Tetragonoderus flavovittatus Waterhouse, 1881

Felix, Ron F. F. L., Farkač, Jan & Suleiman, Ahmed Saeed, 2012, Annotated checklist of the Carabidae (Coleoptera) of the Socotra Archipelago, Insect biodiversity of the Socotra Archipelago. (Acta Entomologica Musei Nationalis Pragae 52), pp. 75-106 : 87

Tetragonoderus flavovittatus Waterhouse, 1881

Material examined (86 spec.). YEMEN: SOCOTRA ISLAND: Homhil plateau, 12°34′N, 54°19′E, 540 m a.s.l., 9.ii.1999, 7 spec., lgt. H. Pohl, det. W. Lorenz ( HLMD) ; Kilisan , 12°29.136′N, 54°19.715′E, 12.ii.1999, 2 spec., lgt. K. van Damme, det. M. Persohn ( HLMD) ; S Socotra, 6.–24.ix.1999, 6 spec., lgt. V. Bejček & K. Šťastný, det. J. Farkač ( CULS, DWCB) ; Ayhaft , 12.ii.2000, 3 spec., lgt. V. Bejček & K. Šťastný, det. J. Farkač ( CULS) ; Noged , 12.318N, 53.678E, 250 m a.s.l., 27.ii.–1.iii.2000, 1 spec., lgt. V. Bejček & K. Šťastný, det. J. Farkač ( JFCP) ; Shuab , 10.iii.2000, 1 spec., lgt. V. Bejček & K. Šťastný, det. J. Farkač ( CULS) ; Noged , Farmihin, 12°24′41″N, 54°13′35″E, 0 m a.s.l., 24.–25.x.2000, 1 spec., lgt. H. Pohl, det. M. Persohn ( HLMD) ; Noged , Farmihin, Steroh, wadi, 12°24′26″N, 54°08′40″E, 24.x.2000, 5 spec., lgt. A. van Harten, det. M. Persohn ( HLMD) ; Noged , 12.318N, 53.678E, 250 m a.s.l., 12.–13.xi.2000, 3 spec., lgt. V. Bejček & K. Šťastný, det. J. Farkač ( CULS) ; Qaariah vill. env., 12°38′05″N, 54°12′39″E, 11 m a.s.l., 28.xi.2003, 1 spec., lgt. P. Kabátek, det. J. Farkač ( JFCP) ; Hadiboh , 10–100 m a.s.l., 21.xi.–12.xii.2003, 3 spec., lgt. P. Kabátek, det. J. Farkač ( JFCP, CULS) ; the same data, 1 spec., lgt. J. Farkač ( JFCP) ; Dixam plateau, Firmihin area, 12°47′40″N, 54°01′53″E, 428 m a.s.l., 3.xii.2003, 6 spec., lgt. & det. J. Farkač ( JFCP, CULS) ; the same data, 3 spec., lgt. P. Kabátek ; Wadi Esgego , Dixam plateau, 12°28′09″N, 54°00′36″E, 300 m a.s.l., 2.–3.xii.2003, 2 spec., lgt. & det. J. Farkač ( JFCP, CULS) ; Homhil protected area, 12°34′27″N, 54°18′32″E, 364 m a.s.l., 28.–29.xi.2003, 3 spec., lgt. & det. J. Farkač ( JFCP) ; Qaareh (waterfall), Noged plain, 12°20′10″N, 53°37′56″E, 57 m a.s.l., 5.–6.xii.2003, 1 spec., lgt. & det. J. Farkač ( JFCP) ; Wadi Ayhaft , 12°36′38″N, 53°48′49′E, 190 m a.s.l., 24.–26.xi.2003, 3 spec., lgt. & det. J. Farkač ( JFCP) ; east of Hadiboh , near Suq, approximately 12°26′N, 54°01′E, 28.x. 2000, 1 spec., lgt. & det. H. Pohl ( HLMD) ; Hadiboh , village and environment, 12°36′57″N, 54°01′01″E, 20.x.–1.xi.2000, 11 spec., lgt. H. Pohl, det. M. Persohn ( HLMD) ; Wadi Di-Negehen , 12°38′33.0″N, 54°03′18.5″E, 20.ii.2009, 1 spec., lgt. & det. R.F.F.L. Felix ( RFBE) ; Lagoon Qaryah , 12°32′22.7″N, 54°17′38.5″E, 24.ii.2009, 1 spec., lgt. & det. R.F.F.L. Felix ( RFBE) ; Hadiboh plaine, 12°39′14.97″N, 54°03′02.68″E, 23.ii.2009, 4 spec., lgt. & det. R.F.F.L. Felix ( RFBE) ; Deyup cave environment, 16.vi.2009, 1 spec., lgt. V. Hula, det. R.F.F.L. Felix ( NMPC) ; Wadi Difarroha , south side, 15.i.2010, 1 spec., lgt. Saldaitis, det. R.F.F.L. Felix ( IRSB) ; Wadi Ayeft , 20.i.2010, 1 spec., lgt. Saldaitis, det. R.F.F.L. Felix ( IRSB) ; Wadi Ayaft , 12°36′21.7″N, 53°59′33.9″E, 265 m a.s.l., 26.x.2010, 6 spec., lgt. & det. R.F.F.L. Felix ( RFBE) ; Mômi plateau, 12°32′40.7″N, 54°21′27.0″E, 505 m a.s.l., 2.xi.2010, 4 spec., lgt. & det. R.F.F.L. Felix ( RFBE) ; Wadi Fa’rhoh , Nogid, 250 m a.s.l., 12°26.464′N, 54°09.327′E, 15/ 16.ii.2001, 4 spec., lgt. V. Neumann, det. D.W. Wrase ( DWCB) .

Diagnosis. Body length 7.3–8.3 mm. Dorsal surface black, smooth and shiny. Elytra with yellow markings consisting of two divided patches on each elytron. Antennae and legs pale with darker femora.

Comments on classification. Tetragonoderus flavovittatus is similar to T. quadrum (Fabricius, 1792) from Yemen mainland which has a narrower pronotum. In T.flavovittatus the aedeagus has four large internal spines while in T. quadrum there are six much smaller spines.

Collection circumstances. The specimens from Wadi Fa’rhoh, Nogid, collected by V. Neumann were found in dry shrub land. In 2009 and 2010 the species was found in similar environments on plains (near Hadiboh, Wadi Ayhaft, and other places), in the day underneath stones and at night running on the soil.

Distribution. A species endemic to Socotra Island.

Harpalini