Senecio doriiformis subsp. orientalis (Fenzl) Matthews

Calvo, Joel, Álvarez, Inés & Aedo, Carlos, 2015, Systematics of Senecio section Crociseris (Compositae, Senecioneae), Phytotaxa 211 (1), pp. 448-450 : 448-450

9b. Senecio doriiformis subsp. orientalis (Fenzl) Matthews in Davis (1975: 434). Senecio nemorensis var. orientalis Fenzl in Tchihatcheff (1860: 294). Senecio doriiformis var. megalophron Fenzl ex Boissier (1875: 407) , nom. illeg. superfl. ( McNeill et al. 2012, ICN Art. 52.1). Senecio doriiformis var. orientalis (Fenzl) Handel-Mazzetti (1913: 437) . TYPE: Turkey, Mersin, région montagneuse supérieure du Taurus, près de Gulek-Maden, [37º16'N 34º46'E], 16 Aug 1855, B. Balansa 654 (lectotype, designated here, G-96309 image!; isolectotypes, C!, E-287936 image!, G-96310 image!, GOET-2037 image!, LE!, WAG-646 image!).

Senecio megalophron Fenzl , nom. nud., in sched. ( McNeill et al. 2012, ICN Art. 38.1) (B-10-0325113!, G-162718!, H-1441895!, LE!, M-30305 image!, UPS!).

Middle cauline leaves 8–13 cm long, 1.7–3.7 cm wide, lanceolate (ratio middle leaf width / middle leaf length = 0.20–0.43), sessile to attenuated into a petiole up to 3 cm long, not asymmetric or weakly so; upper cauline leaves 3.7–10.5 cm long, 0.4–2.5 cm wide, lanceolate (ratio upper leaf width / upper leaf length = 0.11–0.36), sessile, sometimes attenuated into a petiole up to 0.3 cm.

Iconography: — Nordenstam (1989b, tab. 40).

Distribution and habitat: — Iran?, Turkey (southern central-east Anatolia); rocky places, woods of Cedrus libani , on calcareous soils; elevations of 1000–2500 m ( Figure 13 ).

Phenology: — Flowering from June to July.

Discussion: — This subspecies has been misidentified with S. pseudoorientalis . In fact, some sheets of the exsiccate Kotschy 121, under the name S. megalophron , correspond to S. pseudoorientalis (B-10-0325112!, BR-505713!, M-30304 image!, MPU-12833 image!, MPU-12834 image!, S!, WU!). Although they can be easily

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CALVO ET AL.

distinguished (size capitula, number of involucral bracts, etc.), misidentification may occur due to their overlapping ranges in some regions such as in the Taurus Mountains. Consequently, the syntype Kotschy 121 should be rejected as original material.

Nordenstam (1989b) cited a Haussknecht’s collection (herbarium not specified) from Avroman and Shahu (Kermanshah Province, northwestern Iran), under the name S. doriiformis subsp. orientalis . Likewise, Lotfi et al. (2010) recorded this taxon from the Iranian Kurdestan Province (Mozaffarian 74827, TARI), and Kermanshah Province (Tavakoli & Mirabdali 2900, TARI). Unfortunately, we cannot study the mentioned collections, thus, we include Iran with a question mark in the distribution. Taking into account that the nearest population (that from Pira Magrun, northeastern Iraq; about 140 km away) corresponds to S. doriiformis subsp. doriiformis , we have some doubts about the identifications. Further collections are needed to elucidate the proper identity of the Iranian populations.

Selected specimens examined. TURKEY. Adana: Karaisali, Koca Çukur Yaylasi-Katir gediği, 37º25’N, 35º1’E, 29 June 1959, A . Huber-Morath 15840 ( G) . Antalya: Ak Dağ ( S of Geyik Dağ ), 36º52’N, 32º10’E, 28 Aug 1947, P. H . Davis 14331 ( C, G, H, K) . Konya: Tauri alpes, BulgarDagh , versus plumbi fodinas, 37º26’N, 34º20’E, 11 Aug 1853, T . Kotschy 251 ( B, BR, K, LE, S, UPS, WU) . Malatya: Taurus Cataonicus , inter urbem Malatja et vicum Kjachta , 38º3’N, 38º2’E, 16 July 1910, H. F . Handel-Mazzetti 474 ( WU); Dedeyasi, Taslch, 38º13’N, 37º51’E, 17 Aug 1966, H . Peşmen 1069 ( G). Maraş: Akher Dagh, 37º39’N, 36º59’E, July 1907, M . Haradjian 1694 ( G, LE); Akher Dagh, 37º39’N, 36º59’E, Sep 1907, M . Haradjian 1738 ( E, G, K, LE); Akherdagh, 37º39’N, 36º59’E, 15 Sep 1884, B . Post 119 ( G, US) . Mersin: Tauri Alpes, BulgarDagh , in valle Karli Boghas , 37º9’N, 34º27’E, 13 July 1853, T . Kotschy 121 ( G, K, LE, UPS); Güllek, 37º15’N, 34º46’E, July 1896, W . Siehe 667 ( G, K, LE, S, WU). Niğde: in vicinitate oppidi Çamardi, montes Ala Dağlar, loco Demir-kazik dicto, 37º51’N, 35º6’E, 9 Aug 1992, V . Vašák s.n. ( BR). Van: Çatak, Dalbasti köyü, 37º54’N, 42º55’E, 20 July 2002, B . Bani 1255 ( GAZI); Artos Dag above Gevas, 38º15’N, 43º3’E, 14 July 1954, P. H . Davis & O . Polunin 22690 ( ANK); Karduchia, ad fontes hospitium Chana Putkie, 38º7’N, 42º48’E, Sep 1859, T . Kotschy 626 ( S, UPS) .

Figures

FIGURE 13. Distribution map for Senecio doriiformis Candolle subsp. doriiformis (●), and S. doriiformis subsp. orientalis (Fenzl) Matthews (▲).