Anchimacheta costaricae, Sohn, 2014

Sohn, Jae-Cheon, 2014, A taxonomic review of the Neotropical genus Anchimacheta (Lepidoptera: Urodidae) with descriptions of a new congener and an allied, new genus and species from Sri Lanka, Journal of Natural History 48 (43 - 44), pp. 2617-2631 : 2625-2626

Anchimacheta costaricae n. sp.

( Figures 3D and E , 4B and D , 5B )

Types

Holotype: 1 ♀, ‘Est. Cacao, 1000–1400 m | Lado SO Vol [can] Cacao, | P.N.G., Prov [ince] Guan[acaste] | COSTA RICA, C. Chaves, Abr [=April] 1991. | L-N- 323300,375700’, ‘Sohn | Genitalia slide ♀ | SJC – 537 | made on 17. Nov. 2007 ’, deposited in INBIO . Paratypes (1 ♂, 3 ♀): 1 ♀, same locality as holotype, 21–29 May 1992 (E Lopez), INBIO ; 1 ♀, ditto, Nov.–Dec. 1990 (C Chaves & R Epinoza), INBIO , [GSN] SJC-532; 1 ♀, Costa Rica, Prov. Guanacaste, lado oeste del Volcan Cacao , Est. Cacao , Derrumbe , May 1992 (Illcurso Parataxon.), INBIO ; 1 ♂, Costa Rica, Prov. Guanacaste, P. N. Amistad, Est. Las Mellizas , Fca. Cafrosa , 1300 m, April 1991 (M Ramires), INBIO , [GSN] SJC-824.

Diagnosis

Anchimacheta costaricae is similar to A. iodes in forewing coloration and pattern but differs from the latter in having the narrower and more yellowish forewings. However, such superficial differences can often be obscure, depending on condition of specimens. Examination of the genitalia is therefore required for reliable identification. The male genitalia of A. costaricae are distinguished from those of A. iodes ( Figure 4A a and b ) in having a narrower cucullus ( Figure 4B ). In the female genitalia ( Figure 5B b and c ), Anchimacheta costaricae possesses a shorter and trapezoidal antrum (quadrate in A. iodes ; Figure 5A a ).

Description

Head. Vertex pale brownish grey, intermixed with dark brownish grey scales in anterior half; frons pale brownish grey. Antennae 1/2 as long as forewing costa; scape pale brownish grey; flagellomere lustrous dark brown in basal half, pale brownish grey in distal half. Labial palpi 1st segment pale brownish grey; 2nd segment mostly dark brown, brownish grey in ventromedial 1/3, pale brownish grey in ventrobasal 1/3, on interior surface, and around distal end; 3rd segment dark brown on outer surface, narrowly pale brownish grey on inner surface, apical area pale brownish grey.

Thorax. Patagium, tegumen, mesonotum and legs same as A. iodes . Forewing length 4.9–6.6 mm (n = 5), yellowish brown, intermixed with pale reddish brown scales; basal area dark grey; sub-basal line arched, dark brown, broadened to costa; antemedial line slightly arched, dark brown, of even width, broadened in costal area,; postmedian area mottled with dark brownish grey; subterminal and terminal area suffused with dark brownish grey; a small, dark brown spot near to tornus; fringe yellowish grey. Hind wing brownish grey; fringe yellowish grey.

Abdomen. Tergites brownish grey; sternites brownish white. Male and female genitalia ( Figures 4B and D , 5B ) overall similar to iodes except having a digitate cucullus that is 1/2 as narrow as the maximum width of costal process in the male genitalia ( Figure 4B ); antrum broadened anteriorly, 1/2 as long as its maximum width ( Figure 5B b and c ); and corpus bursae oval in the female genitalia.

Distribution

Costa Rica (Guanacaste, Puntarenas) .

Etymology

The species is named after the country where the type specimens were collected.

Figures

Figure 3. Type specimens. (A) Lectotype of Anchimacheta iodes, male; (B) Lectotype of A. capnodes (= A. iodes), male; (C) holotype of A. tolmetes (= A. iodes), male; (D) paratype of A. costaricae, male; (E) holotype of A. costaricae, female; (F) holotype of Glaucotunica tamila, male.

Figure 4. Male genitalia. (A) Genital capsule of Anchimacheta capnodes (= A. iodes), paralectotype (insets: a, b – right valva; c – medial band of gnathos; d – juxta); (B) right valva of A. costaricae, holotype; (C) phallus of A. capnodes (= A. iodes), paralectotype; (D) phallus of A. costaricae, holotype; (E) genital capsule of Glaucotunica tamila, holotype (insets: e – gnathal arms; f – juxta); (F) phallus of G. tamila, holotype (inset: g – enlarged distal third, transmitted light phase contrast image).

Figure 5. Female genitalia. (A) Anchimacheta capnodes (= A. iodes), paralectotype (insets: a – antrum; d – signum); (B) A. costaricae, paratype (insets: b, c – antrum; e – signum).