Bibio giganteus Unger, 1841

Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, Zootaxa 4759 (3), pp. 351-378 : 364-367

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Bibio giganteus Unger, 1841


Bibio giganteus Unger, 1841  

( Figs. 12–14 View FIGURE 12 View FIGURE 13 View FIGURE 14 , 32–33 View FIGURES 32–35. 32 )

Bibio giganteus Unger, 1841: 427   .

Syn. Bibio partschi Heer, 1849: 216   . Syn. n.

Syn. Bibio maculatus Heer 1849: 219   . Syn.n.

Penthetria Partschi (Heer) Pongracz, 1928: 172   . Type material: Holotype (female) of Bibio giganteus   SLJG (77605). This is the specimen depicted by Heer (1849, Pl. XVI, Fig. 10 View FIGURE 10 ). Syntype of Bibio maculatus   SLJG 77474 View Materials   . Holotype (female) of Bibio partschi   NHMW   (2007z013010001).

Additional material: Female ( SLJG 235), wing ( SLJG 35), wing and fragment of the rest of the specimen ( SLJG 13). Heer mentioned one specimen and three wings. Female ( NHMW D 1184), labeled as Bibio obsoletus   . 3 females ( CNHM 1836, 1840, 1842). Female, probably non-type, NHMW 1852.1.1007 (labeled as Bibio partschi   ).


Female:Total length 12.5–17.7 mm (N=5).

Head: ( Fig. 33 View FIGURES 32–35. 32 ) Length 1.4 mm (N=1). Black. Poorly preserved in specimens at hand. Complex eye rather small, rounded. Ocellar tubercle small. Antennal flagellum apparently short and stout, 0.41 mm long by 0.19 mm wide, possibly five- or six-segmented.

Thorax: Length 3.9–4.6 mm (N=4), width 3.2–3.6 mm (N=3). Dark reddish brown. Halter reddish brown.

Wing ( Fig. 32 View FIGURES 32–35. 32 ): Length 10.0– 13.1 mm, width 3.7–4.4 mm (N=5). Brown fumose, costal cell conspicuously darkened, veins darkened, strong throughout. Pterostigma black, oval. Basal R s a little longer than R-M. Bm-Cu meets M 2 some distance beyond furcation of M. Veins M 1, M 2 and CuA 1 apically rather straight, CuA 2 a little curved. Vein measurements (all N=5): Subcosta 5.0– 6.8 mm, basal R 3.4–4.2 mm, distal R 1 2.6–3.3 mm, R s 1.0 – 1.6 mm, R 2–5 4.2–5.8 mm, R-M 0.4–0.6 mm., basal M 3.2–4.0 mm, distal M 0.8–1.4 mm, M 1 3.6–4.9 mm, M 2 3.2–3.9 mm, M-Cu 0.7–1.0 mm, CuA 2.0– 2.8 mm, CuA 1 5.0–6.0 mm, CuA 2 3.0–4.0 mm.

Legs: Dark brown, likely black in life. Protibia stout with a quite short spur. Length of fore tibia 2.8–2.9 mm (N=2), width 0.55–0.64 mm (N=2), length of hind tibia 4.2–4.7 mm (N=3).

Abdomen: Length 9.3–13.4 mm (N=5), width 3.3–4.3 mm (N=4). Tergites dark brown, pleura reddish brown. Cerci small and triangular.

Discussion: The nominal species Bibio giganteus   , Bibio maculatus   and Bibio partschi   may appear distinctive since B. giganteus   has conspicuously coloured wings, the others not. However, the morphometric traits of the wings are quite similar, and it seems most likely that the specimens classified as Bibio maculatus   and Bibio partschi   are Bibio giganteus   specimens without preserved wing pigments. The similarity of B. giganteus   and B. maculatus   was noted already by Heer (1849: 219), who distinguished the two on the basis of B. giganteus   having relatively longer abdomen (but the abdomen is notably swollen in the holotype of B. giganteus   ) and B. maculatus   having marginally narrower wings. Heer (1849: 216) distinguished Bibio partschi   principally by its wider abdomen, this trait is, however, almost certainly an artefact of post-mortem swelling due to partial decay of the specimen before it was covered by sediment. The senior author has also seen material of this species from the late Oligocene site of Rott, Germany (Skartveit & Wedmann in prep.)


Naturhistorisches Museum, Wien


Steiermarkisches Landesmuseum Joanneum


Cincinnati Museum of Natural History














Bibio giganteus Unger, 1841

Skartveit, John & Krizmanić, Katarina 2020

Penthetria Partschi (Heer) Pongracz, 1928: 172

Pongracz, A. 1928: 172