Drilus rectus Schaufuss, 1867

Kobieluszova, Lucie & Kundrata, Robin, 2015, Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini, Zootaxa 4012 (1) : -

Drilus rectus Schaufuss, 1867

( Figs 7 , 27 , 39 )

Drilus rectus Schaufuss, 1867: 84 .

Type material. Not available.

Type locality. Beirut ( Lebanon) and Antiochia (southern Turkey).

Material studied. Male, " Turkey, 975 m, Toros Dağları, 9.6.1987, Lundberg lgt." ( LMBC); male, "South Turkey, prov. Hatay, Harbyie env., 400m, 36°07'N 36°10'E, 31.3.2010, E. & P. Hajdaj lgt." ( PCRK); 2 males, " Turkey, 13.–14.VI.2003, Adana vil. (ca 45 km N), Boztahta env., 37°23' N, 35°15' E, 460 m, Jiří Hájek & Josef Hotový leg."( NMPC); male, " Turkey, 5. 1990, Iskenderun, leg. Krajčík, coll. general, National Museum Prague, Czech Republic " ( NMPC); male, " TR mer., province Hatay, Karaçay, Hatay env., valley of Asi Nehri river, 7.5.2005, J. Vávra leg." ( PCJV).

Diagnosis. Drilus rectus is similarly colored as D. akbesianus , D. nemethi and D. iranicus ( Figs 1, 7, 12 ). It differs from D. akbesianus and D. nemethi in relatively smaller eyes and from D. iranicus in the lightly colored abdomen and shallower transverse groove near the frontal prosternal margin.

Remarks. Drilus rectus occurs in Cyprus, southern Turkey and mainland Levant, which makes this species relatively widespread in comparison to the vast majority of its congeners. Similarly to other widely distributed Drilus species ( D. concolor Ahrens, 1812 and D. flavescens ; personal observation), the different populations and in some cases even the specimens within a single population vary in the body size, coloration, and several morphological characters including e.g. the head surface, shapes of frontal depression, pronotum and paramera ( Schaufuss 1867, Kundrata et al. 2014b). The detailed DNA-based study together with knowledge of the biology and ecology is important for understanding the taxonomic status of the geographically distant D. rectus populations. This species was recently redescribed and figured by Kundrata et al. (2014b) so we are not repeating the description here. Female and immature stages are unknown. A female specimen from Cyprus collected by H. M. Morris and determined as D. rectus (Tab. 1) is deposited in the Natural History Museum in London, but without any further details this determination must be confirmed by future studies.

Distribution. Cyprus, Israel, Lebanon, Turkey (Adana, Hatay; Fig. 55 ). Given the current distribution of this species, its occurence in Syria is highly probable.

Figures

FIGURE 55. The distribution of Drilus species in Asia Minor.

FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm.

FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm.

FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm.