Tylodinus kuscheli Luna-Cozar, 2014

Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. & León-Cortés, Jorge L., 2014, A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico, Zootaxa 3788 (1), pp. 1-63 : 49-50

29. Tylodinus kuscheli Luna-Cozar , sp. nov.

( Figures 82–83 , 122 , 134 )

Diagnosis. Length male, 3.9–4.2 mm, female 4.1–4.2 mm. Width male, 2.1–2.2 mm, female 2.3 mm. Body 2.1x longer than wide, black. Head with punctures similar to punctures on rostrum, clothed with yellow scales; vertex with overlapping scales, uniformly distributed along vertex, similar in color and tonality to head scales; frons concave; rostrum moderately robust, moderately carinate, apical area puncticulate; eyes moderately convex. Pronotum in lateral view strongly convex, outline with anterior constriction, without posterior constriction ( Figure 10 , 83 ); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc with longitudinal impression; surface with dense, deep punctures on flanks, becoming smaller, scattered towards middle, with yellow to light brown scales very dense on lateral area and flanks, anterior area forming a thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark scales on middle and posterior areas of disc ( Figure 14 , 82 ); surface without granules. Elytra black, occasionally black with dark red color along intervals 1–2, basal margin slightly sinuate ( Figure 3 ), with tubercles as follows: I2 with tubercle slightly beyond declivity base; I3 with three tubercles, first subbasal, second on middle of elytra, similar to subbasal tubercle, third on middle of elytral declivity, similar to tubercle on I2; I4 with two tubercles, first very weakly developed between first two tubercles on I3, second on base of elytral declivity, longer than tubercle on I2; I5 with two tubercles at same position as first two tubercles on I3; I6 with two weak tubercles in same position as those on I4; I7 with two small tubercles in same position as tubercles on I5; elytral declivity with tubercle on I4 much longer than tubercle on I2. Metasternum glabrous, moderately concave, metasternal tooth absent. Male abdomen with ventrites 3–5 not in same plane as ventrites 1–2, ventrite 1 moderately concave, squamous, ventrite 2 as long as 3–4 combined, with a transverse line of scales along middle, ventrite 5 longer than ventrite 1, flat, with anterior impression. Femora armed, hind femur reaching elytral apex, premucro absent. Male genitalia ( Figure 122 ) with median lobe in lateral view weakly curved, moderately stout; in ventral view longer than half the length of apodemes, swollen at 0.25 length from base of median lobe, convergent, apex acute; apical process in lateral view large (0.35–0.42 as long as median lobe), slightly curved; in ventral view not constricted beyond median orifice, lateral sides convergent. Female not dissected.

Comments. This species is closely related to T. pappi and T. branstetteri based on the body being clothed mainly with yellow to fulvous scales, moderately dense, dark brown scales that are less sparse, and the median lobe in ventral view distinctly wider at the apical 1/3 (at median orifice).

Geographic distribution. México (Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in various wet forest types between 1300–1600 m.

Derivation of specific name. Patronym, named after Guillermo (Willy) Kuschel ( New Zealand), a wellknown worker on weevils.

Material examined. Total 4 males, 2 females. Holotype male ( CMNC): México: Chiapas, Villa Corso, Ejido Sierra Morena , R . Biosfera La Sepultura , elev. 1487 m (16°9.773’N, 93°3536.211’W), 13.v.2008, oak forest litter, coll. R . S. Anderson. Paratypes: México: Chiapas, Villa Flores, Ejido Sierra Morena , R . Biosfera La Sepultura , elev. 1400 m (16°9'10.4''N, 93°35'25.1''W), 19–20.vii.2003, coll. R . S. Anderson — 1 male ( ECOS); same locality, elev. 1600 m (16°9'10.6''N, 93°35'25.1''W)— 1 female ( CMNC); same locality, elev. 1360 m (16.16028°N, 93.60507°N), 12.v.2008, LLAMA (Wa-A-01-1-40) — 1 female ( CMNC); same locality (16.15969°N, 93.60516°N), 12.v.2008, LLAMA (Wa-A-01-1-23) — 1 male ( ECOS); same locality, elev. 1310 m (16.15404°N, 93.60080°N), 12.v.2008, LLAMA (Wa-A-01-2-17) — 1 male ( CMNC) .

Figures

FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli.

FIGURES 120–129. Ventral and lateral habitus of median lobe: 120. Tylodinus dominicus, 121. Tylodinus noctis, 122. Tylodinus kuscheli, 123. Tylodinus pappi, 124. Tylodinus branstetteri, 125. Tylodinus immundus, 126. Tylodinus intzin, 127. Tylodinus gibbosus, 128. Tylodinus kissingeri, 129. Tylodinus pinguis.

FIGURES 134–135. Maps showing collecting sites. 134. Occurrences in Chiapas, México of: Tylodinus rufus, Tylodinus kuscheli, Tylodinus pappi, Tylodinus jonesi, Tylodinus coapillensis; 135. Occurrences in Chiapas, México of: Tylodinus spiniventris, Tylodinus lum, Tylodinus noctis, Tylodinus pinguis, Tylodinus rugosus.

FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic).

FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth.