Daedalma boliviana boliviana Staudinger

Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, Zootaxa 2898, pp. 1-68 : 15-16

Daedalma boliviana boliviana Staudinger

( Figs. 2C, 2D, 9D, 15A)

Daedalma dinias Hewitson var. rhomboidea Weymer, 1912: 266 , pl. 56, row f (Synonymy established by Pyrcz, 2004: 509); Gaede, 1931: 509.

Daedalma dinias boliviana Hewitson ; Forster, 1964: 178.

Material examined: BOLIVIA: LECTOTYPE male: Río Tanampaya, Bolivia (La Paz), 1894, Garlepp, square pink label saying: Origin; rectangular red label saying : Lectotype, Daedalma dinias v. boliviana Staudinger , designated by Lee D. Miller, 1989, ZMHB (Designated herein) ; PARALECTOTYPE female: Cillat., Bol. , Garl., square label saying: Origin; rectangular label saying : Paralectotype, Daedalma dinias v. boliviana Staudinger , designated by Lee D. Miller, 1989, ZMHB (Designated herein) ; 2 males: no locality, B. Mery leg., TWP ; 1 male: Coroico , X.1993, TWP ; 2 males: Coroico , 30.IX.1993, TWP ; 20 males: Yungas de La Paz, ex. Staudinger & Bang- Haas, 2 TWP , 18 SMTD; 1 male: Prov. Cochabamba, Locotal , 2200–2300 m, 13.VIII.2000, T. Pyrcz leg., TWP ; 1 female: via Cochabamba, 1900 m, VIII.2000, T. Pyrcz leg., TWP ; 3 males and 1 female: Prov. Cochabamba, Río Vinto , 2350 m, 15–17.II.2009, T. Pyrcz & J. Wojtusiak leg., MZUJ ; 1 male: 213, no other data, USNM ; 1 male: col. B. Neumögen, collection Brklyn Mus., 209, USNM ; 2 males: Chaparé, Prov. Cochabamba, El Limbo, 2200 m, USNM ; 1 male: Cochabamba, Carrasco, Chaqui Sacha, Corte , 1835 m 16.XI.2007, A. Cespedes leg., MNKM ; 1 female: [ Bolivia ], M85/990, M. Adams coll., BMNH ; 1 male: Inca Chaca , Cochabamba, 2500 m, no. 5557, J-C. Petit leg. ; 1 female: same data but 5466, JCP ; PERU: 1 male: S-E Peru [Puno], Agualani , 10000 ft, 06.X.[19]02, Ockenden, Rothschild Bequest, BM 1939-1, BMNH ; 1 male: Puno, Oroya to Limbani , I.1901, G. Ockenden, Rothschild Bequest, BM 1939-1, BMNH ; 1 male: [ Bolivia ], M85/936, M. Adams coll., BMNH ; 4 males: Puno, Carcel Punco, Agualanes 8 Km Nord Limbani , 2400–2700 m, 16.X.2004, P. Boyer leg., TWP ; 1 male: Puno, Agualanes, Limbani , X.2004, J. Bottger leg., TWP ; 3 males: Puno, Est de Pacchani , 2500–3000 m, J. Bottger leg., TWP ; 1 female: Uruhuasi, S. Peru , 7000 feet, 04.V.1910, H. & C. Watkins, Adams Bequest, 1912-09, BMNH .

Redescription: MALE ( Fig. 2C): Head: eyes chocolate brown; labial palpi medium brown covered with yellow and brown hair; antennae dorsally medium brown, ventrally light brown, club orangeish. Thorax: blackish brown, legs medium brown. Abdomen: dorsally and laterally blackish brown, ventrally dull brown. Wings: FWD dark brown, basal one-third slightly lighter, lustrous; a large rhomboid light orange patch extending from outer one-third of discal cell, along vein Cu2 to subapical and submarginal area; fringes alternately dark brown and milky white, latter denser in apical area. HWD uniform dark brown, lustrous; fringes alternately milky white and dark brown in apical area, elsewhere dark brown. FWV: ground colour dark brown, duller and slightly lighter in basal one-third; orange patch slightly lighter and smaller than on upperside, towards costa suffused with brown scales, in most individuals enclosing a brown dot in cell Cu1-Cu2; subapical and apical areas chocolate brown, milky white and magenta with two dark brown minute ocelli in R4-R5 and R5-M1 and a short, sinuate dark brown submarginal line. HWV: a nearly nondescript pattern of chocolate brown, crimson brown, chestnut and milky white bands and stripes with a two lunular silvery submarginal patches in M2-M3 and M3-Cu1 and a series of seven small submarginal dark brown ocelli pupiled with milky white, one in each cell and two in Cu2-1A. Genitalia ( Fig. 9D): Uncus thin, long and straight; gnathos adhered to uncus, half its length; saccus intermediate in depth; valvae conical with a sharp apical tip pointing straight; aedeagus straight, smooth, slightly longer than valva.

FEMALE ( Fig. 2D): Female differs from peruviana by larger FWD red-orange patch, which extends slightly further distally; both nominate subspecies and peruviana possess a brown dot enclosed within this patch in cell Cu1-Cu2; orange markings on HWD are somewhat variable but wide submarginal band is more dusted with red and is longer than in peruviana, reaching to vein Cu2. Genitalia ( Fig. 15A): Papillae anales large, their posterior edge irregular, apophyses posteriores very short, triangular, hump on lamella postvaginalis widely rounded when seen from side, almost completely filling interior of sinus vaginalis, wall of lamella postvaginalis strongly sclerotized, anteriorly smooth, with small transverse wrinkles on its posterior part. Pear-like bursa copulatrix, gradually narrowing posteriorly and transforming into ductus bursae. Ductus bursae of same length as antrum, appearing twisted.

Remarks: Staudinger, in the orginal description of D. boliviana , refers to specimens from Bolivia and central Peru (Chanchamayo). Pyrcz (2004) recognised the populations from Amazonas and Pasco (north-central Peru) as a separate subspecies, peruviana, but did not sort out the problem of lectotype. Miller selected one male in the Staudinger collection deposited in ZMHB to be the lectotype, but did not publish this designation. We accept his selection and in order to stabilise the nomenclature, we formally designate this specimen as the Lectotype. Nominate D. boliviana is recognised primarily by a larger FWD orange patch than in the other subspecies. It occurs in southern Peru (Cuzco, affluents of the Río Madre de Dios; Puno) and Bolivia (Yungas de La Paz and Yungas de Cochabamba).