Bethylomimus academus Trjapitzin, 1967

Japoshvili, George, Higashiura, Yoshimitsu & Kamitani, Satoshi, 2016, A review of Japanese Encyrtidae (Hymenoptera), with descriptions of new species, new records and comments on the types described by Japanese authors, Acta Entomologica Musei Nationalis Pragae 56 (1), pp. 345-401 : 363

Bethylomimus academus Trjapitzin, 1967 comb. restit.

Bethylomimus academus Trjapitzin, 1967: 208 . Cerchysiella togashii Tachikawa, 1988: 63–65 , syn. nov.

Material examined. Cerchysiella togashii : PARATYPES: 5 ♀♀ (2 ♀♀ on slides), Kaga city, Ishikawa pref. ex Mesosa longipennis , 6.vii.1987, I. Togashi lgt. ( ELKU).

Distribution. Japan: Honshu ( TACHIKAWA 1963a). Russia ( NOYES 2016).

Comments. According to John Noyes (pers. comm.), Bethylomimus was incorrectly synonymised with Zaommoencyrtus Girault, 1916 , by TRJAPITZIN & GORDTH (1978). It differs from Zaommoencyrtus in having the head hypognathous, the filum spinosum conspicuously converging with the proximal margin of the linea calva and directed towards the junction of the submarginal and marginal veins and lack a strongly swollen parastigma whereas in Zaommoencyrtus the head is more or less prognathous, the filum spinosum is not directed towards the junction of the submarginal and marginal veins and the parastigma is strongly swollen and much wider than the submarginal vein. The characters highlighted by YANG et al. (2013) are incorrect as they are based on an undescribed species, whose placement in Zaommoencyrtus by NOYES et al. (1996) is possibly incorrect. It is likely that species previously placed in Bethylomimus could be quite easily accommodated in that genus. However, as we are confident that C. togashii is a synonym of B. academus (supported by morphological characters and proximity of the type locality of B. academus , Russia: Primorskiy Kray, to Japan) we prefer to reinstate Bethylomimus as a valid genus until the generic limits of the two genera can be investigated in more detail. Further work will probably suggest that Bethylomimus includes diminutive species that actually represent a monophyletic clade derived from within the Cerchysiella clade.