Eupines (Byraxis) hectori ( Broun, 1895 )

Shen, Jia-Wei & Leschen, Richard A. B., 2020, Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae), Zootaxa 4777 (1), pp. 1-84 : 36-37

Eupines (Byraxis) hectori ( Broun, 1895)

Fig. 22 ; map Fig. 51

Bryaxis hectori Broun, 1895: 73 .

Eupines (Byraxis) hectori : combination and subgeneric assignment by Raffray 1904: 208. Raffray 1911: 83, 1924: 157; Nomura and Leschen 2006: 255; Maddison 2010: 417; Shen and Leschen 2019: 264 .

Eupines (Byraxis) diversides Newton, 2017: 13 . New name for Bryaxis diversa Broun, 1893b: 174 , preoccupied by Bryaxis diversa Raffray, 1887: 36 . Bryaxis diversa Broun, 1893b: 174 . Eupines (Byraxis) diversa : combination and subgeneric assignment by Raffray 1904: 207. Raffray 1911: 83, 1924: 157; Hudson 1923: 364, 1934: 183; Nomura and Leschen 2006: 254; Maddison 2010: 417. Shen and Leschen 2019: 264 . Type locality: [AK] Hunua Range, Drury. Type depository: BMNH LT ♂, 5 PLT; NZAC 2 PLT. NEW SYNONYMY.

Type locality: [BP] Tarukenga, near Rotorua.

Type depository: BMNH LT ♂ .

Diagnosis: Head with vertexal foveae distinct and asetose; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7 and A8 unmodified, A9 cylindrical, about 3 × longer than A8, with medium-sized spine surrounded in small anterolateral excavation, A10 conical and slightly longer than A9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of slender median struts, about three-fourths of ventral length. Protrochanters with medium-sized ventral spine, about three-fifths of protrochanteral width; protibiae and mesotibiae unmodified.

Redescription: Body length 1.45–1.55 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, with about 25 facets; vertexal foveae distinct and asetose; antennal tubercles barely visible; postantennal notches weak; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7 and A8 unmodified, A9 cylindrical, about 3 × longer than A8, with medium-sized spine surrounded in small anterolateral excavation, A10 conical and slightly longer than A9. Pronotum slightly wider than long, slightly wider than head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest at apical three-fourths; humeral calli barely visible; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.3 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of slender median struts, about three-fourths of ventral length. Legs with protrochanters modified with medium-sized ventral spine, about three-fifths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.28 mm long, with parameres slightly curved inwards, widest at basal two-thirds, bearing four pairs of long apical setae and pair of short apical setae; median lobe complex, with three long sclerites, left one and middle one about as wide as parameres in dorsal view, strongly curved towards each other, right one straight and long, with pair of slightly sclerotized laminae next to three sclerites at dorsal side, with another pair of strongly sclerotized basal laminae strongly curved at ventral side, about one-third length of parameres.

Comments: The lectotype of E. (B.) hectori was compared with the lectotype and two paralectotypes of E. (B.) diversides (as E. (B.) diversa ), and were found to represent a single species by having identical modifications of A9 ( Figs. 22 B–D), with the latter name treated as the junior synonym. Newton (2017) proposed a new name, E. (B.) diversides for the junior primary homonym, Bryaxis diversa which is now unavailable, and is also here treated as the junior synonym of E. (B.) hectori .

Distribution: North Island. AK, WO, CL, BP, WN.

Additional material examined: 14 ♂♂, CL: 2, Rakitu I., 167 m, 07/Jan/1981, L. Roberts, litter, 81/7; 2, Coromandel, Twin Kauri Tk., 03/Nov/2005, R. Leschen & S. Nomura, flood debris, 37°00'S 175°38'E; 2, JNPC, WN, Levin L Papaitonga, 17/XI/2015, forest floor litter, J. Nunn; WO: 1, Collector, 14/3/1920, A.E. Brookes, Okauia Kaimai, Matamata, A.E. Brookes Collection; BP: 1, Kaimai, May/1920, A.E. Brookes Collection; 6, Kaimai Range, Leyland Tramline, 427 m, 22/Sep/1981, B.M. May, litter, 81/92. Associated females (14 ♀♀): AK: 1, Ex. Broun Collection, Hunua Ra., Maketu, Auckland, 2735, Byraxis diversa Broun, T. Broun Collection, A. E. Brookes Collection; CL: 4, Rakitu I., 167 m, 07/Jan/1981, L. Roberts, litter, 81/7; 1, 2739, Broun; BP: 8, Top Kaimai, 16/12/35, A.E. Brookes Collection.

Figures

FIGURE 22. Diagnostic characters of E. (B.) hectori (Broun, 1895). A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, B–F = 0.2 mm, G–J = 0.1 mm.

FIGURE 51. Geographic distributions of E. (B.) glabrata (Broun, 1886), E. (B.) graceae sp. n., E. (B.) halli (Broun, 1921), E. (B.) hectori (Broun, 1895), E. (B.) hoarei sp. n., E. (B.) huizhenae sp. n., and E. (B.) illustris (Broun, 1914) in New Zealand.