Bembidium (Notholopha), Jeannel, 1962

Maddison, David R., 2014, An unexpected clade of South American ground beetles (Coleoptera, Carabidae, Bembidion), ZooKeys 416, pp. 113-155 : 131

Taxon classification Animalia Coleoptera Carabidae

Subgenus Notholopha Jeannel, 1962

Notholopha Jeannel, 1962; type species Bembidium punctigerum Solier, by original designation.

Pacmophena Jeannel, 1962; type species Bembidium scitulum Erichson, by original designation.

Remarks.

Notholopha consists of 11 described species ( Toledano 2002; Toledano 2008), and several undescribed (two of which are sequenced here). These are small beetles with large, protruding eyes (Fig. 2B, C), with small flagella in the internal sac of the male genitalia, and with brush sclerites. They have the general appearance when running of a member of the Holarctic subgenus Bembidion . Some frequent habitats similar to those of subgenus Bembidion , including dry habitats far from water ( Bembidion stricticolle ), or upper banks of creeks (e.g., Bembidion sp. “Nahuelbuta”). Others occur at high elevation near small rivulets in open, alpine areas (e.g., Bembidion rugosellum and Bembidion melanopodum ), or at the edges of snowfields ( Bembidion sexfoveolatum ).

Jeannel considered Pacmophena and Notholopha to be two subgenera within the genus Notholopha . As the characters that Jeannel used to distinguish the two are minor characters such as surface texture and antennal length, and as it appears that Pacmophena is paraphyletic with respect to the Notholopha (s. str.) (Fig. 7), I consider them synonymous, with Notholopha as the valid name.