes; Japan).

— Scheerpeltz, 1933: 1032 ( Trigonodemus ; catalog).

— Nakane, 1963a: 82 ( Trigonodemus ; characters; habitus photograph; Japan).

— Watanabe, 1990: 253 ( Trigonodemus ; characters; Japan).

— J. Li, 1993: 18 ( Trigonodemus ; China).

— Smetana, 1996d: 8 ( Trigonodemus ; characters; Japan).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: Japan, China.

modestus Smetana, 2000: 300 ( Trigonodemus ; Type locality: China: W. Sichuan: Ganzi Tibet Aut. Pref., Kangding Co., Daxue Shan, Mu Ge Cuo , ob. See, 15 km NW Kangding, 3700 m, 30°09N, 101°52E).

- DISTRIBUTION: China.

montanus Smetana, 1996 h: 241 ( Trigonodemus ; Type locality: China: N. Yunnan, Xue Shan nr. Zhongdian , 4000–4100 m).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

mirabilis Hlisnikovsky´, 1962: 458 ( Klapperichianellia ; Type locality: China: Fukien , Kuatun, 2300 m, 27.40 nördl. Breite, 114.40 östl Länge) .

— Schawaller, 1978: 176, 181, 182, 183 ( Klapperichianellia ; notes; habitus outline; type locality). — Smetana, 1996d: 13 ( Trigonodemus ; characters; China).

— Smetana, 1996h: 245 ( Trigonodemus ; characters in key).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

pictus Smetana, 2000: 295 ( Trigonodemus ; Type locality: China: NW Yunnan, Bai Ma Xue Shan, 35 km S. Dêqên, 4300–4800).

- DISTRIBUTION: China.

puncticollis Smetana, 2000: 298 ( Trigonodemus ; Type locality: China: Sichuan, sud Barkam ). - DISTRIBUTION: China.

schuelkei Smetana, 1996 h: 244 ( Trigonodemus ; Type locality: China: Shaanxi, Qin Ling Shan, 107.56E, 33.45 N, Autoroute   GoogleMaps km 93 S of Zhouzhi, 108 km SW Xian Mountain Forest, 1650 m).

— Smetana, 2000: 302 ( Trigonodemus ; characters in key).

- DISTRIBUTION: China.

striatus LeConte, 1863 a: 57 ( Trigonodemus ; Type locality: Southern States).

— Fauvel, 1878d: 232 [= 1878e: 68] ( Trigonodemus ; characters; Canada; West Virginia).

— Bernhauer and Schubert, 1910: 37 ( Trigonodemus ; catalog).

— Smetana, 1996d: 5 ( Trigonodemus ; characters; Nova Scotia ; Ontario; Quebec; New Hampshire; New York).

— Downie and Arnett, 1996: 434 ( Trigonodemus ; characters; USA).

— Smetana, 2000: 301 ( Trigonodemus ; characters in key).

- DISTRIBUTION: Canada, USA.

Unamis

[5 species; Nearctic Region]

Unamis Casey, 1894: 398 , 400 (species included: truncata ). TYPE SPECIES: Lesteva truncata Casey , fixed by monotypy.

— Bernhauer and Schubert, 1910: 73 (world catalog; 1 species).

— Leng, 1920: 95 (catalog of North American species).

— Blackwelder, 1952: 402 (type species: truncata ).

— Hatch, 1957: 49, 56 (characters; key to species of Pacific Northwest).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1972: 21 (characters; key to species).

— Moore and Legner, 1975: 218 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 198 (characters; notes).

— Campbell and Davies, 1991: 90 (checklist of species; Alaska and Canada).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (5 Nearctic species; notes; characters in key). bjorkmanae Hatch, 1957: 57 ( Unamis ; Type locality: Crater L., Oregon).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: USA.

columbiensis Hatch, 1957: 56 ( Unamis ; Type locality: Sanca, B. C. [ British Columbia ]).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: Canada, USA.

fulvipes Fall, 1922: 12 ( Unamis ; Type locality: California: Humboldt County: Korbel ).

— Hatch, 1957: 57 ( Unamis ; characters; British Columbia ).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: Canada, USA.

giulianii Moore and Legner, 1972: 21 ( Unamis ; Type locality: California: Inyo County, near Finger Lake , 11,300 feet).

- DISTRIBUTION: USA.

truncata Casey, 1885 a: 322 ( Lesteva ; Type locality: California: Middle Sierras ).

— Casey, 1894: 400 ( Unamis ).

— Bernhauer and Schubert, 1910: 73 ( Unamis ; catalog).

— Fall, 1922: 13 ( Unamis ; Colorado specimen provisionally referred to genus).

— Hatch, 1957: 56 ( Unamis ; characters; Idaho; Washington; Oregon).

— Moore and Legner, 1972: 21 ( Unamis ; characters).

- DISTRIBUTION: USA.

Uenohadesina

[1 species; Palaearctic Region]

Uenohadesina Smetana, 2000 a: 285 (species included: styx ). TYPE SPECIES: Uenohadesina styx Smetana , fixed by original designation and monotypy.

styx Smetana, 2000 a: 291 ( Uenohadesina ; Type locality: South Korea: Yong’yeon-gul Cave nr. Hwangji , Changseong-eub, Kangweon-do).

- DISTRIBUTION: South Korea.

Vellica

[1 species; Nearctic Region]

Vellica Casey, 1885 a: 321 (species included: longipennis ). TYPE SPECIES: Vellica longipennis Casey , fixed by monotypy.

— Casey, 1894: 401 (notes).

— Leng, 1920: 95 (catalog of North American species).

— Blackwelder, 1952: 403 (type species: longipennis ).

— Hatch, 1957: 49, 61 (characters; species of Pacific Northwest).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1975: 219 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 199 (characters; notes).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (1 Nearctic species; notes; characters in key). longipennis Casey, 1885 a: 321 ( Vellica ; Type locality: California: Middle Sierras).

— Bernhauer and Schubert, 1910: 74 ( Vellica ; catalog).

— Hatch, 1957: 61 ( Vellica ; characters; British Columbia ).

— Campbell and Davies, 1991: 90 ( Vellica ; checklist of species; Canada).

- DISTRIBUTION: Canada, USA.

Xenicopoda

[1 species; Nearctic Region]

Xenicopoda Moore and Legner, 1971: 51 (species included: helenae ). TYPE SPECIES: Xenicopoda helenae Moore and Legner , fixed by original designation and monotypy.

— Moore and Legner, 1975: 220 (catalog; North America north of Mexico).

— Thayer, 1978: 146 (characters; notes; discussion).

— Moore and Legner, 1979: 200 (characters; notes).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (1 Nearctic species; notes; characters in key). helenae Moore and Legner, 1971: 53 ( Xenicopoda ; Type locality: Mt. Wilson, Calif.).

— Thayer, 1978: 143 ( Xenicopoda ; characters; notes; California).

- DISTRIBUTION: USA.

Tribe Aphaenostemmini

Aphaenostemmini Peyerimhoff, 1914: 248 (genera included: Aphaenostemmus ). TYPE GENUS: Aphaenostemmus (see Newton and Thayer, 1992: 55).

— Scheerpeltz, 1933: 1031 (world catalog supplement).

— Peyerimhoff, 1939: 212 (characters in key).

— Scheerpeltz, 1940: 14 (key to Palaearctic genera).

— Newton and Thayer, 1995: 247–286, 295 (phylogenetic relationships; characters; discussion; tribe of Omaliinae ; includes Aphaenostemmus , Giulianium ).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (characters; notes).

Aphaenostemminae Peyerimhoff, 1914.

— Newton and Thayer, 1992: 55 (summary of current classification; notes).

Aphaenostemmus

[5 species; Oriental and Palaearctic Regions]

Aphaenostemmus Peyerimhoff, 1914: 245 (species included: bordei ). TYPE SPECIES: Aphaenostemmus bordei Peyerimhoff , fixed by monotypy.

— Winkler, 1925: 326 (catalog for Palaearctic region).

— Scheerpeltz, 1933: 1031 (world catalog supplement).

— Normand, 1935: 357 (list of species of Tunisia).

— C. Koch, 1936c: 332 (key to species).

— Peyerimhoff, 1939: 211 (characters; key to North African species).

— Blackwelder, 1952: 58 (type species: bordei ).

— Kocher, 1958: 92 (checklist of species; Morocco).

— Newton and Thayer, 1995: 296 (list of synonyms).

Torretassoella C. Koch, 1936b: 126 (species included: testacea ). TYPE SPECIES: Torretassoella testacea C. Koch , fixed by monotypy.

— C. Koch, 1936c: 332 (synonym of Aphaenostemmus ).

— Peyerimhoff, 1939: 211 (synonym of Aphaenostemmus ).

Paratriga Cameron, 1940c: 209 (species included: indica , associata ). TYPE SPECIES: Paratriga indica Cameron , fixed by original designation.

— Blackwelder, 1952: 294 (type species: indica ).

— Newton and Thayer, 1995: 296 (synonym of Aphaenostemmus ).

associatus Cameron, 1940c: 209 ( Paratriga ; Type locality: United Provinces: Dehra Dun).

— Newton and Thayer, 1995: 296 ( Aphaenostemmus ).

- DISTRIBUTION: India.

bordei Peyerimhoff, 1914: 248 ( Aphaenostemmus ; Type locality: Biskra).

— Scheerpeltz, 1933: 1031 ( Aphaenostemmus ; catalog).

— C. Koch, 1936c: 332 ( Aphaenostemmus ; characters; Algeria).

— Peyerimhoff, 1939: 213 ( Aphaenostemmus ; characters in key).

- DISTRIBUTION: Algeria.

himalayicus Bernhauer, 1943 a: 170 ( Aphaenostemmus ; Type locality: Dehra-Dun: Bhagwantung, Himalaya).

- DISTRIBUTION: India.

indicus Cameron, 1940c: 209 ( Paratriga ; Type locality: United Provinces: Dehra Dun).

— Newton and Thayer, 1995: 296 ( Aphaenostemmus ).

- DISTRIBUTION: India.

testaceus C. Koch, 1936b: 126 View in CoL , 133 (Torretassoella; Type locality: Aegypten: Wadi Digla. Sinai: Wadi Isla; Wadi Luotaie ).

— C. Koch, 1936c: 332 ( Aphaenostemmus View in CoL ; characters; Egypt).

— Peyerimhoff, 1939: 213 ( Aphaenostemmus , transferred from Torretassoella; characters in key).

— Coiffait, 1979a: 163, 179 ( Aphaenostemmus ; illustrations of forebody and aedeagus; Saudi Arabia).

— Gusarov, 1997: 278 ( Aphaenostemmus ; Oman).

- DISTRIBUTION: Egypt, Saudia Arabia, Oman.

Giulianium

[3 species; Nearctic and Palaearctic Regions]

Giulianium Moore, 1976: 56 (species included: campbelli ). TYPE SPECIES: Giulianium campbelli Moore , fixed by original designation and monotypy.

— Moore and Legner, 1979: 256 (characters; notes).

— Smetana and Campbell, 1980: 1069 (characters).

— Newton and Thayer, 1995: 295 ( Aphaenostemmini ).

— Ahn and Ashe, 1999: 159 (characters; key to species).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (3 Nearctic species; notes; characters in key).

alaskanum Ahn and Ashe, 1999: 162 ( Giulianium ; Type locality: USA: Alaska: Seward. Also cited from Japan).

- DISTRIBUTION: USA, Japan.

campbelli Moore, 1976: 57 ( Giulianium ; Type locality: California: Humboldt County: Bear Harbor ).

— Ahn and Ashe, 1999: 162 ( Giulianium ; characters; USA).

- DISTRIBUTION: USA.

newtoni Ahn and Ashe, 1999: 163 ( Giulianium ; Type locality: USA: California: San Mateo Co., Moss Beach ).

- DISTRIBUTION: USA.

Tribe Corneolabiini

Corneolabiini Steel, 1950: 54 (including key to genera; genera included: Corneolabium , Paracorneolabium , Metacorneolabium ). TYPE GENUS: Corneolabium , by original designation.

— Thayer, 1985: 115 (characters; notes).

— Newton and Thayer, 1992: 58 (tribe of Omaliinae ).

— Newton and Thayer, 1995: 247–286, 295 (phylogenetic discussion; characters; discussion; includes Corneolabium , Paracorneolabium ; Metacorneolabium ).

— Klimaszewski, Newton, and Thayer, 1996: 146 (list of New Zealand genera; number of New Zealand species).

Corneolabium

[1 species; Australian Region]

Corneolabium Steel, 1950: 54 (species included: mandibulare ). TYPE SPECIES: Corneolabium mandibulare Steel , fixed by original designation and monotypy.

— Thayer, 1985: 115, 163 (characters; phylogenetic relationships).

mandibulare Steel, 1950: 56 ( Corneolabium ; Type locality: New Zealand: West Plains ).

— Klimaszewski, Newton, and Thayer, 1996: 146 ( Corneolabium ; endemic to New Zealand).

- DISTRIBUTION: New Zealand.

Metacorneolabium

[22 species; Neotropical and Australian Regions]

Metacorneolabium Steel, 1950: 62 (species included: minus , cited as minor ). TYPE SPECIES: Metacorneolabium minus Steel , fixed by original designation and monotypy.

— Thayer, 1985: 116, 122, 135, 144, 163 (characters; revision of and key to species; habitats; phylogeny; biogeography).

Metomalium Coiffait and Saiz, 1968: 449 (species included: chilensis ). TYPE SPECIES: Metomalium chilense Coiffait and Saiz , fixed by original designation and monotypy. — Thayer, 1985: 116 (synonym of Metacorneolabium ).

alulatum Thayer, 1985: 137 ( Metacorneolabium ; Type locality: Australia: Tasmania: W side Lake St. Clair , c. 750 m).

- DISTRIBUTION: Australia.

batyle Thayer, 1985: 155 ( Metacorneolabium ; Type locality: Chile: Concepción Prov. : Ramuncho).

- DISTRIBUTION: Chile.

bipertica Thayer, 1985: 159 ( Metacorneolabium ; Type locality: Chile: Osorno Prov. : Parque Nacional Puyehue, Antillanca road, 965 m).

- DISTRIBUTION: Chile.

chilense Coiffait and Saiz, 1968: 450 ( Metomalium ; cited as chilensis ; Type locality: Chili: Nahuelbuta, Malleco).

— Thayer, 1985: 148 ( Metacorneolabium ; emended to chilense ; characters; Chile).

- DISTRIBUTION: Chile.

convexum Thayer, 1985: 123 ( Metacorneolabium ; Type locality: New Zealand: NC: 8.0 km S [town of] Arthur’s Pass , 670 m).

- DISTRIBUTION: New Zealand.

dactylotum Thayer, 1985: 154 ( Metacorneolabium ; Type locality: Chile: Osorno Prov. : Parque Nacional Puyehue, Antillanca road, 965 m).

- DISTRIBUTION: Chile.

darlingtoni Thayer, 1985: 138 ( Metacorneolabium ; Type locality: Australia: Tasmania: Lower Gordon River   GoogleMaps , 42.31S, 145.45E – 42.32S, 145.47E 7).

- DISTRIBUTION: Australia.

diffusum Thayer, 1985: 156 ( Metacorneolabium ; Type locality: Chile: Llanquihue Prov. : Lago Chapo, 13. 5 km E Correntoso, 310 m).

- DISTRIBUTION: Chile.

echidna Thayer, 1985: 142 ( Metacorneolabium ; Type locality: Australia: Victoria: Mt. Buf-

falo, 500 m).

exochlum Thayer, 1985: 158 (Metacorneolabium; Type locality: Chile: Cautín Prov.: Volcán Villarrica, 1250 m).

- DISTRIBUTION: Chile.

exuberatum Thayer, 1985: 145 (Metacorneolabium; Type locality: Chile: Cautín Prov.: Volcán Villarrica, 1250 m).

- DISTRIBUTION: Chile.

gigas Thayer, 1985: 124 (Metacorneolabium; Type locality: New Zealand: BR: Mt. Murchison, 1420 m).

- DISTRIBUTION: New Zealand.

hokitika Thayer, 1985: 132 (Metacorneolabium; Type locality: New Zealand: WD: Hokitika, Glow Worm Dell, 5 m).

- DISTRIBUTION: New Zealand.

logor Thayer GoogleMaps , 1985: 143 ( Metacorneolabium   GoogleMaps ; Type locality: Australia   GoogleMaps : Tasmania: Lower Gordon River, 42.42S, 145.53E- 42.41.5S, 145.53E).

- DISTRIBUTION: Australia.

minus Steel, 1950: 64 (Metacorneolabium; cited as minor; Type locality: New Zealand: Mount Hutt).

— Thayer, 1985: 130 (Metacorneolabium; emended from minor to minus; characters; New Zealand).

— Emberson, 1998: 32 (Metacorneolabium; collecting notes; Chatham Islands).

- DISTRIBUTION: New Zealand, Chatham Islands.

pilatum Thayer, 1985: 153 (Metacorneolabium; Type locality: Chile: Osorno Prov.: Parque Nacional Puyehue, Volcán Casa Blanca, 1370 m).

- DISTRIBUTION: Chile.

pyriforme Thayer, 1985: 125 (Metacorneolabium; Type locality: New Zealand: BR: Nelson Lakes N. P., Lake Rotoiti, 600 m).

- DISTRIBUTION: New Zealand.

rangipo Thayer, 1985: 133 (Metacorneolabium; Type locality: New Zealand: TO: Rangipo Intake, 36 road km S Turangi, 900 m).

- DISTRIBUTION: New Zealand.

rivalis Thayer, 1985: 147 (Metacorneolabium; Type locality: Chile: Osorno Prov.: 7. 7 km NE Termas de Puyehue, 200 m).

- DISTRIBUTION: Chile.

soror Thayer, 1985: 161 (Metacorneolabium; Type locality: Chile: Malleco Prov.: 12 km E Malalcahuello, 1350 m).

- DISTRIBUTION: Chile.

victas Thayer, 1985: 135 (Metacorneolabium; Type locality: Australia: Tasmania: Rufus Canal, 13. 5 km WNW Derwent Bridge, 800 m).

- DISTRIBUTION: Australia.

zanotium Thayer, 1985: 129 (Metacorneolabium; Type locality: New Zealand: SL: Owaka, Glenomaru Reserve).

- DISTRIBUTION: New Zealand.

Paracorneolabium

[1 species; Australian Region]

Paracorneolabium Steel, 1950: 59 (species included: brouni ). TYPE SPECIES: Paracorneolabium brouni Steel , fixed by original designation and monotypy.

— Thayer, 1985: 115-116, 162-171 (characters; phylogenetic relationships).

brouni Steel, 1950: 60 ( Paracorneolabium ; Type locality: New Zealand: Hollyd.; according to Steel this is probably an abbreviation for Hollyford).

Tribe Coryphiini

Coryphiini Jakobson, 1908 .

— Portevin, 1929: 430 (key to French genera).

— Joy, 1932: 97 (key to British genera).

— Hatch, 1957: 49 (characters; key to genera of Pacific Northwest).

— Arnett, 1963: 241, 259 (key to North American genera; characters; list of North American genera).

— Campbell, 1978a: 22 (characters; key to genera and species; Eudectus , Haida , Pseudohaida excluded; biological notes).

— Moore and Legner, 1979: 161 (characters; tabular key to genera of North America north of Mexico).

— Muona, 1979: 18 (list of Scandinavian species).

— Outerelo and Gamarra, 1985: 82 (key to genera of Iberian Peninsula).

— Watanabe, 1990: 264, 377 (characters; key to genera of Japan; relationships of genera).

— Zerche, 1989: 27 (supplement to Lohse, 1964).

— Zerche, 1990 (monograph; phylogeny; characters; checklist of and keys to taxa of Palaearctic region; natural history notes; systematic position and taxonomic status of some Nearctic taxa changed).

— Newton and Thayer, 1992: 58 (tribe of Omaliinae ).

— Zerche, 1993: 352, 354–357 ( Geodromicus, Liophilydrodes and Philydrodes placed in Hygrogeus group of Anthophagini ; bionomics; Zoogeography).

— Newton and Thayer, 1995: 247–286, 294 (phylogenetic relationships; characters; discussion).

— M. Hansen, 1996: 95 (list of Danish species).

— Newton, Thayer, Ashe, and Chandler, 2000: 285 (key to Nearctic genera).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (characters; notes).

Subtribe Boreaphilina

Boreaphilina Zerche, 1990: 22 , 158 (genera included: Archaeoboreaphilus , Caloboreaphilus , Boreaphilus , Niphetodes, Hypsonothrus ). TYPE GENUS: Boreaphilus (see Newton and Thayer, 1992: 58).

— Newton and Thayer, 1992: 58 (subtribe of Coryphiini ).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (characters; notes).

Boréaphilaires Mulsant and Rey, 1880: 391 (genera included: Coryphium , Eudectus , Boreaphilus ). TYPE GENUS: Boreaphilus (see Newton and Thayer, 1992: 58).

— Newton and Thayer, 1992: 58 (synonym of Boreaphilina ).

Archaeoboreaphilus

[6 species; Palaearctic Region]

Archaeoboreaphilus Zerche, 1990: 158 (species included: nikkoensis , miyamorii , tateoi , fuscus , iwakisanensis , yasutoshii ). TYPE SPECIES: Boreaphilus nikkoensis Sawada , fixed by original designation.

fuscus Watanabe, 1988: 53 ( Coryphium ; Type locality: Japan: Honshu: Yamaguchi Pref.: Azami-ga-take, Kanochô ).

— Watanabe, 1990: 362 ( Coryphium ; characters; Japan).

— Zerche, 1990: 164 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

iwakisanensis Watanabe, 1988: 54 ( Coryphium ; Type locality: Japan: Honshu: Aomori Pref.: Mt. Iwakisan ).

— Watanabe, 1990: 363 ( Coryphium ; characters; Japan).

— Zerche, 1990: 165 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

miyamorii Watanabe, 1988: 49 ( Coryphium ; Type locality: Japan: Hokkaido: Horobetsukôzan, Iburi ).

— Watanabe, 1990: 359 ( Coryphium ; characters; Japan).

nikkoensis Sawada, 1964: 36 ( Boreaphilus ; Type locality: Japan: Honshu: Pref. Tochigi: Chuzenji near Nikko ).

— Watanabe, 1988: 56 ( Coryphium ; characters; Japan).

— Watanabe, 1990: 364 ( Coryphium ; characters; Japan).

— Zerche, 1990: 159 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

tateoi Watanabe, 1988: 52 ( Coryphium ; Type locality: Japan: Honshu: Nara Pref.: Tônomine, Yamato ).

— Watanabe, 1990: 361 ( Coryphium ; characters; Japan).

— Zerche, 1990: 164 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

yasutoshii Watanabe, 1988: 58 ( Coryphium ; Type locality: Japan: Honshu: Gunma Pref.: near Edozawa, Mt. Jôshû-Hotaka ).

— Watanabe, 1990: 366 ( Coryphium ; characters; Japan).

— Zerche, 1990: 166 ( Archaeoboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

Boreaphilus

[23 species; Nearctic and Palaearctic Regions]

Boreaphilus C. Sahlberg, 1832: 433 (species included: henningianus ). TYPE SPECIES: Boreaphilus henningianus C. Sahlberg , fixed by monotypy.

— Zetterstedt, 1838: 72 (characters).

— Erichson, 1840: 899 (characters).

— Haliday, 1841: 186 (characters).

— Schaum, 1852: 31 (list of species; Europe).

— Lacordaire, 1854: 137 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 632 (characters).

— Kraatz, 1857: 39 (characters).

— Jacquelin du Val, 1857: 67 (characters).

— Redtenbacher, 1857: 245 (characters).

— Thomson, 1858: 37 (characters).

— Thomson, 1859: 49 (characters; type species: henningianus ).

— Schaum, 1859: 32 (catalog; European species).

— Thomson, 1861: 183 (characters).

— Gemminger and Harold, 1868: 664 (catalog).

— Jacquelin du Val, 1868: 79 (catalog; European species).

— Fauvel, 1871a: 106 [= 1872: 80] (characters).

— Redtenbacher, 1874: 273 (characters).

— Fauvel, 1875a:V [= 1875b: 207] (catalog).

— Mulsant and Rey, 1880: 401 (characters).

— Heyden, 1880: 82 (list of species of Siberian region).

— Duvivier, 1883: 197 (catalog).

— Seidlitz, 1891: 349 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 118 (list of species of Europe and Caucasus).

— Heyden, 1893: 48 (list of species of Siberian region).

— Ganglbauer, 1895: 699 (characters; key to middle European species).

— Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 84 (world catalog; 8 species).

— Notman, 1918 (characters; notes).

— Leng, 1920: 95 (catalog of North American species).

— Winkler, 1925: 337 (catalog for Palaearctic region).

— Porta, 1926: 33 (characters; species of Italy).

— Scheerpeltz, 1933: 1072 (world catalog supplement).

— Normand, 1935: 358 (list of species of Tunisia).

— Blackwelder, 1952: 83 (type species: henningianus ).

— Smetana, 1959a: 196 (list of species collected in Albania).

— Horion, 1963: 154 (list of central European species).

— Székessy, 1963: 24 (characters).

— Lohse, 1964: 69 (characters; key to central European species).

— Scheerpeltz, 1968a: 19 (catalog of Austrian species).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Moore and Legner, 1975: 186 (catalog; North America north of Mexico).

— Shibata, 1976: 125 (checklist of species of Japan).

— Campbell, 1978a: 24, 58 (key to species; characters; notes).

— Moore and Legner, 1979: 162 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Tóth, 1982: 107 (characters; habitus; species of Hungary).

— Muona and Viramo, 1986: 16 (list of species of northeastern Finland).

— Zanetti, 1987: 434 (characters; habitus; key to species of Italy).

— Zerche, 1989: 123, 129 (characters; notes; distributional dates discussed).

— Watanabe, 1990: 370 (characters; key to species of Japan).

— Zerche, 1990: 169, 286, 306 (characters; key to species of Palaearctic region; phylogeny; distribution).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Zerche, 1993: 344, 350, 355 (modification of key; phylogeny; distribution; distribution in Nearctic and Palaearctic regions compared).

— Siitonen, 1993: 230 (list of Finnish species).

— Zanetti, 1995: 10 (list of species of Italy).

— Downie and Arnett, 1996: 436 (characters; key to species of Northeastern North America ).

— Kashcheev, 1999: 144 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (1 Nearctic species; notes; characters in key).

Chevrieria Heer, 1839: 188 (species included: velox ). TYPE SPECIES: Chevrieria velox Heer , fixed by monotypy.

— Heer, 1841: 571 (notes).

— Schaum, 1852: 30 (list of species; Europe).

— Lacordaire, 1854: 135 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 631 (characters).

— Jacquelin du Val, 1857: 67 (synonym of Boreaphilus ).

— Redtenbacher, 1857: 241 (characters).

— Redtenbacher, 1857: 992 (synonym of Boreaphilus ).

— Schaum, 1859: 32 (synonym of Boreaphilus ).

— Gemminger and Harold, 1868: 664 (synonym of Boreaphilus ).

— Fauvel, 1871a: 106 [= 1872: 80] (synonym of Boreaphilus ).

— Ganglbauer, 1895: 699 (synonym of Boreaphilus ).

— Bernhauer and Schubert, 1910: 84 (synonym of Boreaphilus ).

— Scheerpeltz, 1933: 1072 (synonym of Boreaphilus ).

— Blackwelder, 1952: 99 (type species: velox ).

— Campbell, 1978a: 59 (synonym of Boreaphilus ).

— Zanetti, 1987: 434 (synonym of Boreaphilus ).

— Zerche, 1990: 169 (synonym of Boreaphilus ).

Boreophilus Agassiz, 1847: 49 (emendation of Boreaphilus ). TYPE SPECIES: Boreaphilus henningianus C. Sahlberg , fixed by objective synonymy with Boreaphilus .

— Blackwelder, 1952: 84 (type species).

— Campbell, 1978a: 59 (synonym of Boreaphilus ).

Catocopa Gistel, 1856: 29 . TYPE SPECIES: Chevreiria velox Heer , fixed by monotypy.

— Blackwelder, 1952: 96 (synonym of Boreaphilus ; type species: velox ).

— Zerche, 1990: 169 (synonym of Boreaphilus ).

albanicus Zerche, 1990: 197 ( Boreaphilus ; Type locality: Nordalban.: Alpen Casa Valbones).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Philorinum

Loc

Philorinum nitidulum

Herman, Lee H. 2001
2001
Loc

testaceus

C. Koch 1936: 126
1936
Loc

Aphaenostemmus

Peyerimhoff 1914
1914
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