Tmethis pulchripennis (Serville, 1838)

Ünal, Mustafa, 2016, Pamphagidae (Orthoptera: Acridoidea) from the Palaearctic Region: taxonomy, classification, keys to genera and a review of the tribe Nocarodeini I. Bolívar, Zootaxa 4206 (1), pp. 1-223 : 44-45

Tmethis pulchripennis (Serville, 1838)

( Figs. 17 , 103 , 121–122 , 214–215 )

Tmethis pulchripennis asiaticus Uvarov 1943: 66 .

Tmethis pulchripennis asiaticus Uvarov, 1943 : Karabağ 1958: 114; Weidner 1969: 156; Demirsoy 1977: 36; Otte 1994: 162; Sevgili & Çıplak 2000: 234.

Tmethis pulchripennis asiaticus Uv. : Ramme 1951: 410, 427. Tmethis sp.: Karabağ et al. 1980: 14.

Tmethis pulchripennis (Serville, 1838) : Massa 2013: 440 , 441.

Material examined. As Tmethis pulchripennis pulchripennis : LIBYA: Libyen , Driana, 30 km nö Benghasi, 4.1942, 2♂ (leg. Kirchberg) ; Cyrenaica , 1♂ (leg. Ruhmer) ( MfN).

As Tmethis pulchripennis asiaticus Uvarov, 1943 : PALESTINE: Khedeira , costal zone, 30.5.1921, 1♂ (Holotype) (leg. P. A. Buxton) ( NHMUK) ; TURKEY: Urfa and environs, 7.1931, 1♂, 1♀ (Paratypes) ( NHMUK) ; Mardin, 5.5.1966, 1♂ (leg. A. Demirtola) ; Adıyaman, Besni, Akkuyu Mezrası , 23.6.1972, 1♀ (AİBÜEM); LEBANON : Libanon, Baalbek , 30.5.1961, 1♂ (leg. Kasy & Vartian) ( NMW) ; ISRAEL: Jerusalem, Linnaea V., 1♂, 1♀ (AİBÜEM) ; Israel, Arad, 4.6.1963, 1♀ (leg. Pener & Blondheim); Lahalo , 28.4.1963, 2♂ (leg. Pener et al.) (det. Hollis); Haluza, 15.6.1982, 1♂ (leg. et det. Fishelson) (NHMUK); Golan Höhen, Umg. El Quneitra, Österr. Un-Baon, Camp Faouar , 33.09.45 N, 35.37.30 E, 1981, 1♀ ( NMW) ; Golan Höhen , Umg. El Quneitra, 33.14 N, 35.47 E, 1978, 1♂ ( NMW) ; IRAQ: Assur , Mesopot., 1910, 2♀ (leg. Pietschmann); Mosul , 25.5.1910, 1♀; Hinler, Djeolalabe , 11.6.1910, 1♀ (leg. Pietschmann) ( NMW) ; SYRIA: Ladakia , 1♂ ; Syria, 20 km N.O. v. Damaskus, 16– 23.5.1961, 1♀ (leg. Kasy & Vartian) ( NMW) ; PALESTINE: Wadi Ghuzze , 25.11.1930 – 30.4.1931, 1♂ (leg. N. Scott) ( NHMUK); JORDAN : Jordan Valley, Khor Fasayil , 10.6.1951, 7♂, 5♀ ; Jordan, Ghor Damiya, Basat Thmeila , 2.4.1951, 2♂, 2♀ ; 14.6.1951, 1♀; Ghor Damiya, Ain Khanaizir , 2.6.1951, 2♀; Jordan, Wadi Sbu’eib , 10.4.1951, 1♂ (all leg. A. R. Waterston) ; Mafraq, 1956, 1♂ (W. H. Reay) (det. Ragge as T. pulchripennis ); IRAN: Naft-i-ghah, Somar , 6.5.1973, 2♂ (leg. A. A. Soltani) (all in NHMUK).

Remarks. Massa (2013: 441) synonymized Tmethis pulchripennis asiaticus Uvarov, 1943 with the nominotypical subspecies because of the high variability of T. pulchripennis . It is a fact that most of the Asian specimens have yellow tibia as in T. pulchripennis asiaticus . But in some populations it is possible to see the specimens with yellow and reddish orange tibia even in the same habitat. e.g. While the male has yellow hind tibia, the female with orange red hind tibia both of which collected from Jaffa, Israel (NMW) or one of the three females from Syria has red hind tibia (MfN) or two females collected from Israel with reddish-orange hind tibia in the NHMUK and one female from Palestine has also slightly reddish orange hind tibia (NHMUK). However the Libyan specimens examined here are recognizable as the nominotypical subspecies by the smaller size and the distinct red hind tibia (different shades of red from the Asian specimens with reddish hind tibia) from T. p. asiaticus . Therefore I prefer to give the materials separately for further research.

Figures

FIGURES 17 – 31. Male phallic complex. a phallus lateral, b phallus dorsal, c epiphallus, d phallus posterior view. 17 Tmethis pulchripennis (asiaticus); 18 Atrichtmethis semenovi; 19 Melanotmethis fuscipennis fuscipennis; 20 Pezotmethis tartarus tartarus; 21 Glyphanus obtusus; 22 Asiotmethis limbatus; 23 Glyphotmethis escherichi escherichi; 24 Utubius syriacus; 25 Strumiger desertorum desertorum; 26 Thrinchus campanulatus campanulatus; 27 Filchnerella beicki; 28 Mongolotmethis gobiensis gobiensis; 29 Beybienkia lithophila; 30 Eotmethis nasutus; 31 Haplotropis brunneriana. Scales 1 mm (short scale for a, b, c; long scale for d).

FIGURES 93 – 105. Body parts. 93 – 94 mid tibia in male, 93 Glyphotmethis dimorphus dimorphus; 94 Pamphagus caprai; 95 male hind wing of Prionotropis maculinervis. 96 – 97 female head antero-lateral view 96 Pezotmethis tartarus tartarus; 97 Paranocarodes turkmen. 98 – 99 male vertex, 98 Haplotropis brunneriana; 99 Pezotmethis tartarus tartarus. 100 – 101 female pronotum, 100 Thrinchus turcmenus; 101 Haplotropis brunneriana. 102 – 103 apical part of left hind tibia in female, 102 Thrinchus turcmenus; 103 Tmethis pulchripennis (asiaticus). 104 – 105 female prosternum, 104 Glyphotmethis escherichi inermis; 105 Rhinotmethis beybienkoi. Abbreviations: a: preocellar foveola, b: supraocellar foveola, c: supraocular foveola.

FIGURES 106 – 128. Body parts. 106 – 108 male pronotum, 106 Eremopeza gibbera gibbera; 107 Eremocharis bampura bampura; 108 Asiotmethis limbatus. 109 – 110 male arolium, 109 Prionotropis hystrix; 110 Tuarega insignis. 111 – 114 subtympanal lobe in female, 111 Prionotropis maculinervis; 112 Prionotropis hystrix; 113 Iranotmethis cyanipennis iranicus; 114 Eremocharis subsulcata beccalonii. 115 – 116 female hind femur, 115 Iranotmethis cyanipennis iranicus; 116 Eremotmethis carinatus. 117 – 118 female vertex, 117 Iranotmethis persa zagrosi; 118 Eremopeza reducta. 119 – 120 male pronotum, 119 Eremocharis granulosa brachycera; 120 Tuarega insignis. 121 – 124 male vertex, 121 and 122 Tmethis pulchripennis (asiaticus); 123 and 124 Atrichotmethis semenovi. 125 – 126 female pronotum, 125 Atrichotmethis semenovi; 126 Melanotmethis fuscipennis fuscipennis. 127 – 128 metazona of pronotum in female, 127 Glyphanus obtusus; 128 Pezotmethis tartarus tartarus.

FIGURES 205 – 220. Habitus. 205 – 206 Eremocharis ancestralis, 205 male holotype; 206 female paratype. 207 – 208 Eremocharis bampura bampura, 207 male paratype; 208 female paratype. 209 – 210 Eremocharis bampura khorasana, 209 male paratype; 210 female paratype. 211 – 212 Dhofaria splendens, 211 male paratype; 212 female paratype. 213 Tuarega insignis, male. 214 – 215 Tmethis pulchripennis (asiaticus), 214 male; 215 female. 216 Tmethis cisti (gracilis). 217 – 218 Atrichotmethis semenovi, 217 male; 218 female. 219 – 220 Melanotmethis fuscipennis fuscipennis, 219 male; 220 female.