Frullania ericoides var. verrucosa, (Kamim.) Hentschel et von Konrat, comb., 2015

Hentschel, Jörn, Konrat, Matt Von, Söderström, Lars, Hagborg, Anders, Larraín, Juan, Sukkharak, Phiangphak, Uribe, Jaime & Zhang, Li, 2015, Notes on Early Land Plants Today. 72. Infrageneric classification and new combinations, new names, new synonyms in Frullania (Marchantiophyta), Phytotaxa 220 (2), pp. 127-142 : 134-135

Frullania (subg. Frullania sect. Frullania ) ericoides var. verrucosa (Kamim.) Hentschel et von Konrat, comb.

nov. Basionym:— Frullania squarrosa var. verrucosa Kamim., J. Hattori Bot. Lab. 24: 19, 1961 ( Kamimura 1961).

Type:— JAPAN. Kochi: Tosa, 20 February 1948, Kamimura 1321 (NICH-46849, isotype). Note:— Kamimura (1961) noted that the most diagnostic character of this variety is the verruculose cuticle of the leaf cells together with the propagules, which sometimes are seen on the ventral surface of the leaves. Frullania ericoides ( Nees 1833: 346) Montagne (1839: 51) is a polymorphic species and Hentschel et al. (2009) indicated that at least two independent lineages are present. Further investigation of the complex is required.

Frullania (subg. Microfrullania) fertilis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 235, 1855 ( De Notaris 1857). Type:— CHILE. Valparaiso: Puccio ( FH).

= Frullania magellanica var. diminutiva Herzog, Arch. Esc. Farm. Fac. Sci. Med. Cordoba 7: 29, 1938 ( Herzog & Hosseus 1938), syn. nov. Type:— CHILE. Valdivia: Quitaluto, Herzog 679 ex p. (JE!, lectotype here designated); Villarica: Pucón, Herzog 204!; Chiloé: Llanquihué, Petrohué, Herzog 522). Note:—Only two of the three syntypes mentioned by Herzog and Hosseus (1938) are located at JE. One of them, Hosseus 204, was revised by G. G. Hässel de Menéndez in 1981 as belonging to F. fertilis . Due to the fact that Hosseus 679 ex p. is fertile (c.per., c.spor, ♂) the second syntype was chosen as the lectotype. Since F. magellanica frequently has been confused with F. fertilis, Hässel de Menéndez & Rubies (2009) considered several reports of F. magellanica F.Weber et Nees in Gottsche et al. (1845: 446) and its varieties doubtful, including the records of F. magellanica var. diminutiva by Herzog & Hosseus (1938). Engel (1978), who had not seen the type specimen of F. magellanica cited F. fertilis as its synonym. A detailed account of both species is presented by Hässel de Menéndez (1983). To decide on the status of F. magellanica subsp. tristaniana (S.W. Arnell 1958: 9) Váňa et J.J.Engel (2013: 59) further investigations are needed.

Frullania (subg. Frullania sect. Australes ) fugax (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 445, 1845 ( Gottsche et al. 1845). Basionym:— Jungermannia fugax Hook.f. et Taylor, London J. Bot. 4: 87, 1845 ( Hooker & Taylor 1845). Type:— NEW ZEALAND. On Rimelia (Parmelia) reticulata , 1844, J. D. Hooker (ex herb. Lehm., S!) (lectotype here designated [packet of lower right hand corner on sheet]).

= Frullania yorkiana Steph., Sp. Hepat. (Stephani) 4: 558, 1911 ( Stephani 1911), syn. nov. Type:— AUSTRALIA. Queensland: Cape York, August 1897, Micholitz, Herbarium F. Stephani no. 18236 (G-00066920, lectotype designated by Hattori 1979: 152). Note:— Hattori (1979) noted the similarity with F. fugax and stated that it possibly is a synonym.

Frullania (subg. Diastaloba) gracilis (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 452, 1845 ( Gottsche et al. 1845). Basionym:— Jungermannia gracilis Reinw., Blume et Nees , Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12(1): 221, 1824 [1825] ( Reinwardt et al. 1824). Type:— INDONESIA. Java: s. loc. spec. (type not located; missing in STR and other searched herbaria). Note:— Frullania gracilis belongs to a polymorphous and taxonomically difficult species complex (e.g., Verdoorn 1929, Hattori 1974a, 1974b, 1975c) and needs further attention. Particularly, the type of Jungermannia gracilis has never been examined.

= Frullania minor var. integribracteola S.Hatt., J. Hattori Bot. Lab. 39: 293, 1975 ( Hattori 1975a), syn. nov. Type:— INDONESIA. Sumatra, Mt. Singgalang, July 1955, Meijer 1723 (L, holotype; NICH, isotype).

= Frullania minor subsp. recurviloba S.Hatt., J. Hattori Bot. Lab. 39: 293, 1975 ( Hattori 1975a), syn. nov. Type:— INDONESIA. Sumatra, Mt. Korinchi, 29 July 1956, Meijer B-9055 (L, holotype; NICH isotype).

Frullania (subg. Frullania) jovetiana von Konrat et Hentschel , nom. nov. pro Frullania pseudericoides S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12: 132, 1986 ( Hattori 1986c), nom. illeg. (ICN Art. 53.1; hom. illeg. [non S.Hatt. 1982]). Type:— INDIA. Kurz 1409, Herbarium F. Stephani no. 15963 (G-00115327, holotype; NICH, isotype). Blocking name:— Frullania pseudericoides S.Hatt., J. Hattori Bot. Lab. 51: 256, 1982 (Hattori 1982). Note:—The type is stored in G under the herbarium name “ Frullania jovetiana ”. It is possible that the name in Hattori (1986c) is a mistake for what was intended to be this name.