Takobia maxillare (Braasch & Soldan , 1983)
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https://dx.doi.org/10.3897/zookeys.1071.71582 |
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lsid:zoobank.org:pub:BEA57970-A281-49DE-9F9B-D88910A73824 |
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https://treatment.plazi.org/id/C615D200-1E6F-5C11-9A49-DE22590F3564 |
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scientific name |
Takobia maxillare (Braasch & Soldan , 1983) |
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Takobia maxillare (Braasch & Soldan, 1983)
Figs 1A, D View Figure 1 ; 3 View Figure 3 , 4 View Figure 4
Differential diagnosis.
Takobia maxillare can be easily separated from other related species by the combination of the following characters: 1) maxillary palp highly developed with the segment I widened apically and segment II straight; 2) labrum dorsally covered with numerous setae, none of them arranged in a row; 3) right prostheca reduced, apically bifid; 4) labial palp segment III quadrangular, asymmetrical, with a short projection lateroapically; 5) claw edentate, subequal to 1/2 of corresponding tarsus; 6) paraproct with a short prolongation bent ventrally.
Description of nymph.
Length. Female body 6.8-9.1 mm; cerci 4.6-5.4 mm; median caudal filament 3.4-4.5 mm; male body 5.0-7.2 mm; cerci 4.2-5.8 mm; median caudal filament 2.5-3.5 mm.
Coloration and texture. General coloration brown (Fig. 1A, D View Figure 1 ). Head uniformly brown with vermiform marks visible on vertex and frons in some specimens. Turbinate eyes in male nymphs purple-brown. Legs ecru. Thorax brown with some areas of darker coloration. Abdominal tergites medium brown without any pattern. Abdominal sternites light brown. Cerci ecru to light brown without bands or pattern. Original coloration probably faded after more than 40 years of storage in alcohol. Surface of body shagreened, most pronounced on head capsule and thorax (Fig. 4C View Figure 4 ).
Head. Antennae close to each other, with a narrow interantennal carina; scape and pedicel with V-shaped scale insertions and sparse setae. Dorsal surface of labrum (Fig. 3A View Figure 3 ) evenly covered with numerous long setae and scattered small fine setae, distolateral arc of more prominent setae not distinguishable, almost no setae present along midline; ventral surface with row of ca. ten submarginal small, pointed setae laterally; distal margin fringed with ca. 12-17 short, followed by 8-12 long, feathered setae. Right mandible (Figs 3C, D View Figure 3 ; 4A View Figure 4 ) with sparse fine setae; incisors composed of eight pointed denticles (in nymphs long after molting, denticles become worn out and rounded), outer and inner incisor group with four denticles each; row of short fine setae along inner margin of incisors present; prostheca reduced and apically asymmetrically bifid (this bifurcation very inconspicuous, see Fig. 4A View Figure 4 ), slightly feathered; margin between prostheca and mola with tuft of fringed setae. Left mandible (Fig. 3B, E View Figure 3 ) with sparse fine setae; incisors composed of seven apically pointed denticles, outer and inner incisor groups not distinctly separated; prostheca with denticles and comb-shaped structure; margin between prostheca and mola with short, fringed setae. Hypopharynx apically covered with thin setae; lingua with central small protuberance; superlingua slightly longer than lingua. Maxilla (Fig. 3F View Figure 3 ) with incisors composed of three elongated and curved teeth; crown with two rows of setae, ventral one with small setae, dorsal row with three long stout dentisetae (apical dentiseta similar to maxillary teeth, relatively narrow); maxillary palp very long, nearly 3 × longer than galeolacinia, two-segmented, length of segment I nearly double length of galeolacinia, length of segment II subequal to segment I; segment I widened apically and slightly curved outward; segment II apically rounded; both segments with numerous thin setae, most dense along inner margin. Labium (Fig. 3G View Figure 3 ) with glossae subequal to paraglossae; both inner and outer margins of glossae with row of pointed setae, dorsal surface of glossae with well-defined group of fine setae subapically; ventral surface of glossae with group of long setae extending from basal part of glossa along its inner margin to apex; paraglossae with two rows of long, stout setae apically; labial palp three-segmented; segment I slightly shorter than segments II and III combined; segment II with very small medioapical protuberance and dorsal oblique row generally of six long setae; segment III asymmetrical, with medioapical part widely rounded and short projection lateroapically; all segments of labial palp with hair-like setae, present only occasionally on segments I and II, most dense on ventral surface of segment III; several distinct stout pointed setae present along apical part of segment III.
Thorax. Forelegs (Fig. 3H View Figure 3 ). Trochanter with ca. five marginal spine-like setae. Femur dorsally with one row of 16-18 medium, stout setae; additional dorsoapical setal patch formed by another 6-9 stout, medium setae; ventral margin with numerous stout, pointed short setae, some of these setae on lateral margin subparallel to ventral margin, villopore absent; lateral margin with occasional short hair-like setae and V-shaped scale bases with scales (not figured in Fig. 3H View Figure 3 ). Tibia with many setae along ventral margin and group of setae apically; scarce setae also elsewhere on surface of tibia and along dorsal margin; tibiopatellar suture present; lateral margins with scales and numerous scale bases. Tarsus with row of ca. 20 small, pointed setae on ventral margin; lateral margins with numerous scale bases. Tarsal claw (Fig. 4B View Figure 4 ) very slightly hooked, without any teeth, apical setae present, minute; length of tarsal claw subequal to 1/2 of tarsus; Mid and hindleg (Fig. 3I, J View Figure 3 ) similar to foreleg, except setae along ventral margin of femora less robust and more scarce in mid and hindlegs compared to forelegs. Hindwing pads present.
Abdomen. Tergites (Fig. 4E View Figure 4 ) not shagreened, with numerous V-shaped scale bases, scales (rounded apically), and thin hair-like setae; distal margin of tergite I with few occasional small triangular spines, tergites II-X with well-developed row of triangular spines (length:width ratio of spines in middle part of segment IV ca. 3:1-3:2); row of triangular spines on tergite X not interrupted in middle. Sternites (Fig. 4F View Figure 4 ) with scales, scale bases and setae similar to tergites; distal margin of sternites III-IX with row of triangular spines, this row interrupted in middle in sternite III and also interrupted in places of emerging gonostyli in male nymphs; triangular spines in lateral parts of sternite generally narrower than in central part of same sternite. Gills (Fig. 3K-Q View Figure 3 ) on segments I-VII, slightly asymmetrical, widened in distal portion, widely rounded apically; dorsal surface with scales and scale bases submarginally (Fig. 4D View Figure 4 ); tracheation faintly visible; margins serrated in distal 1/2, with row of fine setae; gill VII similar to gills II to VI. Paraproct (Fig. 4G View Figure 4 ) with abundant scales and scale bases (of same shape as on tergites and sternites); distinct prolongation bent dorsally; paraproct margin with ca. six or seven triangular spines laterally from prolongation and numerous slightly smaller spines medially from prolongation; prolongation margined with ca. ten elongated medium spines, with spines also on ventral surface; cercotractor with scales and scale bases, margin with triangular spines.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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