Reichardtia pedator (Sharp, 1876)

Lackner, Tomas & Leschen, Richard A. B., 2017, A monograph of the Australopacific Saprininae (Coleoptera, Histeridae), ZooKeys 689, pp. 1-263 : 55-58

Reichardtia pedator (Sharp, 1876) Figs 240, 241-252, 253-255, 256-262, 758

Saprinus pedator Sharp, 1876: 25.

Type locality.

New Zealand: Auckland.

Type material examined.

Saprinus pedator Sharp, 1876: Lectotype, present designation: 1 spec., with " Saprinus / pedator / N. Zeal. Type / D.S." written on the actual mounting card with the specimen, followed by: "Auckland / New Zealand" (printed); followed by: "Sharp Coll. / 1905-313"; followed by: “Type” (red-margined printed round label); followed by: " Saprinus pedator / Sharp, 1876 / LECTOTYPE 2014 / Des. Lackner & Leschen" (red label, written) (BMNH). Paralectotypes, present designation: 1 spec., " Saprinus pedator / D.S. / Otago Hulton 1878" (written on the actual mounting card), followed by: "Sharp Coll. / 1905-313"; followed by: " Saprinus pedator / Sharp, 1876 / PARALECTOTYPE 2014 / Des. Lackner & Leschen" (red label, written) (BMNH). 1 spec., " Saprinus pedator / Sharp / Ind. Typ. / N. Zeald D.S." (written on the actual mounting card); followed by: “Auckland” (red label, written); followed by: "Auckland / New Zealand" (printed); followed by: " Saprinus pedator / Sharp, 1876 / PARALECTOTYPE 2014 / Des. Lackner & Leschen" (red label, written) (BMNH). 1 spec., " Saprinus pedator / Sharp. / Otago Hulton 1878" (written on the actual mounting card); followed by: "Sharp Coll. / 1905-313"; followed by: " Saprinus pedator / Sharp, 1876 / PARALECTOTYPE 2014 / Des. Lackner & Leschen" (red label, written) (BMNH). 1 spec., with the following labels: " Saprinus / pedator / N. Zeald. Ind. Typ. / D.S." (written); followed by: "G. Lewis Coll. / B.M. 1926-369" (printed); followed by: " Saprinus pedator / Sharp, 1876 / PARALECTOTYPE 2014 / Des. Lackner & Leschen" (red label, written) (BMNH). Sharp (1876) does not mention the exact number of specimens in his original description and the lectotype designation serves to fix the identity of the species.

Additional material examined.

NEW ZEALAND. North Island: ND: 1 spec., Waipapakauri, 7.xi.1991, G. Kuschel leg. (NZAC); 1 ♀, Cape Maria, 20.i.1963, L. Price leg. (NZAC). AK: 1 spec., Ruakaka Beach, N. Auckland, 22.ix.[19]32, C.E. Clarke coll. (BMNH); 1 spec., Muriwri Beach, 12.ix.[19]32, C.E. Clarke coll. (BMNH). TK: 1 spec., Taranaki, New Zealand, Broun Coll. (BMNH). WI: 1 spec., Himatangi Beach, 20.iii.1988, M.O.E. Peters leg. (AMNZ). WN: 1 spec., New Zealand, Parapararaumu, 17.ii.[19]24, O’Connor, G.V. Hudson, in dead fish on beach (BMNH); 1 spec., Wellington, ii.1902, J.J. Walker (BMNH); 1 spec., Manawatu Estuary, 25.iii.1984 (CJN); 1 spec., Waikawa Beach, 2.iv.1995 (CJN); 1 spec., Waikanae Beach, 9.ii.1995 (CJN). South Island. NN: 3 specs., Farewell Spit, Outer beach, 9.ii.1981, J.W. Early leg. (LUNZ); 7 specs., Wharariki Beach, 7.ii.1981, J.W. Early leg. (LUNZ, 3 exs. in coll. TLAN); 1 ♂, Tahunanui, Beach Nelson 21.viii.1976, J.C. Watt leg. (NZAC). SC: 1 spec., Temuka Beach, 22.iii.2008 (CJN). SL: 4 specs., South Island, Curio Bay, 50 km E of Invercargill, sandy beach, 4.xii.2000, Erber (ZMHUB).

DN: 2 specs., Kuri Bush, near Taieri Mouth, 3.xi.2001 (CJN); 2 specs., Sandfly Bay, Otago Peninsula, 18.i.2004 (CJN); 1 spec., Smails beach, Dunedin, 16.iii.1999 (CJN); 1 spec., Boulder Beach, Otago Peninsula, 5.i.2005 (CJN); ditto, but 29.xii.2001 (CJN); 1 spec., Waikouati, 14.i.2002 (CJN). Unknown localities: 1 spec., N. Zealand, no further data, G. Lewis Coll. (BMNH); 1 spec., New Zealand, no further data, Broun Coll. (BMNH); 1 spec; New Zealand, 25.ii.1917, Broun Coll. (BMNH).

Biology.

Littoral species found on beach under kelp, dead fish, often buried in the sand.

Distribution.

Endemic to New Zealand (Fig. 758).

Re-description.

Body length: PEL: 3.25-4.25 mm; APW: 1.25-1.60 mm; PPW: 2.35-3.00 mm; EL: 2.15-2.90 mm; EW: 2.60-3.35 mm.

Body (Fig. 240) rectangular oval, elytral humeri prominent, strongly convex, cuticle light to dark brown, shining, without metallic luster, entirely glabrous; legs, antennae and maxillary palpi similarly colored.

Antennal scape (Fig. 241) slightly thickened, with numerous long setae; antennal club (Fig. 242) comparatively small, rounded, entirely covered in dense short sensilla, intermingled with sparse longer erect sensilla; sensory structures of antennal club (Fig. 255) in form of a single ventral sensory area with pear-shaped vesicle situated beneath it.

Mandibles (Fig. 253) massively thickened with rounded outer margin, strongly carinate dorsally, bluntly pointed, sub-apical tooth on inner margin of left mandible large, blunt; labrum (Fig. 254) faintly convex dorsally, sparsely microscopically punctate, anterior margin slightly emarginate; labral fold tiny, not conspicuous; setae of lateral fringe conspicuously short; labrum with two labral setae; terminal labial palpomere elongated, its width about one-third its length; palpal organ present on both labial and maxillary palpi; mentum (Fig. 243) sub-trapezoid, anterior angles slightly produced, anterior margin with shallow median excavation and tiny median notch (Fig. 243), surface around it with four long setae, lateral margins with several microscopic setae, surface of mentum imbricate; cardo of maxilla with few short setae on lateral margin; stipes triangular, with four long setae; lacinia without lacinial hook (=uncus); terminal maxillary palpomere elongated, its width about one-fourth its length, about three times as long as penultimate palpomere.

Clypeus (Fig. 241) large, rectangular, strongly convex, sloping down anteriorly, rounded laterally, glabrous, surface between clypeus and frontal disc strongly concave; supraorbital and frontal striae absent; frontal disc (Fig. 241) about twice as broad as long, with conspicuous median fovea; eyes slightly flattened, but visible from above.

Pronotal sides (Fig. 240) rounded, apical angles blunt, marginal pronotal stria complete, carinate, somewhat weakened behind head; disc strongly convex, glabrous; pronotal hypomeron with long yellow setae; scutellum small, but visible.

Elytral epipleura punctate; marginal epipleural stria thin, faintly impressed; marginal elytral stria well impressed, continuous along elytral apex as apical elytral stria, connected with sutural elytral stria; humeral elytral stria well impressed, occasionally continuous with inner subhumeral stria forming a stria parallel to first dorsal elytral; dorsal elytral striae deeply impressed, first the longest, reaching approximately half of elytral length apically, striae 2-4 shorter, reaching about one-third of elytral length apically; sutural elytral stria complete, deeply impressed, continuous with apical as well as 4th dorsal elytral striae. Except for scattered punctures near elytral apex entire elytral disc glabrous.

Propygidium (Fig. 244) transverse, about three times as broad as long, completely exposed, sparsely punctate, punctures separated by several times their diameter, interspaces between punctures with fine alutaceous microsculpture; pygidium (Fig. 244) almost smooth, only on basal third with several scattered shallow almost inconspicuous punctures.

Anterior margin of median portion of prosternum (Fig. 245) bisinuate, setose, anterior angles somewhat produced; marginal prosternal stria absent; prosternal apophysis strongly constricted between procoxae, prosternal process knife-like, strongly widening and sloping down anteriorly, surface imbricate, with scattered punctures fringed with setae.

Anterior margin of mesoventrite (Fig. 246) straight; discal marginal mesoventral stria well impressed, fringed with several short setae; disc convex, only with microscopic punctation, strongly constricted between mesocoxae; meso-metaventral suture deeply impressed, meso-metaventral sutural stria well impressed, overriding meso-metaventral suture; intercoxal disc of metaventrite broad, smooth, in males with concave median longitudinal depression. Lateral metaventral stria well impressed, strongly curved outwardly; lateral disc of metaventrite (Fig. 247) convex, with round shallow large setigerous punctures; metepisternum + fused metepimeron (Fig. 247) with even denser setigerous punctures than those of lateral disc of metaventrite, punctures almost disappear on fused metepimeron; lateral metepisternal stria intermittent, present only on metepisternum.

Intercoxal disc of first abdominal ventrite in males with large depression medially, in females depression much smaller, lateral striae shortened apically; glabrous.

Protibia (Fig. 248) widening apically, outer margin with a row of about 17 densely-set straight long lamellate denticles growing in size apically; setae of outer row moderately dense, long and regular; setae of median row approximate to outer row, but shorter and sparser; protarsal groove deep; anterior protibial stria complete, carinate, next to it another row of much shorter sparse setae present, apically setae growing in length; protibial spur short, hooked, growing out near tarsal insertion; outer part of posterior surface of protibia (Fig. 249) with dense moderately long strongly sclerotized setae, demarcation line between outer, median and inner part of posterior surface of protibia absent; inner margin with double row of sparse sclerotized setae growing in size and becoming denser apically.

Mesotibia (Fig. 250) strongly thickened, outer margin with a dense row of thick denticles growing in size apically; setae of outer row strongly sclerotized, dense, growing in size but becoming thinner apically; setae of median row rather distanced from outer row, thinner and sparser; between the two rows several additional setae present; posterior mesotibial stria inconspicuous (absent?); anterior surface of mesotibia convex, with several dense rows of short denticles, becoming sparser and thinner towards inner mesotibial margin; anterior mesotibial stria inconspicuous (absent?); mesotibial spur inconspicuous; first and second tarsomeres ventrally with four long strongly sclerotized setae; third and fourth tarsomeres with only two such setae; fifth tarsomere devoid of setae ventrally; first two tarsomeres dorsally with two strongly sclerotized setae; third, fourth and fifth tarsomeres dorsally with only single seta; claws of apical tarsomere bent, shorter than half its length; metatibia (Figs 251-252) basically similar to mesotibia, but even more thickened, denticles of outer margin and anterior surface much denser and stouter than those of mesotibia.

Male genitalia. Eighth sternite (Figs 256-257) fused longitudinally; vela absent; apex of eighth sternite laterally with several short setae; eighth tergite and eighth sternite not fused laterally (Fig. 258). Ninth tergite (Figs 259-260) longitudinally fused medially; spiculum gastrale (Fig. 259) gradually dilated on apical half, basal end strongly dilated, resembling flukes of a whale. Aedeagus (Figs 261-262) slightly thickened, constricted before apex; basal piece of aedeagus very short, ratio of its length: length of parameres 1: 7; parameres of aedeagus fused almost along their basal two-thirds; aedeagus only slightly curved from lateral view, apex of aedeagus flattened dorso-ventrally.