Nocaracris sabulosus Ramme, 1951

Ünal, Mustafa, 2016, Pamphagidae (Orthoptera: Acridoidea) from the Palaearctic Region: taxonomy, classification, keys to genera and a review of the tribe Nocarodeini I. Bolívar, Zootaxa 4206 (1), pp. 1-223 : 176-177

Nocaracris sabulosus Ramme, 1951 stat. rev.

( Figs. 601 , 705–706 , 846 )

Nocaracris sp.: Uvarov 1930: 373.

Nocaracris sabulosa n. sp.: Ramme 1951: 279, 304, 427.

Nocaracris sabulosa Ramme, 1951 : Karabağ 1958: 121; Weidner 1969: 160; Otte 1994: 172. Nocaracris bodenheimeri n. f. flavipes: Ramme 1951: 301.

Paranocaracris rubripes bodenheimeri forma flavipes Ramme, 1951: Presa & Garcia 1983: 23. Nocaracris bodenheimeri flavipes Ramme 1951 : Otte 1994: 172.

Paranocaracris citripes sabulosa ( Ramme, 1951) : Demirsoy 1973: 423; Demirsoy 1977: 79; Presa & Garcia 1983: 23. Paranocaracris rubripes burri f. acarinata f. n.: Demirsoy 1973: 426, partim; Demirsoy 1977: 83, partim. Paranocaracris citripes citripes Uvarov, 1949 : Önder et al. 1999: 165.

Type locality. Turkey, Çorum. Holotype: female ( NHMUK).

Material examined. TURKEY: Çorum, 12.7.1930, 1♀ (Holotype) (leg. M. Sureya) ( NHMUK) ; Amasia, 1860, 1♂, 2♀ (leg. Mann) (det. Ramme as Nocaracris bodenheimeri n.f. flavipes) ( MfN) ; Amasia, 1860, 2♂, 1♀ (leg. Mann), 1♀ (leg. Lederer) (det. Ramme as Nocaracris bodenheimeri f. flavipes) ( NMW) ; Amasya, Duruca , 400 m, 2.5.1992, 6♂, 2♀ (leg. M. Ünal) ; Amasya, Akdağ , 1620 m, 5.7.2006, 2♀ (leg. M. Ünal) ; Amasya, Akdağ , 1225–1600 m, 1.7.2014, 1♀ in alcohol, ibidem , 1660 m, 2♀ in alcohol (leg. M. Ünal); Amasya, Ferhat Kayası , 954 m, April 2008, 1 ♀ (leg. İ. Eker) (AİBÜEM) ; Turkey, [Tokat], Niksar , 900 ft, 29.7.1959, 1♂ (leg. K. M. Guichard) ( NHMUK) ; Asia Minor, Amasia, 1♂ (det. as rubripes ) ( MNCN) ; Gamsun [Samsun], 1♂ (leg. Steindachner) ( NMW) .

Distribution. Central part of N. Turkey: Çorum, Amasya, Tokat, Samsun provinces ( Fig. 846 ).

Description of male: Body ( Fig. 705 ) slender, compressed laterally. Fastigium of vertex ( Fig. 601 ) short and wide, slightly depressed and sloping. Frontal ridge slightly narrowed just below the median ocellus. Vertex ( Fig. 601 ) narrow, 1.3 times narrower than vertical diameter and very slightly narrower than transversal diameter of eye; supraocular foveola distinct, open type, semiclosed in some males; median carinula long and grooved; convex in lateral view. Antennae with 12 segments. Pronotum ( Figs. 601 , 705 ) compressed laterally, roof-shaped; lateral carinae weak, more distinct in metazona of some males; median carina slightly raised, with distinct longitudinal sulcus, wider in anterior part; slightly convex or straight in lateral view. Anterior margin of prosternum raised collar-like with a blunt median projection, in some males tridentate or tongue-shaped but never sharply pointed. Mesosternal interspace 2.2 times wider than its length and clearly narrower than mesosternal lobes. Hind femur ( Fig. 705 ) 2.6 times longer than its height, proximal part widened and sharply narrowing towards genicular lobes, dorsal margin straight in proximal part, then sloping backwards, finelly serrated; ventral margin convex. Hind tibia with 9 inner, 9–11 outer spines. Abdominal tergites ( Fig. 705 ) simple; median carina very slightly raised, not protruded posteriorly. Mesonotum and metanotum with grooved median carinae. Tympanum absent. Arch of zygoma narrow; posterior lobes of zygoma relatively short and narrow; apodemes narrow with distinct apical notch; pseudolophi separated with 10–13 spines.

Coloration. Body brown with many small cream and black spots. Ventral half of head, fore and mid legs and antenna light brown; dorsal surface of head blackened except the cream basal part of median carinula of vertex. Pronotum with small cream and black tubercles; typical light band on paranota creamish-brown. Typical light band on abdomen distinct; lateral surface of abdomen dark brown. Outer surfaces of hind femur and tibia light creamishbrown with small dark spots; inner and ventral surfaces of hind femur blackened, dorsal margin of inner surface body color. Inner surface and inner half of dorsal surface of hind tibia orange, turning to reddish-orange apically with black tipped spines, outer half of dorsal surface yellow with outer spines, in one male tibia yellow without orange. Tarsus reddish-orange. Sternites light brownish-cream or yellowish-cream; anterior margins of abdominal sternites and subgenital plate with a distinct brown band transversally.

Measurements (mm) of both sexes. body length: male 21–23.8, female 36–37.8; pronotum length: male 4.6– 4.9, female 7.1–8.2; pronotum height: male 4.8–5.3, female 8.5–9.6; pronotum width anterior: male 4.3–4.5, female 6.7–7.2; pronotum width posterior: male 5.9–6.4, female 9.6–10.6; hind femur length: male 8.6–9, female 12.4–14.5; hind femur height: male 3.3–3.7, female 4.7–5.2.

Remarks. Ramme (1951: 301) described a new form N. bodenheimeri f. flavipes which is undoubtedly conspecific with N. sabulosus by the orange or yellow hind tibia, the compressed body laterally, the roof-shaped pronotum without or with weak lateral carinae, some of which are the reasons of its placement in the N. tridentatus sp. group while N. bodenheimeri Uvarov is placed in another species group (see the key).

Demirsoy (1973, 1977) cited this species as “( Ramme 1951: 304)” with its type locality “ Çorum ” for two taxa, Paranocaracris citripes sabulosa Ramme ( Demirsoy 1973: 423; 1977: 79) and Paranocaracris rubripes burri f. acarinata ( Demirsoy 1973: 426; 1977: 83). While the Çorum population belongs to N. sabulosus , the specimens from Ilgaz Dağı belong to Nocararis niethammeri Ramme, 1951 .

The previously unknown male is described here.

According to the ICZN (Articles 31.2, 34.2) the species name of this taxon is combined.

Figures

FIGURE 846. Distribution of the species of genera Nocaracris and Turkanocaracris.

FIGURES 584 – 610. Body parts. 584 – 585 female hind femur, 584 Nocaracris karadagi, paratype; 585 Nocaracris monticolus, paratype. 586 – 587 Nocaracris karadagi, female paratype, 586 longitudinal sulcus of pronotum; 587 pronotal carinae. 588 – 589 Nocaracris palandoken, female paratype, 588 longitudinal sulcus of pronotum; 589 pronotal carinae. 590 – 591 male hind femur, 590 Nocaracris latipes, paratype; 591 Nocaracris monticolus, holotype. 592 – 593 Nocaracris elegans, female, 592 vertex and eye; 593 prosternum. 594 – 595 Nocaracris tridentatus, female, 594 vertex and eye; 595 prosternum. 596 – 597 Nocaracris tecticollis, female holotype, 596 body high; 597 head frontal view; 598 Nocaracris citripes, paratype, body low. 599 – 600 female vertex, 599 Nocaracris citripes; 600 Nocaracris karshitoros, paratype. 601 – 602 male pronotal carinae, 601 Nocaracris sabulosus; 602 Nocaracris karshitoros. 603 Nocaracris dilekensis, female tympanum. 604 – 606 Nocaracris goektepe, 604 without tympanum, female; 605 male prosternum; 606 female prosternum. 607 – 609 Nocaracris bicoloripes, 607 without tympanum, female; 608 male prosternum; 609 female prosternum. 610 Nocaracris subrubratus, female prosternum.

FIGURES 695 – 708. Habitus. 695 – 696 Nocaracris van, 695 male holotype dorsal; 696 female paratype dorsal. 697 – 698 Nocaracris citripes, 697 male holotype; 698 female paratype. 699 – 700 Nocaracris bicoloripes, 699 male; 700 female holotype. 701 – 702 Nocaracris dilekensis, 701 female holotype; 702 ditto dorsal. 703 – 704 Nocaracris tecticollis, 703 male; 704 female holotype. 705 – 706 Nocaracris sabulosus, 705 male; 706 female. 707 – 708 Nocaracris karshitoros, 707 male holotype, mirror view; 708 female paratype.