Eupholidoptera pallipes Willemse & Kruseman, 1976

Willemse, Luc, Tilmans, Jos, Kotitsa, Nefeli, Trichas, Apostolos, Heller, Klaus-Gerhard, Chobanov, Dragan & Ode, Baudewijn, 2023, A review of Eupholidoptera (Orthoptera, Tettigoniidae) from Crete, Gavdos, Gavdopoula, and Andikithira, ZooKeys 1151, pp. 67-158 : 67

publication ID	https://dx.doi.org/10.3897/zookeys.1151.97514
publication LSID	lsid:zoobank.org:pub:5FEDE55D-C9AF-47D5-9125-9F1758AE2A18
persistent identifier	https://treatment.plazi.org/id/C590E8C4-2654-5503-811C-C9719D1E0287
treatment provided by	ZooKeys by Pensoft
scientific name	Eupholidoptera pallipes Willemse & Kruseman, 1976
status

Treatment

Eupholidoptera pallipes Willemse & Kruseman, 1976 View in CoL

Figs 9 View Figures 8–10 , 13 View Figures 11–24 , 27 View Figures 25–38 , 41 View Figures 39–52 , 55 View Figures 53–66 , 71 View Figures 69–82 , 85 View Figures 83–96 , 99 View Figures 97–110 , 113 View Figures 111–125 , 128 View Figures 126–139 , 142 View Figures 140–153 , 156 View Figures 154–167 , 170 View Figures 168–181 , 184 View Figures 182–197 , 200 View Figures 198–212 , 252 View Figures 247–253

Eupholidoptera pallipes Willemse & Kruseman, 1976: 135.

Eupholidoptera pallipes Morphological description. Willemse and Kruseman 1976: 135, 136.

Examined specimens.

Holotype, allotype, 5 ♂ (paratypes) (for details see Suppl. material 2).

Diagnostic features.

Frontal part of head (Fig. 13 View Figures 11–24 ) pale with two large black patches joined or not into a transverse band; pronotum (Fig. 27 View Figures 25–38 ) pale except for small black patch in the rear of the side flap; abdomen pale, proximal margins tergites black. Male - stridulatory file with 94 teeth (including proximal and distal ones), density of teeth in middle two thirds of the file 22 teeth per mm; anal tergite (Figs 71 View Figures 69–82 , 85 View Figures 83–96 , 99 View Figures 97–110 ) wide with central groove, distally bend downward, forming two spines pointing downward and inward almost touching each other separated by rectangular excision; cerci (Figs 113 View Figures 111–125 , 128 View Figures 126–139 ) 5 × longer than wide, cylindrical, central third conical, weakly curved inward and upward, armed with sharply pointed inward curved sub-basal tooth; subgenital plate (Figs 142 View Figures 140–153 , 156 View Figures 154–167 ) as wide as long, proximally widest, sides rimmed almost up to apex, in profile distally narrowing, straight, pointing backward, tip apical lobes truncate with protuberance on the inner margin and strong upward and backward pointing curved spine at base of stylus, with V-shaped excision along one third of total length; styli (Fig. 170 View Figures 168–181 ) long, more than half as long as cerci, 3 × longer than wide, cylindrical, inserted at tip of apical lobes, pointing backward and outward; titillator (Figs 184 View Figures 182–197 , 200 View Figures 198–212 ) symmetrical, weakly sclerotised, basal arms short, apical arms completely fused, widening in basal third, gradually narrowing in apical two thirds, near tip widening again, tip rounded at either side with a tiny thorn, in profile equally wide, straight, in apical half weakly curved upward. Female (Fig. 252 View Figures 247–253 ) - subgenital plate (Figs 41 View Figures 39–52 , 55 View Figures 53–66 ) as long as wide, proximally convex, apical lobes rectangularly rounded separated by wide concave median excision along quarter of total length, in profile rhomboid, apically truncate, upper angle widely rounded.

Measurements.

See Tables 6 View Table 6 , 7 View Table 7 .

Bioacoustics.

The song of this species has not yet been recorded.

Differential diagnosis.

Males differ from congenerics in the stout, straight cerci (Figs 113 View Figures 111–125 , 128 View Figures 126–139 ) with a sub-basal rectangular side-tooth, in the subgenital plate (Figs 142 View Figures 140–153 , 156 View Figures 154–167 ) narrowing into truncate tips with an upward and backward pointing spine, a protuberance on the inner margin of the excision, in the long, apically inserted, backward pointing styli (Fig. 170 View Figures 168–181 ), in the anal tergite (Figs 71 View Figures 69–82 , 85 View Figures 83–96 , 99 View Figures 97–110 ), medially bent downward forming an excision adjoined by two inward and downward pointing spines and in the titillator (Figs 184 View Figures 182–197 , 200 View Figures 198–212 ) with short basal arms and completely fused apical arms, apically widened, tip rounded at either side with a tiny thorn. Females differ from congenerics in the subgenital plate (Figs 41 View Figures 39–52 , 55 View Figures 53–66 ), as long as wide, convex hind margin centrally with wide excision a deep as quarter of the length. Eupholidoptera pallipes closely resembles E. gemellata but males differ in the apical arms of the titillator apically being fused in E. pallipes (Figs 184 View Figures 182–197 , 200 View Figures 198–212 ), its tip with a tiny lateral thorn, in E. gemellata (Figs 183 View Figures 182–197 , 199 View Figures 198–212 ) being adjoined the tip being bare. Females of both species differ in the shape and the hind margin of the subgenital plate (compare Figs 40 View Figures 39–52 , 41 View Figures 39–52 ). In colouration E. pallipes differs from congenerics, except E. gemellata , in the general pale colouration in particular of the legs and the narrow anterior transverse black band in the abdominal tergites (Fig. 9 View Figures 8–10 ). For more details differentiating E. pallipes from other Cretan Eupholidoptera see Table 5 View Table 5 .

Distribution.

The type series was collected in 1973 on Mt. Lefka at the saddle of Linoseli above Xyloskalo between 1600 m and 1800 m. Additional specimens were collected in pitfall traps operated in the summer of 1991 more to the east on Mt. Lefka above Limnia (Fig. 255 View Figures 254, 255 ). On 13 October 2017 a female was photographed just below the Pakhnes peak at 2440 m. For a complete list of localities, specimens and repositories see Suppl. material 1.

Habitat.

Rocky mountain slopes with phrygana between 1600 and 2440 m.

Phenology.

The type series was collected 5 August 1973. Pitfalls above Limnia trapped adults between early August and early September and during the entire month of October whereas a trap operated between early June and early July only contained nymphs.