Angustopila elevata (Thompson & Upatham, 1997)

Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne, 2023, Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae), ZooKeys 1147, pp. 1-177 : 1

Angustopila elevata (Thompson & Upatham, 1997)

Figs 12 , 13

Systenostoma elevata Thompson & Upatham, 1997: 232-233, figs 39-43.

Angustopila elevata - Jochum et al., 2014: 27.

Angustopila elevata - Páll-Gergely et al., 2015a: 33, fig. 11 (silhouette for size comparison with other species).

Angustopila subelevata Páll-Gergely et al., 2015a: 39, fig. 4. new synonym.

Angustopila subelevata Páll-Gergely et al., 2017: 332, figs 1B, 2A-G, 7E, F.

Type locality.

"Thailand, Chiang Mae Province, Doi Chiang Dao (Mountain), 7 km west of Chiang Dao; 600 m altitude (19°24.3'N, 98°54.2'E)" ( A. elevata ); China, Guangxi (广西), Hechi Shi (河池市), Bama Xian (巴马县), cliffs at the southern edge of Jiaole Cun (交乐村), 590 m, 24°7.045'N, 107°7.847'E " ( A. subelevata ).

Material examined.

China • 23 shells; Guangxi, Bose Shi, Leye Xian, ca. 2 km from Leye towards Huaping Zhen , cave above Niuping junction (locality code: 2013/85); 24°47.61'N 106°31.89'E; 590 m a.s.l.; 8 Sep. 2013; A. Hunyadi & M. Szekeres leg.; coll. HA • 3 shells; Guangxi, Hechi Shi, Huanjiang Maonanzu Zizhixian, south of Mulun Guojiaji Ziran Baohuqu , rocks above Dongning (locality code: 2013/99); 25°05.97'N 107°57.64'E; 530 m a.s.l.; 17 September 2013; A. Hunyadi & M. Szekeres leg.; coll. HA .

Myanmar • 1 figured shell; Shan State, 7.4 km from the centre of Hoponh towards Namsang along road no. 4, ca. 5 km north, Parpant Cave (locality code: 2018/36); 20°50.96'N, 97°14.26'E; 1170 m a.s.l.; 6 Oct. 2018; A. Hunyadi, K. Okubo & J.U. Otani leg.; HNHM 103484 • 6 shells + 2 j/b shells; same data as for preceding shell; coll. HA • 1 shell; Shan State, Hopong, Hopong Spring and cave (locality code: JG/2019/2); 20°49.05'N, 97°13.49'E; 2 Feb. 2019; J. Grego leg.; coll. JG .

Thailand • 1 figured shell; Tak Province, 6 km NNW Tha Song Yang , left side of road no. 105 (locality code: 2015/23); 17°15.36'N, 98°12.65'E; 140 m a.s.l.; 15 Feb. 2015; A. Hunyadi leg.; HNHM 100183 • 7 shells; same data as for preceding shell; coll. HA • 1 figured shell; Chiang Mai Province, north-eastern part of Doi Chiang Dao, Wat Tham Chiang Dao northwest 2 km; 19°24.02'N, 98°54.68'E; 835 m a.s.l.; 7 Feb. 2015; A. Hunyadi leg.; HNHM 100184; • 11 shells; same data as for preceding shell; coll. HA • 18 shells + 4 j/b shells; Chiang Rai Province, 7 to 9 km SSW of Mae Sai, around Wat Tham Pla (locality code: 2015/18); 20°19.72'N, 99°51.82'E; 400 m a.s.l.; 12 Feb. 2015; A. Hunyadi leg.; coll. HA • 1 figured shell; Nakhon Si Thammarat Province, Wat Tham Thong Panara, vicinity of the temple (locality code: 2015/31); 08°25.28'N, 99°22.76'E; 40 m a.s.l.; 19 Feb. 2015; A. Hunyadi leg.; HNHM 100180; • 7 adult + 2 j/b shells; same data as for preceding shell; coll. HA .

Laos • 56 adult + 6 j/b shells; Luang Namtha Province, 20.6 km southeast from centre of Vieng Phou Kha, 1.5 km south of Ban Phou Lek , right side of road (locality code: 2019/121); 20°32.21'N, 101°08.10'E; 785 m a.s.l.; 8 Oct. 2019; A. Hunyadi leg.; coll. HA • 70 adult + 22 j/b shells; Luang Namtha Province, 19.8 km southeast from centre of Vieng Phou Kha, south from Ban Phou Lek, southern edge of quarry (locality code: 2019/122); 20°32.54'N, 101°07.96'E; 780 m a.s.l.; 8 Oct. 2019; A. Hunyadi leg.; coll. HA • 21 shells; Udomxai Province, 10 km south of centre of Na Mor , 3.8 km east-southeast of Na Xay , rock wall facing north (locality code: 2019/119); 20°53.37'N, 101°48.96'E; 660 m a.s.l.; 7 Oct. 2019; A. Hunyadi leg.; coll. HA • 141 adult shells + 21 j/b shells; Luang Namtha Province, 43.8 km from centre of Luang Namtha towards Vieng Phou Kha, Phou Lan , 100 m from left side of road (locality code: 2019/123); 20°44.53'N, 101°11.10'E; 770 m a.s.l.; 8 Oct. 2019; A. Hunyadi leg.; coll. HA • 12 adult shells; Luang Prabang Province, 3.1 km from centre of Nong Khiaw towards Pak Xeng, Tham Pha Toke , around the cave (locality code: 2019/114); 20°33.22'N, 102°37.72'E; 345 m a.s.l.; 5 Oct. 2019; A. Hunyadi leg.; coll. HA • 3 adult shells; Luang Namtha Province, 48 km southwest from centre of Luang Namtha towards Vieng Phou Kha, Nam Eng, Tham Kao Rao (locality code: 2019/120); 20°43.45'N, 101°09.26'E; 745 m a.s.l.; 7 Oct. 2019; A. Hunyadi leg.; coll. HA • 1 shell; Khammouane Province, Tham Kong Lor (cave), 60 km direct N of Thakhek, at the base of limestone rocks above Recreational Area (locality code: La.2); 17°57.39'N, 104°45.65'E; 200 m a.s.l.; Mar. 2010; A. Reischütz leg.; NHMW-MO-112005 • 2 complete + 1 broken shells; same data as for preceding; coll. RE • 1 shell; Vientiane Province, Vang Vieng , rocks at Vang Vieng Resort , by river Nam Song , limestone rocks at Tham Chang cave (locality code: La.12); 18°54.66'N, 102°26.42'E; 280 m a.s.l.; Mar. 2010; A. Reischütz leg.; coll. RE • 1 shell; Vientiane Province, 4.5 km west from centre of Vang Vieng, Phone Ngeun, Tham Khan Kham (locality code: 2019/133); 18°55.53'N, 102°24.95'E; 280 m a.s.l.; 14 Oct. 2019; A. Hunyadi leg.; coll. HA .

Vietnam • 18 adult shells; Sơn La Province, Thu ận Chau District , 900 m southeast from centre of Co M ạ, rock wall above unpaved road (locality code: 2020/18); 21°21.19'N, 103°31.68'E; 1300 m a.s.l.; 8 Feb. 2020; A. Hunyadi leg.; coll. HA • 1 shell; Sơn La Province, Mộc Châu, Hang Dơi, around the entrance of the cave (locality code: 2020/26); 20°50.96'N, 104°38.34'E; 865 m a.s.l.; 11 Feb. 2020; A. Hunyadi leg.; coll. HA • 4 adult shells; Lạng Sơn Province, Bình Gia, eastern edge of the village, Di Chi Keo Leng , rock wall (locality code: 2020/48); 21°56.30'N, 106°23.82'E; 375 m a.s.l.; 17 Feb. 2020; A. Hunyadi leg.; coll. HA • 1 shell; Sơn La Province, Qu ỳnh Nhai district , 20 km north from cross to Thu ận Châu, Chi ềng Khoang, cave above the village (locality code: 2020/9); 21°33.44'N, 103°40.91'E; 315 m a.s.l.; 7 Feb. 2020; A. Hunyadi leg.; coll. HA .

Diagnosis.

A small to medium-sized Angustopila species with relatively irregularly coiled, slightly concave-conical shell, and a toothless, rather subquadrate aperture. Individuals in most populations possess widely spaced radial ribs the on the ventral side and on the body whorl in apertural view.

Measurements (in mm).

H = 0.8-0.97, D = 0.74-0.88, H/D*100 = 95.4-123 (n = 58), RUD = 24.1-31.6 (n = 20).

Differential diagnosis.

See also under A. apiostoma sp. nov., A. halongensis sp. nov., A. oostoma sp. nov., and A. werneri sp. nov.

Distribution.

A. elevata is the most widely distributed Angustopila species, inhabiting Myanmar’s Shan State, northern and southern Thailand, northern Laos, northern Vietnam, and southern China (Guangxi) (see Fig. 14 ).

Remarks.

When Angustopila subelevata was described, we relied on the original description of A. elevata only. Although it was evident that the two species were very similar, we considered the differences ( A. elevata had a more slender shell, a deeper umbilicus, and was lacking spiral striae on its base) and the nearly 1000-km distance sufficient to describe A. subelevata as new. The newly collected shells from the type locality of A. elevata showed that it had weak spiral striae on the base. Furthermore, the geographical populations examined between the type localities of A. elevata and A. subelevata indicated that only minor conchological differences exist. Although the area spans nearly 2000 kilometres in a straight line from southern Thailand to Guangxi Province in China, the populations listed here show little variation in terms of conchological characters. Thus, we now treat them as a single species.

Figures

Figure 12. Angustopila elevata (Thompson & Upatham, 1997) (specimen from the type locality, HNHM 100184). Apertural (A), ventral (B), lateral (C) and apical (D) sides of the shell; granular sculpture on the protoconch (E), ventral (F) and frontal (G) surface of the body whorl.

Figure 13. Angustopila elevata (Thompson & Upatham, 1997) A Perpant Cave (Myanmar) B Chiang Dao C Tha Song Yang D Panara E 2019 / 123 F 2019 / 119 G 2020 / 18 H 2020 / 26 I Cong Troi Cave J 2020 / 52 K 2020 / 48 L A. subelevata holotype M La. 2.

Figure 14. Distribution of Angustopila elevata (Thompson & Upatham, 1997).