Harpognatus robynsii, Wesmael, 1834, 1833

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

robynsii Wesmael, 1834 , see: angusticolle Stephens, 1834 .

sikkimense Zerche, 1990: 62 ( Coryphium ; Type locality: India: Sikkim, Lachen , 8950 ft.).

- DISTRIBUTION: India.

smetanaorum Zerche, 1993: 325 ( Coryphium ; Type locality: Nepal: Bhakta B. /Syabnu,

2200–3350 m).

- DISTRIBUTION: Nepal.

tateoi Watanabe, 1988 , see: Archaeoboreaphilus .

tchyldebayevi Kashcheev, 1999: 143 ( Coryphium ; Type locality: Zailiiskii Alatau, pereval

Kysylauc).

- DISTRIBUTION: Kazakhstan.

turcicum Zerche, 1993: 322 ( Coryphium ; Type locality: Turquie: Artvin).

- DISTRIBUTION: Turkey.

uludaghense Fagel, 1971: 121 ( Coryphium ; Type locality: Anatolie occidentale: vilayet de

Bursa, Uludagh, 1800–1900 m).

— Zerche, 1990: 50 ( Coryphium ; characters; Turkey).

- DISTRIBUTION: Turkey.

venustus Filatova, 1981 , see: Coryphiocnemus .

yasutoshii Watanabe, 1988 , see: Archaeoboreaphilus .

Ephelinus

[3 species; Nearctic Region]

Ephelinus Cockerell, 1906: 241 (replacement name for Ephelis Fauvel ). TYPE SPECIES: Coryphium pallidum LeConte , by subsequent designation by R. Lucas, 1920: 267.

— Bernhauer and Schubert, 1910: 83 (world catalog; 3 species).

— Leng, 1920: 95 (catalog of North American species).

— Scheerpeltz, 1933: 1070 (world catalog supplement).

— Blackwelder, 1952: 149 (type species).

— Hatch, 1957: 52, 75 (characters; key to species of Pacific Northwest).

— Moore and Legner, 1975: 188 (catalog; North America north of Mexico).

— Campbell, 1978a: 24, 64 (key to species; characters; notes).

— Moore and Legner, 1979: 163 (characters; notes).

— Frank, 1986a: 365 (checklist of species of Florida).

— Zerche, 1990: 261 (characters discussed; Coryphiini ).

— Downie and Arnett, 1996: 435 (characters; key to species of Northeastern North America ).

— Lundgren, 1998: 38 (list of species from Florida; records for other States listed for each species).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 ( Nearctic species; notes; characters in key).

Ephelis Fauvel, 1878d: 219 [= 1878e: 55] (preoccupied by Lederer, 1863; species included: pallida , guttata, notata ). TYPE SPECIES: Corphium pallidum LeConte , fixed by objective synonymy with Ephelinus for which R. Lucas fixed the type species.

— Duvivier, 1883: 197 (catalog).

— Bernhauer and Schubert, 1910: 83 (synonym of Ephelinus ).

— Blatchley, 1910: 481 (characters).

— Scheerpeltz, 1933: 1070 (synonym of Ephelinus ).

— Blackwelder, 1952: 149 (type species).

— Campbell, 1978a: 64 (synonym of Ephelinus ).

arizonensis Bernhauer, 1912 , see: Coryphium .

brunneum Hatch, 1957 , see: Coryphium .

guttatus LeConte, 1863 a: 58 ( Coryphium ; Type locality: Southern States ).

— Fauvel, 1878d: 220 [= 1878e: 56] ( Ephelis ; characters; Texas).

— Bernhauer and Schubert, 1910: 83 ( Ephelinus ; catalog). — Campbell, 1978a: 66 ( Ephelinus ; characters; District of Columbia; North Carolina; South Carolina; Tennessee; Texas; Virginia).

— Downie and Arnett, 1996: 435 ( Ephelinus ; characters; USA).

- DISTRIBUTION: USA.

notatus LeConte, 1863 a: 58 ( Coryphium ; Type locality: Pennsylvania).

— Fauvel, 1878d: 221 [= 1878e: 57] ( Ephelis ; characters; Michigan; Pennsylvania; Illinois; South Carolina).

— Bernhauer and Schubert, 1910: 83 ( Ephelinus ; catalog).

— Blatchley, 1910: 482 ( Ephelis ; characters; Indiana).

— Campbell, 1978a: 68 ( Ephelinus ; characters; Alabama; District of Columbia; Florida; Illinois; Maryland; Massachusetts; Michigan; Mississippi; Missouri; Ohio; Pennsylvania; Texas; Virginia).

— Downie and Arnett, 1996: 435 ( Ephelinus ; characters; USA).

- DISTRIBUTION: USA.

pallidus LeConte, 1863 a: 57 ( Coryphium ; Type locality: Kansas).

— Fauvel, 1878d: 220 [= 1878e: 56] ( Ephelis ; characters; Kansas).

— Bernhauer and Schubert, 1910: 83 ( Ephelinus ; catalog).

— Campbell, 1978a: 71 ( Ephelinus ; characters; Illinois; Kansas).

— Downie and Arnett, 1996: 435 ( Ephelinus ; characters; USA).

- DISTRIBUTION: USA.

Eudectus

[7 species; Palaearctic and Nearctic Regions]

Eudectus Redtenbacher, 1857: 245 (species included: giraudi ). TYPE SPECIES: Eudectus giraudi Redtenbacher , fixed by monotypy.

— Kraatz, 1857: 40 (near Boreaphilus ).

— Kraatz, 1857d: 972 ( Eudectus ; characters; Germany).

— Schaum, 1859: 32 (catalog; European species).

— Thomson, 1861: 184 (characters).

— Gemminger and Harold, 1868: 664 (catalog).

— Jacquelin du Val, 1868: 79 (catalog; European species).

— Fauvel, 1871a: 104 [= 1872: 78] (characters).

— Redtenbacher, 1874: 272 (characters).

— Fauvel, 1875a:V [= 1875b: 207] (catalog).

— Mulsant and Rey, 1880: 398 (characters).

— Duvivier, 1883: 198 (catalog).

— Leconte, 1884: 4 (characters).

— Fowler, 1888: 408 (notes).

— Heyden, Reitter, and Weise, 1891: 118 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 703 (characters).

— Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus).

— Reitter, 1909: 181 (characters; key to German species).

— Bernhauer and Schubert, 1910: 82 (world catalog; 5 species).

— Petri, 1912: 55 (list of species of Siebenbürgen).

— Leng, 1920: 95 (catalog of North American species).

— Winkler, 1925: 336 (catalog for Palaearctic region).

— Roubal, 1930: 319 (catalog; Slovakia).

— Joy, 1932: 104 (key to British species).

— Scheerpeltz, 1933: 1070 (world catalog supplement).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 126 (characters; species of Sweden).

— Tottenham, 1949: 358, 408 (type species: giraudi Redtenbacher ; list of British species).

— Blackwelder, 1952: 155 (type species: giraudi ).

— Tottenham, 1954: 36 (characters; key to British species).

— Székessy, 1963: 23 (characters).

— Lohse, 1964: 68 (characters; key to central European species).

— Scheerpeltz, 1968a: 19 (catalog of Austrian species).

— Steel, 1970a: 22 (larval characters; biological notes).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Moore and Legner, 1975: 189 (catalog; North America north of Mexico).

— Shibata, 1976: 124 (checklist of species of Japan).

— Pope, 1977: 24 (list of British species).

— Topp, 1978: 308 (larval characters).

— Campbell, 1978: 1, 2 (placed in Anthophagini ).

— Burakowski, Mroczkowski, and Stefańska, 1979: 77 (catalog; Poland).

— Moore and Legner, 1979: 164 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Tóth, 1982: 104 (characters; habitus; key to species of Hungary).

— Muona and Viramo, 1986: 16 (list of species of northeastern Finland).

— Zanetti, 1987: 424 (characters; habitus; species of Italy).

— Lohse and Lucht, 1989: 122 (characters).

— Zerche, 1989: 127 (placed in Coryphiini ; characters; notes).

— Watanabe, 1990: 351 (characters; species of Japan).

— Zerche, 1990: 142 (placed in Coryphiini ; characters; key to species of Palaearctic region; distribution; phylogenetic position).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Boháč, 1993: 42 (list of species of Czech Republic and Slovakia).

— Zerche, 1993: 340, 342 (modification of key; phylogeny).

— Zanetti, 1995: 10 (list of species of Italy).

— Lundgren, 1998: 38 (list of species from Florida; records for other States listed for each species).

— Kashcheev, 1999: 144 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (1 Nearctic species; notes; characters in key).

Eudectoides Campbell, 1978: 2 , 11 (species included: crassicornis ). TYPE SPECIES: Eudectus crassicornis LeConte , fixed by original designation.

— Frank, 1986a: 365 (checklist of species of Florida).

— Zerche, 1990: 17, 144 (synonym of Eudectus ).

— Lundgren, 1998: 38 (list of species from Florida; records for other States listed for each species).

altaiensis Zerche, 1990: 155 ( Eudectus ; Type locality: Altai).

- DISTRIBUTION: Altai region.

crassicornis LeConte, 1884: 4 ( Eudectus ; Type locality: Opelousas, La.).

— Bernhauer and Schubert, 1910: 82 ( Eudectus ; catalog).

— Campbell, 1978: 12 ( Eudectoides ; characters; Florida; Louisiana).

— Zerche, 1990: 299 ( Eudectus ; note).

— Zerche, 1993: 342 ( Eudectus ; characters; phylogenetic position; USA).

— Lundgren, 1998: 38 ( Eudectus ; Florida).

- DISTRIBUTION: USA.

gerhardti Pietsch, 1894 , see: giraudi Redtenbacher, 1857 .

giraudi Redtenbacher, 1857: 246 ( Eudectus ; Type locality: bei München).

— Kraatz, 1857d: 973 ( Eudectus ; characters; Germany).

— Thomson, 1861: 185 ( Eudectus ; characters; Scandinavia).

— Fauvel, 1871a: 105 [= 1872: 79] ( Eudectus ; characters; Scandinavia; Germany; Switzerland; Britain).

— Redtenbacher, 1874: 273 ( Eudectus ; characters).

— Seidlitz, 1875: 242 ( Coryphium ; characters; Baltic region).

— Mulsant and Rey, 1880: 400 ( Eudectus ; characters; France).

— Seidlitz, 1891a: 372 ( Eudectus ; characters; Europe).

— Ganglbauer, 1895: 703 ( Eudectus ; characters; Germany; Switzerland; Scotland; Sweden).

— Reitter, 1909: 181 ( Eudectus ; characters; Germany).

— Bernhauer and Schubert, 1910: 83 ( Eudectus ; catalog).

— Scheerpeltz, 1933: 1070 ( Eudectus ; catalog).

— Palm, 1948: 127 ( Eudectus ; characters; Sweden; Norway; Finland).

— Horion, 1963: 150 ( Eudectus ; Altai region; Finland; Sweden; Norway; Britain; Germany; Czech Republic; Slovakia; Poland; Hungary; Romania; France; Switzerland; Italy; Austria).

— Lohse, 1964: 68 ( Eudectus ; characters; central Europe).

— Tóth, 1982: 104 ( Eudectus ; characters).

— Zanetti, 1987: 426 ( Eudectus ; characters; Italy).

— Lucht, 1987: 90 ( Eudectus ; checklist; Germany; Poland; Czechoslovakia; Austria; France).

— Zerche, 1990: 147 ( Eudectus ; characters; phylogenetic position; Norway; Sweden; Finland; Russia; Germany; Poland; Czechoslovakia; Romania; Switzerland; Italy; Austria).

— Zerche, 1993: 353 ( Eudectus ; Poland; Austria).

— Boháč, 1993: 42 ( Eudectus ; Slovakia; Czech Republic).

— Zanetti, 1995: 10 ( Eudectus ; Italy).

- DISTRIBUTION: Europe, Russia.

gerhardti Pietsch, 1894: 17 ( Eudectus ; Type locality: Reinerz in Schlesien).

— Ganglbauer, 1895: 703 ( Eudectus ; characters; Schlesien).

— Reitter, 1909: 181 ( Eudectus ; characters; Germany).

— Bernhauer and Schubert, 1910: 82 ( Eudectus ; catalog).

— Scheerpeltz, 1933: 1070 ( Eudectus ; catalog).

— Lohse, 1964: 68 ( Eudectus ; characters; central Europe).

— Lucht, 1987: 91 ( Eudectus ; checklist; Poland; Czechoslovakia).

— Zerche, 1989: 127 ( Eudectus ; synonym of giraudi ).

— Zerche, 1990: 148 ( Eudectus ; synonym of giraudi ).

— Boháč, 1993: 42 ( Eudectus ; cited as valid species; Slovakia?).

kulczynskii Rybiński, 1902: 4 ( Eudectus ; Type locality: Tatra Gebirge).

— Bernhauer and Schubert, 1910: 83 ( Eudectus ; catalog).

— Scheerpeltz, 1933: 1070 ( Eudectus ; synonym of giraudi ).

— Tóth, 1982: 105 ( Eudectus ; cited as valid species; characters).

— Zerche, 1990: 148 ( Eudectus ; synonym of giraudi ).

— Boháč, 1993: 42 ( Eudectus ; cited as valid species; Slovakia).

japonicus Zerche, 1990: 157 ( Eudectus ; Type locality: Japan: Hokkaido: Kotani ).

- DISTRIBUTION: Japan.

kulczynskii Rybiński, 1902 , see: giraudi Redtenbacher, 1857 .

novayaezemlyae Munster, 1925, see: whitei Sharp, 1871 .

reductus Zerche, 1993: 341 ( Eudectus ; Type locality: o. Vrangelja [ostrov Vrangelja: Wrangel- Insel]).

— Zerche, 1993: 344 ( Eudectus ; phylogenetic position).

— Ryabukhin, 1999: 30 ( Eudectus ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Wrangel Island.

rufulus Weise, 1877: 96 ( Eudectus ; variety of giraudi ; Type locality: Hagi).

— Weise, 1884a: 271 ( Eudectus ; cited as species; characters; Japan).

— Sharp, 1889: 473 ( Eudectus ; cited as species; Japan).

— Bernhauer and Schubert, 1910: 83 ( Eudectus ; catalog).

— Nakane, 1963a: 83 ( Eudectus ; characters; habitus photograph; Japan).

— Watanabe, 1990: 353 ( Eudectus ; characters; Japan).

— Zerche, 1990: 146 ( Eudectus ; characters; phylogenetic position; Japan).

— Zerche, 1993: 343 ( Eudectus ; phylogenetic position).

- DISTRIBUTION: Japan.

whitei Sharp, 1871b: 73 ( Eudectus ; Type locality: Britain: Ben-a-Bhuird).

— Fowler, 1888: 409 ( Eudectus ; characters; Britain).

— Ganglbauer, 1895: 703 ( Eudectus ; variety of giraudi ).

— Bernhauer and Schubert, 1910: 83 ( Eudectus ; variety of giraudi ).

— Scheerpeltz, 1933: 1070 ( Eudectus ; catalog).

— Steel, 1970a: 22 ( Eudectus ; larval characters; biological notes).

— Pope, 1977: 24 ( Eudectus ; Britain).

— Zerche, 1990: 153 ( Eudectus ; characters; phylogenetic position; Britain; Russia).

— Zerche, 1993: 344 ( Eudectus ; phylogenetic position).

— Ryabukhin, 1999: 30 ( Eudectus ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Europe.

novayaezemlyae Munster, 1925: 6 ( Eudectus ; Type locality: Novaja Semlja, Ökland; [Note: See lectotype designation by Zerche, 1990]. Also cited from Mashigin Fj.; Gribovii Fj.).

— Scheerpeltz, 1933: 1070 ( Eudectus ; catalog).

— A. Strand, 1951: 104 ( Eudectus ; cited as novayae zemlyae; synonym of whitei ).

— Pope, 1977: 24 ( Eudectus ; synonym of whitei ).

— Zerche, 1990: 153 ( Eudectus ; lectotype designation: Novaj. Semlja; synonym of whitei ).

Haida

[4 species; Nearctic Region]

Haida Keen, 1897: 285 (species included: keeni ). TYPE SPECIES: Haida keeni Keen , fixed by monotypy.

— Bernhauer and Schubert, 1910: 50 (world catalog; 1 species).

— Leng, 1920: 94 (catalog of North American species).

— Blackwelder, 1952: 179 (type species: keeni ).

— Hatch, 1957: 51, 73 (characters; species of Pacific Northwest).

— Moore and Legner, 1975: 192 (catalog; North America north of Mexico).

— Campbell, 1978: 3 (key to species; characters; should be placed in Anthophagini ).

— Moore and Legner, 1979: 165 (characters; notes).

— Zerche, 1990: 250, 261 (removal from Coryphiini by Campbell, 1978, unjustified).

— Newton and Thayer, 1995: 294 (should be in Coryphiini ; list of New World genera).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (4 Nearctic species; notes; characters in key). bisulcata Campbell, 1978: 7 ( Haida ; Type locality: Cal.: El Dorado Co: Desol. Valley, Primitive Area below Eagle Lake, 6700 ft.).

- DISTRIBUTION: USA.

callosum Mäklin, 1852: 323 ( Omalium ; Type locality: insula Sitkha).

— Fauvel, 1878d: 208 [= 1878e: 44] ( Omalium , cited as Homalium ; characters; Alaska).

— Bernhauer and Schubert, 1910: 52 ( Omalium ; subgenus Omalium ; catalog).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 ( Haida ).

- DISTRIBUTION: USA.

insulcata Campbell, 1978: 8 ( Haida ; Type locality: Oregon: Wasco Co.: 11. 1 mi. SE jct.

US 26, Ore. 216, 2500 ft.).

- DISTRIBUTION: USA.

keeni Keen, 1897: 285 ( Haida ; Type locality: Massett, Queen Charlotte Island ).

— Bernhauer and Schubert, 1910: 50 ( Haida ; catalog).

— Brown, 1944: 5 ( Haida ; characters; notes; British Colombia).

— Hatch, 1957: 73 ( Haida ; characters; British Columbia ; Washington; Oregon).

— Campbell, 1978: 4 ( Haida ; characters; British Columbia ; Alaska; Oregon; Washington).

— Campbell and Davies, 1991: 91 ( Haida ; checklist of species; Alaska and Canada).

- DISTRIBUTION: Canada, USA.

Holoboreaphilus

[1 species; Nearctic and Palaearctic Regions]

Holoboreaphilus Campbell, 1978 a: 54 (species included: nordenskioldi ). TYPE SPECIES: Boreaphilus nordenskioldi Mäklin , fixed by original designation.

— Zerche, 1990: 96 (characters; subfossil records; distribution; characters analysis; phylogenetic relationships).

— Downie and Arnett, 1996: 436 (characters; species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (1 Nearctic species; notes; characters in key).

nearcticus Blair, 1933 , see: nordenskioldi Mäklin, 1878 .

nordenskioldi Mäklin, 1878: 25 [= 1878a: 301] ( Boreaphilus ; Type locality: Insel Sopotschnoj , Sibir. Jeniseie, 70°5′ N; [Note: See lectotype designation by Zerche, 1990]. Originally cited from Mesenkin, 71°20′ n. lat; Briachowskij; Dudino, 69°15′ n. lat; Sopotschnoj, 70°5′ n. lat.).

— J. Sahlberg, 1880: 106 ( Boreaphilus ; characters; Siberia).

— Mäklin, 1881: 42 ( Boreaphilus ; characters; Mesenkin, lat. bor. 71°20′; Dudino, lat. bor. 69°15′).

— Poppius, 1909: 6 ( Boreaphilus ; Lena Valley, Siberia ).

— Bernhauer and Schubert, 1910: 84 ( Boreaphilus ; catalog).

— Munster, 1925: 7 ( Boreaphilus ; Novaya Zemlya).

— Scheerpeltz, 1933: 1072 ( Boreaphilus ; catalog).

— Steel, 1964a: 250 ( Boreaphilus ; Canada; Alaska).

— Campbell, 1978a: 55 ( Holoboreaphilus ; characters; Northwest Territories; Quebec; Yukon Territory; Alaska).

— Zerche, 1990: 97 ( Holoboreaphilus ; lectotype designation: Ins. Sopotschn. Sibir. Jeniseie; correction of spelling of name; characters; Russia) .

— Elias, 1991: 556 ( Holoboreaphilus ; Quaternary fossil collection; Montana).

— Campbell and Davies, 1991: 89 ( Holoboreaphilus ; checklist of species; Alaska and Canada).

— Zerche, 1993: 353 ( Holoboreaphilus ; Alaska).

— Downie and Arnett, 1996: 436 ( Holoboreaphilus ; characters; Canada; USA).

— Ryabukhin, 1999: 33 ( Holoboreaphilus ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Russia, Canada, USA.

nearcticus Blair, 1933: 95 ( Boreaphilus ; Type locality: Akpatok , Island, Ungava Bay).

— Steel, 1964a: 250 ( Boreaphilus ; synonym of nordenskioldi ).

— Campbell, 1978a: 55 ( Holoboreaphilus ; synonym of nordenskioldi ).

— Zerche, 1990: 97 ( Holoboreaphilus ; synonym of nordenskioldi ).

Murathus

[1 species; Palaearctic Region]

Murathus Kashcheev, 1999: 144 (species included: montanus ). TYPE SPECIES: Murathus montanus Kashcheev , fixed by original designation and monotypy.

montanus Kashcheev, 1999: 146 ( Murathus ; Type locality: Tian’shan’, Khr. Kungei Alatau, ushchel’e Kul’bastau, 2500-3200 m).

- DISTRIBUTION: Kazakhstan.

Niphetodes

[5 species; Palaearctic Region] Niphetodes Miller, 1868: 16 (species included: redtenbacheri ). TYPE SPECIES: Niphetodes redtenbacheri Miller , fixed by monotypy.

— Redtenbacher, 1874: 550 (characters).

— Fauvel, 1875a:V [= 1875b: 207] (catalog).

— Duvivier, 1883: 198 (catalog).

— Heyden, Reitter, and Weise, 1891: 118 (subgenus of Boreaphilus ; list of species of Europe and Caucasus).

— Ganglbauer, 1896: 174 (characters; notes; key to species).

— Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 85 (world catalog; 7 species).

— Petri, 1912: 56 (list of species of Siebenbürgen).

— Winkler, 1925: 337 (catalog for Palaearctic region).

— Roubal, 1930: 320 (catalog; Slovakia).

— Scheerpeltz, 1933: 1073 (world catalog supplement).

— Blackwelder, 1952: 262 (type species: redtenbacheri ).

— Horion, 1963: 156 (list of central European species).

— Székessy, 1963: 24 (characters).

— Lohse, 1964: 70 (characters).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Burakowski, Mroczkowski, and Stefańska, 1979: 79 (catalog; Poland).

— Tóth, 1982: 108 (characters; habitus; key to species of Hungary).

— Lohse and Lucht, 1989: 123 (characters).

— Zerche, 1989: 129 (complex of 3 genera; not in central Europe).

— Zerche, 1990: 101 (characters; key to species; characters analysis; phylogenetic relationships; distribution).

— Zerche, 1993: 335 (phylogenetic relationships).

— Kashcheev, 1999: 146 (characters).

apfelbecki Ganglbauer, 1892 , see: Pareudectus .

cyrneus Orousset and Lecoq, 1988 , see: Niphetodops .

deubeli Ganglbauer, 1896 , see: Hypsonothrus .

eppelsheimi Ganglbauer, 1896 , see: Pareudectus .

huppenthali Lokay, 1912, see: Hypsonothrus .

kreissli Zerche, 1990: 104 ( Niphetodes ; Type locality: Buleasee [ Bîlea Lac]).

- DISTRIBUTION: Romania.

leonhardi Bernhauer, 1905 , see: Pareudectus .

redtenbacheri Miller, 1868: 17 ( Niphetodes ; Type locality: Czerna hora; [Note: See lectotype designation by Zerche, 1990]).

— Redtenbacher, 1874: 550 ( Niphetodes ; characters).

— Ganglbauer, 1895: 700 ( Boreaphilus ; subgenus of Niphetodes ; characters; east Carpathians). — Ganglbauer, 1896: 177 ( Niphetodes ; characters; notes).

— Bernhauer and Schubert, 1910: 85 ( Niphetodes ; catalog).

— Scheerpeltz, 1933: 1073 ( Niphetodes ; catalog).

— Horion, 1963: 156 ( Niphetodes ; Carpathians).

— Tóth, 1982: 110 ( Niphetodes ; characters).

— Zerche, 1990: 110 ( Niphetodes ; lectotype designation; characters; Ukrainian Carpathians).

- DISTRIBUTION: Ukrainian Carpathians.

sahlbergi Apfelbeck, 1906 , see: Pareudectus .

schoenmanni Zerche, 1990: 113 ( Niphetodes ; Type locality: Rodnaergebirge).

- DISTRIBUTION: Romania.

semicarinatus Zerche, 1990: 106 ( Niphetodes ; Type locality: Rodnaer - Gebirge).

- DISTRIBUTION: Romania.

spaethi Ganglbauer, 1900: 259 ( Niphetodes ; Type locality: Negoi [Note: See lectotype designation by Zerche, 1990]. Also cited from Siebenbürgen. Rodnaergebirge, Černahora). — Bernhauer and Schubert, 1910: 85 ( Niphetodes ; catalog).

— Horion, 1963: 156 ( Niphetodes ; Romania; Slovakia).

— Tóth, 1982: 110 ( Niphetodes ; characters).

— Zerche, 1990: 108 ( Niphetodes ; lectotype designation: Negoi ; characters; Romania) .

- DISTRIBUTION: Romania.

Occiephelinus

[1 species; Nearctic Region]

Occiephelinus Hatch, 1957: 52 , 75 (species included: vandykei ). TYPE SPECIES: Occiephelinus vandykei Hatch , fixed by original designation and monotypy.

— Moore and Legner, 1975: 198 (catalog; North America north of Mexico).

— Campbell, 1978a: 26 (synonym of Coryphium ).

— Moore and Legner, 1979: 166 (characters; notes).

— Watanabe, 1988: 46 (synonym of Coryphium ).

— Watanabe, 1990: 355 (synonym of Coryphium ).

— Zerche, 1990: 31 (valid genus; removed from synonym with Coryphium ).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (1 Nearctic species; notes; characters in key). vandykei Hatch, 1957: 75 ( Occiephelinus ; Type locality: Washington: Mt. Rainier, Paradise

Park, alt. 6000 ft.).

— Campbell, 1978a: 31 ( Coryphium ; characters; Oregon; Washington).

- DISTRIBUTION: USA.

Ophthalmoniphetodes

[15 species; Palaearctic Region]

Ophthalmoniphetodes Zerche, 1990: 115 (species included: heidemariae , doeblerae , uhligi , behnei , ilievi , macrocephalus , musalensis , rhilensis , longicornis , gracilis ). TYPE SPECIES: Ophthalmoniphetodes longicornis Zerche , fixed by original designation.

— Zerche, 1993: 336, 339 (modification of key; phylogeny).

behnei Zerche, 1990: 123 ( Ophthalmoniphetodes ; Type locality: Sofia, Vitoscha-Gebirge, Malak Resen, 2000 m).

— Zerche, 1993: 353 ( Ophthalmoniphetodes ; Bulgaria).

- DISTRIBUTION: Bulgaria.

doeblerae Zerche, 1990: 121 ( Ophthalmoniphetodes ; Type locality: SW-Bulg.: Rila-Gebirge, Hütte Maljovica, 2100 m).

- DISTRIBUTION: Bulgaria.

falakroensis Assing and Wunderle, 1999 a: 477 ( Ophthalmoniphetodes ; Type locality:

GR. [NE], Makedhonia, Dráma, Falakró, 1800 m).

- DISTRIBUTION: Greece.

gracilis Zerche, 1990: 131 ( Ophthalmoniphetodes ; Type locality: Bulgarien: Pirin-Geb., Hütte Wichren , 2050 m).

- DISTRIBUTION: Bulgaria.

heidemariae Zerche, 1990: 119 ( Ophthalmoniphetodes ; Type locality: Bulgaria: Pirin-Gebirge , Hütte Wichren, 2050 m).

- DISTRIBUTION: Bulgaria.

ilievi Zerche, 1990: 124 ( Ophthalmoniphetodes ; Type locality: Sofia, Vitoscha-Geb. bei Aleko , 2000 m).

- DISTRIBUTION: Bulgaria.

longicornis Zerche, 1990: 130 ( Ophthalmoniphetodes ; Type locality: Bulgarien: Pirin-Gebirge , Hütte Wichren, 2050 m).

- DISTRIBUTION: Bulgaria.

macrocephalus Zerche, 1990: 126 ( Ophthalmoniphetodes ; Type locality: Bulgarien: Pirin-Gebirge , Hütte Wichren, 2050 m).

- DISTRIBUTION: Bulgaria.

maljovicensis Zerche, 1993: 338 ( Ophthalmoniphetodes ; Type locality: Bulg., Rila-Geb., Umg. Hütte Maljovica, 2100 m).

- DISTRIBUTION: Bulgaria.

musalensis Zerche, 1990: 127 ( Ophthalmoniphetodes ; Type locality: SW-Bulgarien, Rilapangeoensis Assing and Wunderle, 1999a: 478 ( Ophthalmoniphetodes ; Type locality:

GR. [NE], Makedhonia, NW Kavála, Pangéo, 1900 m).

- DISTRIBUTION: Greece.

piger Zerche, 1993: 337 ( Ophthalmoniphetodes ; Type locality: SW-Bulg., Rila-Geb., Hütte Musala, 2300 m).

- DISTRIBUTION: Bulgaria.

rhilensis Zerche, 1990: 128 ( Ophthalmoniphetodes ; Type locality: SW-Bulgarien, Rila-Gebirge, Umgebung Granschar, 2185 m).

- DISTRIBUTION: Bulgaria.

uhligi Zerche, 1990: 122 ( Ophthalmoniphetodes ; Type locality: Bulgarien: Sitnjakowo [Schloss bei Borovez], 1730 m, Tschetér Tepé [Tschadjar Tepe]).

- DISTRIBUTION: Bulgaria.

zerchei Assing and Wunderle, 1999 a: 475 ( Ophthalmoniphetodes ; Type locality: GR. [NE], Makedhonia, Dráma, Falakró, 1800 m).

- DISTRIBUTION: Greece.

Pareudectus

[3 species; Palaearctic Region]

Pareudectus Zerche, 1990: 132 (species included: leonhardi , apfelbecki , eppelsheimi ). TYPE SPECIES: Niphetodes apfelbecki Ganglbauer , fixed by original designation.

apfelbecki Ganglbauer, 1892: 17 ( Niphetodes ; Type locality: Sarajevo; ca. 1500 m; [Note: See lectotype designation by Zerche, 1990]).

— Ganglbauer, 1895: 701 ( Boreaphilus ; subgenus Niphetodes ; characters; Bosnia).

— Ganglbauer, 1896: 177, 178 ( Niphetodes ; characters; notes).

— Bernhauer and Schubert, 1910: 85 ( Niphetodes ; catalog).

— Zerche, 1990: 136 ( Pareudectus ; lectotype designation; characters; Yugoslavia) .

— Zerche, 1993: 353 ( Pareudectus ; Bosnia-Hercegovina).

- DISTRIBUTION: Bosnia Hercegovina.

eppelsheimi Ganglbauer, 1896: 177 , 178 ( Niphetodes ; Type locality: Rodnaer Alpen, Gipfel des Koronjis; [Note: See lectotype designation by Zerche, 1990]).

— Bernhauer and Schubert, 1910: 85 ( Niphetodes ; catalog).

— Horion, 1963: 156 ( Niphetodes ; Slovakia).

— Tóth, 1982: 109 ( Niphetodes ; characters).

— Zerche, 1990: 139 ( Pareudectus ; lectotype designation; characters; USSR; Romania) .

- DISTRIBUTION: Romania, Ukraine.

leonhardi Bernhauer, 1905b: 588 ( Niphetodes ; Type locality: Bjelašnica-Planina an der Grenze Bosniens und der Herzegowina; [Note: See lectotype designation by Zerche, 1990]).

— Bernhauer and Schubert, 1910: 85 ( Niphetodes ; catalog).

— Scheerpeltz, 1933: 1073 ( Niphetodes ; catalog).

— Zerche, 1990: 134 ( Pareudectus ; lectotype designation; characters; Yugoslavia) .

- DISTRIBUTION: Bosnia Hercegovina.

sahlbergi Apfelbeck, 1906: 247 ( Niphetodes ; Type locality: Bosna: palnini Ljubični na jugoistoku Bosne na turskoj megji na Treskovici kod Sarajeva [Note: See lectotype designation by Zerche, 1990]).

— Apfelbeck, 1907: 641 ( Niphetodes ; Bosnia).

— Bernhauer and Schubert, 1910: 85 ( Niphetodes ; catalog).

— Zerche, 1990: 134 ( Pareudectus ; lectotype designation: Trescavica pl; synonym of leonhardi ).

winkleri Bernhauer, 1929 g: 180 ( Niphetodes ; Type locality: Orjengebirge in der nördlichen Herzegowina).

— Scheerpeltz, 1933: 1073 ( Niphetodes ; catalog).

— Zerche, 1990: 134 ( Pareudectus ; synonym of leonhardi ).

Planeboreaphilus

[1 species; Palaearctic Region]

Planeboreaphilus Shibata, 1970: 58 (species included: coriaceoides ). TYPE SPECIES: Planeboreaphilus coriaceoides Shibata , fixed by original designation and monotypy.

— Shibata, 1976: 126 (checklist of species of Japan).

— Campbell, 1978a: 26 (synonym of Coryphium ).

— Watanabe, 1988: 46 (synonym of Coryphium ).

— Watanabe, 1990: 355 (cited as Plaeboreaphilus; synonym of Coryphium ).

— Zerche, 1990: 84 (characters; cited as valid genus; species of Palaearctic region; character analysis; phylogenetic relationships).

coriaceoides Shibata, 1970: 59 ( Planeboreaphilus ; Type locality: Japan: Yamanashi Pref.: near Hirogawara, 2400 m. alt., South Alps ).

— Campbell, 1978a: 26 ( Coryphium ; [Note: Not explicitly transferred by Campbell but done so implicitly because it is the type species of Planeboreaphilus which he synonymized with Coryphium ]).

— Watanabe, 1988: 60 ( Coryphium ; characters; Japan).

— Watanabe, 1990: 369 ( Coryphium ; characters; Japan).

— Zerche, 1990: 85 ( Planeboreaphilus ; characters; Japan).

- DISTRIBUTION: Japan.

Platycoryphium

[1 species; Palaearctic Region]

Platycoryphium Zerche, 1990: 26 (species included: parvulum ). TYPE SPECIES: Coryphium parvulum Poppius , fixed by original designation and monotypy.

parvulum Poppius, 1904: 107 ( Coryphium ; Type locality: Lena-Gebiet in Nordost-Sibirien: Aldan-Mündung).

— Poppius, 1909: 6 ( Coryphium ; Lena Valley, Siberia ).

— Bernhauer and Schubert, 1910: 84 ( Coryphium ; catalog).

— Tikhomirova, 1973a: 141 ( Coryphium ; species uncertain; checklist; northern Siberia).

— Zerche, 1990: 27 ( Platycoryphium ; characters; Russia).

— Ryabukhin, 1999: 32 ( Platycoryphium ; catalog; habitat; distribution; northeastern Russia). - DISTRIBUTION: Russia.

Pseudohaida

[1 species; Nearctic Region]

Pseudohaida Hatch, 1957: 51 , 73 (species included: rothi , ingrata ). TYPE SPECIES: Pseudohaida rothi Hatch , fixed by original designation.

— Moore and Legner, 1975: 213 (catalog; North America north of Mexico).

— Campbell, 1978: 3, 9 (moved to Anthophagini ; key to species; characters).

— Moore and Legner, 1979: 167 (characters; notes).

— Zerche, 1900: 250, 261 (character discussion; removal from Coryphiini by Campbell, 1978,

unjustified).

— Newton, Thayer, Ashe, and Chandler, 2000: 341 (1 Nearctic species; notes; characters in key). ingrata Hatch, 1957 , see: Subhaida .

rothi Hatch, 1957: 74 ( Pseudohaida ; Type locality: Oregon: Corvallis , watershed).

— Campbell, 1978: 10 ( Pseudohaida ; characters; Oregon).

— Campbell and Winchester, 1993: 83 ( Pseudohaida ; British Columbia ).

- DISTRIBUTION: Canada, USA.

Subhaida

[7 species; Nearctic Region]

Subhaida Hatch, 1957: 52 , 74 (species included: rainieri ). TYPE SPECIES: Subhaida rainieri

Hatch, fixed by original designation and monotypy.

— Moore and Legner, 1975: 215 (catalog; North America north of Mexico).

— Campbell, 1978a: 24, 39 (key to species; characters; notes).

— Moore and Legner, 1979: 168 (characters; notes).

— Zerche, 1900: 20 (characters discussed).

— Campbell and Davies, 1991: 89 (checklist of species; Alaska and Canada).

— Newton, Thayer, Ashe, and Chandler, 2000: 342 (7 Nearctic species; notes; characters in key). aptera Campbell, 1978 a: 44 ( Subhaida ; Type locality: Colorado: Delta Co. : 16 mi. N. Cedaredge, 6 mi. E. Hwy 65, nr. Peak Reservoir, 10,000 ft.).

- DISTRIBUTION: USA.

californica Campbell, 1978 a: 42 ( Subhaida ; Type locality: California: Mono Co. : 6 miles SW Toms Place, 9000 ft.).

- DISTRIBUTION: USA.

ingrata Hatch, 1957: 74 ( Pseudohaida ; Type locality: Oregon: Corvallis, watershed. Also cited from British Columbia ).

— Campbell, 1978a: 46 ( Subhaida ; characters; Alberta; British Columbia ; Alaska; California; Oregon; Washington).

- DISTRIBUTION: Canada, USA.

monticola Campbell, 1978 a: 45 ( Subhaida ; Type locality: Alberta: Consolation Lk., Banff

Nat. Pk., 64–7000 ft.).

- DISTRIBUTION: Canada.

rainieri Hatch, 1957: 74 ( Subhaida ; Type locality: Washington: Mt. Rainier, Paradise Park , 6000 ft.).

— Campbell, 1978a: 40 ( Subhaida ; characters; Oregon; Washington).

- DISTRIBUTION: USA.

sinuata Campbell, 1978 a: 49 ( Subhaida ; Type locality: Oregon: Klamath Co.: 9 mi. NE Bly, Deming Creek, 5000–5800 ft. Also cited from California).

- DISTRIBUTION: USA.

utahensis Campbell, 1978 a: 48 ( Subhaida ; Type locality: Utah: Cache Co.: Logan Canyon ).

- DISTRIBUTION: USA.

Tribe Eusphalerini

Eusphalerini Hatch, 1957: 82 (genera included: Eusphalerum , Onibathum ). TYPE GENUS: Eusphalerum (see Newton and Thayer, 1992: 58).

— Arnett, 1963: 242, 261 (key to North American genera; characters; list of North American genera).

— Moore and Legner, 1979: 170 (characters).

— Muona, 1979: 17 (list of Scandinavian species).

— Watanabe, 1990: 179, 377 (characters; key to genera of Japan; relationships of genera).

— Newton and Thayer, 1992: 58 (tribe of Omaliinae ).

— Newton and Thayer, 1995: 247–286, 294 (phylogenetic relationships; characters; includes only Eusphalerum ).

— M. Hansen, 1996: 94 (list of Danish species).

— Newton, Thayer, Ashe, and Chandler, 2000: 338 (characters).

Anthobiates Mulsant and Rey, 1880: 290 (genera included: Eusphalerum , Anthobium ). TYPE GENUS: Anthobium (misidentification; see Newton and Thayer, 1992: 58).

— Newton and Thayer, 1992: 58 (synonym of Eusphalerini ).

Anthobiini Portevin, 1929: 450 (genera included: Anthobium ). TYPE GENUS: Anthobium (misidentification; see Newton and Thayer, 1992: 58).

— Joy, 1932: 95 (key to British genera).

— Outerelo and Gamarra, 1985: 85 (key to genera of Iberian Peninsula).

— Newton and Thayer, 1992: 58 (synonym of Eusphalerini ).

Eusphalerum

[227 species; Nearctic, Palaearctic , and Oriental Regions]

Eusphalerum Kraatz, 1857d: 1003 (species included: triviale ; oblitum doubtfully included). TYPE SPECIES: Anthobium triviale Erichson , fixed by monotypy.

— Erichson, 1839a: 637 (characters; [Note: Misidentified as Anthobium according to Blackwelder, 1952: 56]).

— Erichson, 1840: 890 (characters; [Note: Misidentified as Anthobium according to Blackwelder, 1952: 56]).

— Schaum, 1852: 30 (cited as Anthobium ; list of species; Europe).

— Fairmaire and Laboulbène, 1856: 646 (misidentified as Anthobium ; characters).

— Gistel, 1856: 267, 390 (list of species; cited as Anthobium ).

— Redtenbacher, 1857: 254 (misidentified as Anthobium ; characters; key to Austrian species).

— Redtenbacher, 1857: 995 (characters).

— G. Waterhouse, 1858: 32 (catalog of British species).

— Schaum, 1859: 32 (catalog; European species).

— Jacquelin du Val, 1868: 81 (catalog; European species).

— LeConte, 1861a: 70 (cited as Anthobium ; characters).

— Gredler, 1863: 125 (list of species from Tirol).

— LeConte, 1863: 26 (cited as Anthobium ; list of North American species).

— Wencker and Silbermann, 1866: 36 (list of species; collecting notes; France).

— Wencker and Silbermann, 1866: 37 (cited as Anthobium ; list of species; collecting notes; France).

— Gemminger and Harold, 1868: 668 (catalog).

— Gemminger and Harold, 1868: 669 (catalog; cited as Anthobium ).

— Fauvel, 1871a: 60 [= 1872: 34] (characters; key to species).

— Sharp, 1871d: 14 (list of British species; cited both as Eusphalerum and Anthobium ).

— Redtenbacher, 1874: 278 (characters).

— Fauvel, 1875a: III [= 1875b: 205] (synonym of Anthobium ).

— Fauvel, 1878d: 198 [= 1878e: 34] (cited as Anthobium ; key to North American species).

— Mulsant and Rey, 1880: 290 (characters; cited as Anthobium ; key to French species).

— Heyden, 1880: 83 (cited as Anthobium ; list of species of Siberian region).

— Duvivier, 1883: 200 (catalog; cited as Anthobium and Eusphalerum ).

— Fowler, 1888: 424 (notes).

— Heyden, Reitter, and Weise, 1891: 121 (cited as Anthobium ; list of species of Europe and Caucasus).

— Casey, 1894: 426 (cited as Anthobium ; notes; key to some North American species).

— Seidlitz, 1891: 346 (cited as Anthobium with Eusphalerum as subgenus; key to some European species).

— Seidlitz, 1891a: 368 (cited as Anthobium with Eusphalerum as subgenus; key to some European species).

— Heyden, 1893: 49 (cited as Anthobium ; list of species of Siberian region).

— Ganglbauer, 1895: 745 (cited as Anthobium ; characters; key to middle European species).

— Fauvel, 1897d: 243 (catalog of species of Barbary; cited as Anthobium ).

— Heyden, Reitter, and Weise, 1906: 128 (cited as Anthobium ; list of species of Europe and Caucasus).

— Sainte-Claire Deville, 1907: 21 (characters; key to species of Seine Basin, France).

— Reitter, 1909: 195 (cited as Anthobium ; characters; key to German species).

— Bernhauer and Schubert, 1910: 37 (world catalog; cited as Anthobium with Eusphalerum as subgenus; 95 species).

— Blatchley, 1910: 481 (characters; cited as Anthobium ).

— Petri, 1912: 51 (cited as Anthobium ; list of species of Siebenbürgen).

— Johansen, 1914: 621 (characters; cited as Anthobium ; key to Danish species).

— Winkler, 1925: 326 (catalog for Palaearctic region; cited as Anthobium ).

— Cameron, 1925: 10 (catalog of Indian species; cited as Anthobium ).

— Porta, 1926: 12 (cited as Anthobium ; characters; key to species of Italy).

— Cameron, 1930: 134 (cited as Anthobium ; characters; key to British Indian species).

— Joy, 1932: 96 (cited as Anthobium ; key to British species).

— Joy, 1932: 103 (key to British species).

— Scheerpeltz, 1933: 1032 (synonym of Anthobium ).

— Normand, 1935: 357 (cited as Anthobium ; list of species of Tunisia).

— C. Koch, 1938a: 324 (cited as Anthobium ; key to European species).

— Tottenham, 1939d: 225 ( Eusphalerum misidentified as Anthobium by previous authors).

— Kloet and Hincks, 1945: 160 (list of British species; cited with Anthobium of authors as synonym; type species: primulae ).

— Palm, 1948: 56 (cited as Anthobium ; characters; key to species of Sweden).

— Tottenham, 1949: 354, 405 (type species: primulae Stephens = triviale Erichson ; list of British species; notes; Anthobium of authors is synonym).

— Tottenham, 1954: 18 (characters; key to British species).

— Hatch, 1957: 52, 82 (characters; key to species of Pacific Northwest).

— Freude, 1957: 102 (cited as Anthobium ; key to some species of Germany).

— Kocher, 1958: 93 (cited as Anthobium ; checklist of species; Morocco).

— Coiffait, 1959c (subgeneric and species group classification; key to species of France and nearby regions; list of species).

— Ferreira, 1962a: 9 (cited as Anthobium ; list of Portuguese species).

— Horion, 1963: 32 (cited as Anthobium ; list of central European species).

— Székessy, 1963: 12 (cited as Anthobium ; characters).

— Lohse, 1964: 29 (characters; key to central European species).

— Scheerpeltz, 1968a: 5 (cited as Anthobium ; catalog of Austrian species).

— Steel, 1970a: 8 (larval characters; biological notes).

— Shibata, 1973c: 28 (checklist of species of Taiwan).

— Tikhomirova, 1973a: 137 (checklist of species of USSR).

— Moore and Legner, 1975: 190 (catalog; North America north of Mexico).

— Scheerpeltz, 1976a: 99 (cited as Anthobium ; key to species of Himalayan region).

— Shibata, 1976: 98 (checklist of species of Japan).

— Pope, 1977: 23 (list of British species).

— Klinger and Maschwitz, 1977: 402 (morphology and chemical secretions of defensive glands).

— Topp, 1978: 309 (larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 16 (catalog; Poland).

— Moore and Legner, 1979: 170 (characters; notes).

— Muona, 1979: 17 (list of Scandinavian species).

— Uhlig, 1979: 242 (some species collected in Germany).

— Klinger, 1980 (comparative morphology of abdominal defensive gland and discussion of subgeneric classification).

— Uhlig, Vogel, and Sieber, 1980: 240 (some species collected in Germany).

— Spahr, 1981: 94 (references to specimens from amber and copal).

— Uhlig and Vogler, 1981: 84 (list of some species collected in Germany; habitat and collecting notes).

— Zanetti, 1982 (revision of subgenera and species Italy and central Europe; discussion of female genitalic characters).

— Tóth, 1982: 35 (characters; habitus; key to species of Hungary).

— Klinger, 1983 (natural history notes; Germany).

— Muona and Viramo, 1986: 15 (list of species of northeastern Finland).

— Segers, 1986: 18 (checklist of species; Belgium).

— Dettner, 1987: 28 (defensive glands).

— Zanetti, 1987: 46 (characters; habitus; species group classification; key to species of Italy in English and Italian).

— Lohse and Lucht, 1989: 123 (supplement for key to species of Lohse, 1964).

— Watanabe, 1990: 179 (characters; key to species of Japan).

— Dettner and Reissenweber, 1991 (defensive secretions: chemistry and systematics).

— Angelini, 1991: 191 (list of species from high elevations of Calabria, Italy). — Zanetti, 1991: 291 (key to species of Soviet central Asia). — Campbell and Davies, 1991: 87 (checklist of species; Alaska and Canada). — Silfverberg, 1992: 21 (list of species of Fennoscandia, Denmark, and the Baltic States). — Zanetti, 1993 (key to species of Caucasus, Turkey, Iran, and Lebanon; new species; synonyms; catalog). — Siitonen, 1993: 230 (list of Finnish species). — Welch, 1993 (ovariole structure and development). — Boháč, 1993: 40 (list of species of Czech Republic and Slovakia). — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species). — Zanetti, 1995: 5 (list of species of Italy). — M. Hansen, 1996: 94 (list of Danish species). — M. Hansen, Mahler, Palm, and Pedersen, 1996: 240 (additions to list of Danish species). — Downie and Arnett, 1996: 438 (characters; key to species of Northeastern North America ). — M. Hansen, Palm, Pedersen, and Runge, 1998: 70 (list of species collected in 1997; Denmark). — Zanetti, 1998: 154 (key to species from Taiwan). — Assing, Frisch, Kahlen, et al., 1998: 121 (supplement to key for species in Lohse, 1964). — Newton, Thayer, Ashe, and Chandler, 2000: 338 (27 Nearctic species; notes; characters in key). — Naomi, Kuranishi, Saito, and Maruyama, 2000: 102 (collected from Kamchatka).

Abinothum Tottenham, 1939d: 225 (proposed for “ Anthobium pars sensu Kraatz”; subgenus of Eusphalerum ). TYPE SPECIES: Anthobium longipenne Erichson , by original designation and monotypy. — Blackwelder, 1952: 32 (type species: longipenne ). — Zanetti, 1982: 98 (synonym of Eusphalerum ). — Zanetti, 1987: 56 (synonym of Eusphalerum ). — Boháč, 1993: 40 (subgenus of Eusphalerum ).

Onibathum Tottenham, 1939d: 225 (proposed for “ Anthobium pars sensu Kraatz”; subgenus of Eusphalerum ). TYPE SPECIES: Silpha minutum Fabricius , by original designation. — Tottenham, 1949: 354, 405 (type species: minutum Fabricius ; list of British species; subgenus of Eusphalerum ). — Blackwelder, 1952: 275 (type species: minutum ). — Hatch, 1957: 82 (subgenus of Eusphalerum ; characters; key to species of Pacific Northwest). — Zanetti, 1982: 98 (synonym of Eusphalerum ). — Zanetti, 1987: 56 (synonym of Eusphalerum ).

Pareusphalerum Coiffait, 1959c: 216 , 248 (subgenus of Eusphalerum ; species included: viertli , atrum , florale). TYPE SPECIES: Omalium atrum Heer , fixed by original designation. — Zanetti, 1982: 99 (subgenus of Eusphalerum ). — Zanetti, 1987: 50, 152 (subgenus of Eusphalerum ; characters; key to species of Italy). — Zanetti, 1993: 260 ( Eusphalerum ; subgenus of Eusphalerum ; characters). — Boháč, 1993: 40 (subgenus of Eusphalerum ). — Assing, Frisch, Kahlen, et al., 1998: 122 (subgenus of Eusphalerum ; characters).

abdominale Gravenhorst, 1806 , see: semicoleoptratum Panzer, 1795 .

adultum Kiesenwetter, 1851 , see: torquatum Marsham, 1802 .

adustum Heer, 1839 , see: limbatum Erichson, 1840 .

adustum Kiesenwetter, 1851 , see: torquatum Marsham, 1802 .

aetolicum Kraatz, 1858b: 129 ( Anthobium ; Type locality: Griechenland: Ätolische Alpen). — Bernhauer and Schubert, 1910: 38 ( Anthobium ; catalog; Dalmatia; Greece; Hercegovina; Austria). — Scheerpeltz, 1933: 1032 ( Anthobium ; catalog; Caucasus). — Tikhomirova, 1973a: 137 ( Eusphalerum ; subgenus Abinotum ; Caucasus). - DISTRIBUTION: Greece, Bosnia Hercegovina, Caucasus.

afghanicum Herman, 2001: 36 ( Eusphalerum ; replacement name for nuristanicum Coiffait ). - DISTRIBUTION: Afghanistan.

nuristanicum Coiffait, 1982b: 77 ( Eusphalerum ; subgenus Eusphalerum ; [preoccupied]; Type locality: Afghanistan: Nouristan, Kamdesch, 2000 m).

— Herman, 2001: 36 ( Eusphalerum ; synonym of afghanicum ).

africanum Roubal, 1915 , see: luteicorne Erichson, 1840 .

akikoae Watanabe, 1999: 263 ( Eusphalerum ; subgenus Eusphalerum ; Type locality: China: Guangxi Province: Xing’an Xian, Mt. Miao’er Shan, near Tieshan Ping).

- DISTRIBUTION: China.

albertae Zanetti, 1981: 207 ( Eusphalerum ; subgenus Eusphalerum ; Type locality: Svizzera: Laquintal, dint. Simplon-Village, Valais, 1000 m).

— Zanetti, 1987: 132 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Lohse and Lucht, 1989: 125 ( Eusphalerum ; characters).

— Zanetti, 1995: 5 ( Eusphalerum ; Italy).

- DISTRIBUTION: Italy, Switzerland.

albipile Fauvel, 1900d: 219 ( Anthobium ; Type locality: Tyrol).

— Bernhauer and Schubert, 1910: 38 ( Anthobium ; catalog).

— Zanetti, 1982: 105 ( Eusphalerum ; characters; Italy).

— Zanetti, 1987: 116 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Zanetti, 1995: 5 ( Eusphalerum ; Italy).

- DISTRIBUTION: Austria, Italy.

alpestre Heer, 1839 , see: robustum Heer, 1839 .

alpestre Motschulsky, 1857 , see: alpinum Heer, 1839 .

alpinum Heer, 1839: 180 ( Omalium ; [preoccupied]; Type locality: Glarneralpen; Berneralpen; Moloja; Valserberg; Genferalpen).

— Heer, 1841: 569 ( Omalium ; characters; Switzerland).

— Fairmaire and Laboulbène, 1856: 649 ( Anthobium ; characters; France).

— Kraatz, 1857d: 1012 ( Anthobium ; synonym of alpinum ).

— Fauvel, 1869: 494 ( Anthobium ; synonym of montanum ).

— Fauvel, 1871a: 74 [= 1872: 48] ( Eusphalerum ; characters; France; Germany; Switzerland; Italy; Austria).

— Mulsant and Rey, 1880: 349 ( Anthobium ; characters; France).

— Ganglbauer, 1895: 755 ( Anthobium ; characters; central Europe).

— Reitter, 1909: 197 ( Anthobium ; characters; Germany).

— Bernhauer and Schubert, 1910: 38 ( Anthobium ; catalog).

— Portevin, 1929: 454 ( Anthobium ; characters; France).

— Scheerpeltz, 1933: 1032 ( Anthobium ; catalog).

— Freude, 1957: 106 ( Anthobium ; Germany).

— Smetana, 1958: 57 ( Anthobium ; notes).

— Coiffait, 1959c: 220 ( Eusphalerum ; characters; France).

— Horion, 1963: 33 ( Anthobium ; Pyrenees; Balkans; Germany; Austria).

— Lohse, 1964: 31 ( Eusphalerum ; subgenus Abinothum ; characters; central Europe).

— Scheerpeltz, 1968a: 5 ( Anthobium ; catalog; Austria).

— Tikhomirova, 1973a: 137 ( Eusphalerum ; checklist; USSR).

— Zerche, 1977a: 8 ( Eusphalerum ; Germany).

— Burakowski, Mroczkowski, and Stefańska, 1979: 16 ( Eusphalerum ; catalog; Poland).

— Blum, 1979: 242 ( Eusphalerum ; elytral reduction; abdominal mobility; morphology).

— Klinger, 1980: 455 ( Eusphalerum ; subgenus Abinothum ; characters of abdominal defensive gland).

— Herger and Uhlig, 1981: 81 ( Eusphalerum ; habitat notes; Switzerland).

— Zanetti, 1982: 108 ( Eusphalerum ; characters; central Europe; Romania).

— Tóth, 1982: 39 ( Eusphalerum ; characters).

— Klinger, 1983: 38 ( Eusphalerum ; natural history notes; Germany).

— Zanetti, 1984: 75 ( Eusphalerum ; Bulgaria).

— Terlutter, 1984: 10, 36 ( Eusphalerum ; notes; Germany). — Zanetti, 1987: 95 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Lucht, 1987: 85 ( Eusphalerum ; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— K. Koch, 1989: 217 ( Eusphalerum ; habitat).

— Boháč, 1993: 40 ( Eusphalerum ; subgenus Abinothium ; subspecies alpinum ; Czech Republic). — Zanetti, 1995: 5 ( Eusphalerum ; Italy).

— Paśnik, 1998: 69 ( Eusphalerum ; habitat; Poland).

— Herman, 2001: 36 ( Eusphalerum ; nomen protectum).

- DISTRIBUTION: Europe, Algeria.

montanum Erichson, 1840: 897 ( Anthobium ; [Note: This name is older than alpinum ]; Type locality: alpi Austriae Helvetiaeque ).

— Heer, 1841: 569 ( Omalium ; synonym of alpinum ).

— P. Lucas, 1846: 128 ( Anthobium ; notes; Algeria).

— Märkel and Kiesenwetter, 1848a: 328 ( Anthophagus ; notes; Carpathian Alps).

— Redtenbacher, 1849: 755 ( Anthobium ; characters; Austria).

— Redtenbacher, 1857: 256 ( Anthobium ; characters; Austria).

— Kraatz, 1857d: 1012 ( Anthobium ; characters; Germany).

— Fauvel, 1865a: 319 [= 1865b: 69] ( Anthobium ; habitat; France).

— Redtenbacher, 1874: 280 ( Anthobium ; characters; Austria).

— Mulsant and Rey, 1880: 353 ( Anthobium ; characters; France).

— Ganglbauer, 1895: 755 ( Anthobium ; synonym of alpinum ).

— Bernhauer and Schubert, 1910: 38 ( Anthobium ; synonym of alpinum ).

— Scheerpeltz, 1933: 1032 ( Anthobium ; synonym of alpinum ).

— Zanetti, 1987: 95 ( Eusphalerum ; synonym of alpinum ).

luteipenne Erichson, 1840: 898 ( Anthobium ; Type locality: Saxonia montana).

— Redtenbacher, 1849: 829 ( Anthobium ; characters).

— Kiesenwetter, 1851a: 439 ( Anthobium ; notes; Silesia).

— Fairmaire and Laboulbène, 1856: 652 ( Anthobium ; characters; France).

— Redtenbacher, 1857: 256 ( Anthobium ; characters; Germany).

— Kraatz, 1857d: 1013 ( Anthobium ; characters; Germany).

— Fauvel, 1871a: 74 [= 1872: 48] ( Eusphalerum ; synonym of alpinum ).

— Redtenbacher, 1874: 281 ( Anthobium ; characters).

— Seidlitz, 1875: 238 ( Anthobium ; characters; Baltic region).

— Mulsant and Rey, 1880: 350 ( Anthobium ; synonym of alpinum ).

— Ganglbauer, 1895: 755 ( Anthobium ; synonym of alpinum ).

— Bernhauer and Schubert, 1910: 38 ( Anthobium ; synonym of alpinum ).

— Scheerpeltz, 1933: 1032 ( Anthobium ; synonym of alpinum ).

— Zanetti, 1987: 95 ( Eusphalerum ; synonym of alpinum ).

alpestre Motschulsky, 1857 a: 491 ( Anthobium ; [preoccupied]; Type locality: Alpes de la Carniole).

— Fauvel, 1865c: 18 [= 1866: 15] ( Anthobium ; synonym of montanum ).

— Fauvel, 1871a: 74 [= 1872: 48] ( Eusphalerum ; synonym of alpinum ).

— Ganglbauer, 1895: 755 ( Anthobium ; synonym of alpinum ).

— Bernhauer and Schubert, 1910: 38 ( Anthobium ; synonym of alpinum ).

subjectum Mulsant and Rey, 1880: 352 ( Anthobium ; Type locality: montagnes du Var,

au nord de Fréjus).

— Ganglbauer, 1895: 755 ( Anthobium ; synonym of alpinum ).

— Bernhauer and Schubert, 1910: 38 ( Anthobium ; synonym of alpinum ).

— Zanetti, 1987: 95 ( Eusphalerum ; synonym of alpinum ).

obenbergeri Zanetti, 1982: 109 ( Eusphalerum ; subspecies of alpinum ; Type locality: Slovacchia centr., Monti Nízké Tatry).

— Zanetti, 1987: 95 ( Eusphalerum ; subgenus Eusphalerum ; subspecies of alpinum ; characters; Italy).