Gracilipsodes koghiensis, Malm & Johanson, 2008

Malm, Tobias & Johanson, Kjell A., 2008, Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini), Zoological Journal of the Linnean Society 153 (3), pp. 425-452 : 435-438

GRACILIPSODES KOGHIENSIS SP. NOV. ( FIGS 6 , 13 )

Diagnosis: Gracilipsodes koghiensis is distinguished from G. lanceolatus sp. nov., G. grandis sp. nov., G. psocopterus , and G. similis by the tibial spur formula of 2, 2, 2, and by the presence of a pair of pre-apical lateral processes of tergum X, originating from the apical half of the segment. It is separated from G. aureus sp. nov. and G. robustus sp. nov. by hind tibia with a higher number of spines, in tergum X, in the genitalia, by the longer lateral processes, and in the forewing, by longer crossvein m–cu oriented at a sharp angle to M. It is separated from G. aurorus sp. nov. by having a paler body colour, by the higher number of hind tibial spines, by the tergum X being apically slit rather than notched, and by the apical part of the inferior appendage having a higher number of mesal spines. It is distinguishable from G. aoupiniensis sp. nov. by having longer superior appendages and, in tergum X, by the deeper apical slit.

Description, male: Body brown. Forewing medium brown, with a few pale spots. Tibial spur formula 2, 2, 2. Hind tibiae each with 20–25 small black spines (N = 9). Forewing ( Fig. 6A ): length 7.0– 7.7 mm (N = 9), apically rounded; forks 1 and 5 present; crossvein m–cu longer than crossvein r–m, forms sharp angle with M. Hindwing ( Fig. 6B ): length 4.8–5.5 mm (N = 9), apex narrowly rounded; forks 1, 3, and 5 present; crossvein r–m straight; costa with 23 hamuli.

Genitalia: Segment IX annular, laterally setose, narrowest at ventral part; tergite IX broader than sternite IX in lateral view ( Fig. 6C ), anterior margins of segment IX weakly produced anteriorly immediately below mid height ( Fig. 6C ). Superior appendages setose, flattened dorsoventrally, longer than tergum X, and with apices nearly acute in dorsal view ( Fig. 6D ). Tergum X membraneous; median process entire, basally wide, apex rounded, dorsal margin concave in lateral view ( Fig. 6C ), apex with deep slit in dorsal and ventral views ( Fig. 6D, E ); pre-apical lateral processes as long as or longer than median process, produced at apical half, sclerotized, digitate, directed posterad, and slightly curved ventrally ( Fig. 6C ). Inferior appendages each bipartite, setose; basal part broad in ventral view ( Fig. 6E ), mesally concave, produced posteromesally into a broad, apically rounded lip, bearing small dorsal spines ( Fig. 6C, E ); apical part of each inferior appendage digitate, curving mesally; in lateral view, wide, parallel-sided, apex rounded, and bearing 50–60 mesal spines ( Fig. 6C ); mesal process with six strong spines ( Fig. 6D ). Harpago: weakly sclerotized, slender, apex nearly acute ( Fig. 6C ). Phallic apparatus simple ( Fig. 6F, G ); phallicata basally tubular, fused with phallobase, apex truncate; strongly bent ventrad at apical half; apical part with pair of sclerotized, slender, lateral processes, being directed ventrally and tapering distally ( Fig. 6F ); phallotremal sclerite U-shaped ( Fig. 6G ).

Holotype (male): New Caledonia, Province Sud, Mt Koghi , 462 m a.s.l., 2–5 December 2001, Malaise trap, loc#7 (Johanson, Pape & Viklund).

Paratypes: One male, same data as holotype ; 11 males, ditto, except 15–16November 2001 , net, loc#7; one male, ditto, except 6 December 2001 , light trap; one male, ditto, except c. 300 m south of Koghi Restaurant , 22.18288°S, 166.50490°E, 457 m a.s.l., 2–16 Novemeber 2003, Malaise trap, loc#001 (Johanson) ; two males: ditto, except 16–26 November 2003 ; one male: ditto, except 22.18288°S, 166.50393°E, 447 m a.s.l., 2–16 November 2003 , Malaise trap, loc#002; one male, ditto, except 16–26 November 2003 ; two males, ditto, except 22.18288°S, 166.50245°E, 427 m a.s.l., 2–16 November 2003 , Malaise trap, loc#003; three males, ditto, except 16–26 November 2003 ; four males, ditto, except 22.18288°S, 166.50167°E, 417 m a.s.l., 16–26 November 2003 , Malaise trap, loc#004; one male, ditto, except c. 800 m south of Koghi Restaurant , 22.18311°S, 166.50564°E, 460 m a.s.l., 10–26November 2003 , Malaise trap, loc#019; one male, ditto, except 22.18365°S, 166.50451°E, 440 m a.s.l., 11–26 November 2003 , Malaise trap, loc#021.

Distribution: Province Sud, Mt Koghi.

Etymology: The name koghiensis is derived from the type locality, Mt Koghi.

Figures

Figure 6. Gracilipsodes koghiensis sp. nov. A, forewing. B, hindwing. C, genitalia, lateral. D, genitalia, dorsal. E, genitalia, ventral. F, phallus, lateral. G, phallus, ventral.

Figure 13. Maps of New Caledonian Grande Terre with plotted distributions of the different species. A, Gracilipsodes aoupiniensis sp. nov. (triangles), Gracilipsodes aureus sp. nov. (white circles), Gracilipsodes grandis sp. nov. (squares), Gracilipsodes koghiensis sp. nov. (diamond), Gracilipsodes lanceolatus sp. nov. (black circle). B, Gracilipsodes aurorus sp. nov. (triangle), Gracilipsodes robustus sp. nov. (black diamond), Gracilipsodes psocopterus Sykora, 1967 (circles), Gracilipsodes similis Ward (2001) (squares).