Bryanites Valentine, 1987

Liebherr, James K., 2017, Bryanites graeffii sp. n. (Coleoptera, Carabidae): museum rediscovery of a relict species from Samoa, Zoosystematics and Evolution 93 (1), pp. 1-11 : 3

Bryanites Valentine, 1987

Type species.

Bryanites samoaensis Valentine (by original designation).

Diagnosis.

These beetles can be diagnosed from other Pacific platynine taxa by the much reduced subapical sinuation of the elytra, with the elytral margins straight to convex near the apex (Fig. 1 ). The pronotum is broadly orbiculate, with the lateral margins convex to only slightly concave anterad the rounded hind angles. The pronotum has the basal seta present and situated anterad the rounded hind angle. The elytra are moderately narrowed basally, with the basal elytral groove meeting the lateral elytral depression at an angle; i.e. the humeri are angulate. The male aedeagus exhibits characters typical of tribe Platynini (Fig. 4 ): 1, both parameres are rounded apically, or conchoid in shape; 2, the right paramere is smaller than the left, and the aedeagus lies with the right side ventral in repose; 3, the basal bulb is closed and bears a sagittal crest; and 4, the internal sac is tubular and unarmored, with slightly more melanized spicules present apically near the gonopore.

Beetles of all three Bryanites species are brachypterous, with the wing rudiments narrow and elongate stenopterous straps, the length of the rudiments more than 4 × the rudiment breadth. Beetle size is moderate to large, ranging from 11.7 mm in Bryanites barri (measurement of Barr 1987) to 16.2 mm in Bryanites graeffii .

Key to the Adults of Bryanites Valentine

1.	Beetles of moderate size, standardized body length 11.7-12.5 mm; pronotum quadrisetose both lateral and basal setae present; parascutellar seta present	2
-	Larger beetles, standardized body length 16.2 mm; pronotum bisetose, only basal seta present both sides; parascutellar seta absent	Bryanites graeffii sp. n.
2.	Third elytral interval with 6 setae along length, the anterior seta associated with the third stria, the trailing five setae associated with the second stria; elytral basal groove meeting lateral elytral depression at right angle; pronotum without any evident laterobasal depressions, at most a slight depression near hind margin (left side of holotype)	Bryanites samoaensis Valentine
-	Third elytral interval with 3 setae along length, the anterior seta associated with the third stria, the posterior two setae associated with the second stria; elytral basal groove meeting lateral elytral depression at acute angle; pronotum with broad, shallow linear laterobasal depressions extended from hind angles toward center of pronotal disc	Bryanites barri Valentine

Figures

Figure 1. Male holotype, Bryanites graeffii, dorsal view.

Figure 4. Male aedeagus, internal sac everted, of Bryanites graeffii male holotype, right lateral view. Abbreviations include: ae, aedeagal median lobe; is, internal sac; lp, left paramere; rp, right paramere.