Colemanus Fisher, 2015

Fisher, J. Ray, Tucker, Erika M. & Sharkey, Michael J., 2015, Colemanus keeleyorum (Braconidae, Ichneutinae s. l.): a new genus and species of Eocene wasp from the Green River Formation of western North America, Journal of Hymenoptera Research 44, pp. 57-67 : 61

Colemanus Fisher gen. n.

Figs 1 , 2 , 3

Type species.

Colemanus keeleyorum Fisher, sp. n.

Diagnosis.

There are several similarities between Colemanus and other braconids, specifically Cardiochilinae and Cheloninae . Like Colemanus , some Cheloninae have a recurved 3RSb and an evenly curved 1M. However, chelonines possess a metasomal carapace. Colemanus contortus comb. n. lacks a carapace ( Brues 1933) and although the metasoma of C. keeleyorum sp. n. is not completely visible, it seems apparent that it also lacks a metasomal carapace (Fig. 1A ). Thus, the lack of a metasomal carapace distinguishes Colemanus from Cheloninae . Colemanus also resembles many Cardiochilinae in having a recuved 3RSb and heavily sculptured mesosoma, but can be readily distinguished by 3RSb remaining tubular as it reaches the wing margin.

Colemanus can be distinguished from other Ichneutinae s. l. by the presence of a curved 3RSb; fore wing 1cu-a curved downward, not angled toward wing margin; hind wing M+Cu positioned in the posterior half of the wing; and a heavily sculptured mesosoma.

Etymology.

Named for bodybuilder Ronnie Coleman, who was famous for his back; referring to the robust and sculptured nature of the mesosomal dorsum.

Figures

Figure 1. Fossil of Colemanus keeleyorum sp. n.: A whole fossil B close-up of mesosoma C close-up of left pterostigma, note preserved microtrichia.

Figure 2. Reconstructions of Colemanus keeleyorum sp. n.: A dorsal mesosoma, note heavy sculpturing B wings, note last abscissa of forewing radial sector (3 RSb) is recurved.

Figure 3. Placement of Colemanus gen. n. with known relationships of extant taxa: A-C represent the three possible relationships between included Proteropini. Numbers on branches represent total tree length when Colemanus is placed at that location. Circled numbers are most parsimonious placements (most parsimonious in orange; second-most in blue). Number boxes are total tree lengths of that topology when Colemanus is excluded. Note that lowest tree lengths are always achieved when Colemanus is placed within Proteropini.