Caulomorphus kociani, Hlaváč, 2020

Hlaváč, Peter, 2020, Review of the tribe Typoderini (Coleoptera: Curculionidae) from Caucasus with the description of a new species of Caulomorphus from Armenia, Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 60 (2), pp. 629-638 : 635-636

Caulomorphus kociani sp. nov.

( Figs 1c , 6a–c , 7a–e )

Type locality. Armenia, Goris, 39.516611N, 46.322847E, 1700 m a.s.l. Type material. Hඈඅඈඍඒඉൾ: J, ‘ ARMENIA, Goris, E slope, deciduous forest, leaf litter sifting, 1700 m, 25.V.2015, 39.516611N 46.322847E, M. Kocian lgt.[p]’ ( NMPC). Pൺඋൺඍඒඉൾඌ:(21 JJ 13 ♀♀): 2 JJ, the same data as holotype ( CPHP); 3 JJ 1 ♀, ‘ ARMENIA, Tatev env., Vorotan Valley, sifting in macchia, 1300 m, 24.V.2015, 39.394394N 46.247672E, M. Kocian lgt. [p]’ ( CPHP); 1 J 2♀♀, ‘ ARMENIA, Syunik r., 5 km N of Shurnukh, 07.06.2015, F.Pavel lgt.[p] / 39°24′15.7″N, 046°24′37.8″E [p]’ ( CJKH); 1 J, ‘ ARMENIA – Syunik pr., Davit Bek env., N of Karmrakar, 7.6.2017, lgt. Jan Pelikán / 39°19′00″N, 46°28′16″E, 1474 m, kořeny [roots] Corylus . [p]’ ( CJPH); 8 JJ, 3 spec., ‘ Armenia, 25 km S Kapan, Gomarants Ps., 39°01′32″N, 46°21′59″E, 2190 m, oak forest, 7.VII.2016, V.Assing [p]’ ( BMNH, CPHP, OUMNH, ZMHB, CLFC); 1 J, ‘ Armenia [33]-WSW, Kapan, above Shishkert, 39°03′48″N, 46°21′36″E, 2040 m, forest litter sift., 12.vii.2018, V. Assing [p]’ ( ZMHB); 2 JJ 3 ♀♀, ‘ Armenia [27]-WSW, Kapan, W Karajan, 39°09′22″N, 46°06′13″E, 2050 m, mixed forest, 10.VII.2018, V.Assing [p]’ ( ZMHB); 1 J 2♀♀, ‘ Armenia [38]- 20 km SW Sisian Darbas, 39°25′55″N, 46°06′57″E, 1680 m, forest margin, 16.VII.2018, V. Assing [p]’ ( ZMHB); 2 JJ 2 ♀♀, ‘ Armenia [28]- ca 30 km W Kapan, 39°15′13″N, 46°04′16″E, 2040 m, forest margin, 11.VII.2018, V. Assing [p]’ ( ZMHB, PCLF).

Description. Body ( Fig. 1c ) convex. Length 3.10–4.10 mm, maximum width of elytra 1.00– 1.20 mm. Head and pronotum shiny, whole body, antennae and legs unicoloured reddish-brown, elytra with dense setae. Rostrum curved downwards, with five longitudinal carinae, with short apical setae, scrobe deep, straight, not visible in dorsal view. Antennae ( Fig. 7a ) long, with dense pubes- cence, scape thin and parallel-sided at base and expanded towards apex, about 1.10× as long as funicule, funicule with 7 antennomeres; antennomeres II and III elongate and expanded towards apex, antennomere II 1.40–1.60× as long as III, antennomeres IV–VI subequal, about as long as wide, shortest, about 0.6× as long as antennomere III, antennomere VII 1.25× as long as VI and subequal to VIII, antennal club about twice as long as antennomere II and about 1.75× as long as wide. Pronotum 0.90–1.05× as long as wide, 1.00–1.05× as long as head, widest in middle, basal margin straight, with closely arranged large and subequal punctures, with thin but well-defined median carina. Prosternum rugosely sculptured, with uneven large punctures, hypomerae fused with median part of prosternum; procoxae touching, isthmus interrupted. Mesoventrite evenly, densely punctate, in middle about as long as metaventrite. Metaventrite with even and large puctures, punctures much larger than those on mesoventrite. Mesocoxae well-separated, width of isthmus about ⅓ of mesocoxa diameter, posterior meso- and anterior metaventral processes truncate. Metacoxae widely separated by large, posterior metaventral process with well-defined median notch. Ventrite I about 1.3× longer than II, both punctate, with golden setation ( Fig. 7b ). Elytra long, subparallel-sid- ed, on sides slightly rounded in apical third, 1.40–1.75× as long as wide, 2.15–2.30× as long as pronotum, with eight striae composed of large punctures of equal diameter, distance between each two punctures within row slightly less than diameter of puncture, elytral interstriae elevated, with even micropunctures bearing short, golden setae. Male terminalia. Aedeagus ( Figs 6a–b ) wide, flattened dorsally, before apex narrowed, laterally slightly curved, with acuminate apex, with two subequal sclerites, male sternite IX ( Fig. 6c ) curved, asymmetrical, hemisternite rounded at apex, base slightly concave, lacking apical setae. Female terminalia. Spermatheca ( Fig. 7c ), female sternite VIII ( Fig. 7c ), gonocoxite ( Fig. 7d ).

Differential diagnosis. Caulomorphus kociani sp. nov. and the most similar species C. besucheti are readily separated from its congeners by sharing the subparallel-sided elytra, symmetrical, wide, flattened and laterally curved aedeagus and pronotum lacking median carina, however the new species differs from C. besucheti in different shape and different internal structures of the aedeagus (see Figs 5a–b ).

Etymology. Patronymic, named after Matúš Kocian (Prague, Czech Republic), specialist on Staphylinidae and the collector of part of the type series.

Distribution. Armenia.

Figures

Fig. 1. Dorsal habitus. a – Adexius scrobipennis Gyllenhal, 1834; b – Anchonidium caucasicum Motschulsky, 1845; c – Caulomorphus kociani sp. nov.; d – Anchonidium perpensum Faust, 1886; e – Aparopion costatum (Fåhraeus, 1843); f – Pseudaparopion kadleci Borovec, Osella & Zuppa, 2002.

Fig. 6. Caulomorphus kociani sp. nov. a – aedeagus, dorsal view; b – aedeagus, lateral view; c – sternite IX and hemisternites.Scale bars:0.2 mm.

Fig. 7. Caulomorphus kociani sp. nov., a – antenna; b – abdominal ventrites I–V; c – spermatheca; d – sternite VIII; e – gonocoxite. Scale bars: 0.2 mm.

Fig. 5. Caulomorphus besucheti Osella, 1970: a – aedeagus, dorsal view; b – aedeagus, lateral view; c – sternite IX and hemisternites; Caulomorphus talyschensis Reitter, 1897: d – aedeagus, dorsal view; e – aedeagus, lateral view; f – sternite IX and hemisternite. Scale bars: 0.2 mm.