Rhaconotinus (Rhaconotinus) albosetosus Belokobylskij

Belokobylskij, Sergey A. & Achterberg, Cornelis Van, 2021, Review of the braconid parasitoid subfamily Doryctinae (Hymenoptera, Braconidae) from the United Arab Emirates and Yemen, European Journal of Taxonomy 765 (1), pp. 1-143: 97-101

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Rhaconotinus (Rhaconotinus) albosetosus Belokobylskij

sp. nov.

Rhaconotinus (Rhaconotinus) albosetosus Belokobylskij   sp. nov.


Figs 43–44 View Fig View Fig

Comparative diagnosis

This species is similar to Rh. micholitzi (Belokobylskij, 2001)   comb. nov. from Papua New Guinea ( Belokobylskij 2001b), but differs from latter in having the vertex entirely transversely striate (mainly granulate and only laterally with short striation in Rh. micholitzi   ), frons entirely coarsely reticulaterugose (mainly granulate in Rh. micholitzi   ), precoxal sulcus coarsely crenulate (smooth in Rh. micholitzi   ), pterostigma pale in basal third (entirely dark brown in Rh. micholitzi   ), mediocubital vein (M+CU1) of fore wing distinctly curved in apical half (weakly curved in Rh. micholitzi   ), nervulus (cu-a) distinctly postfurcal (almost interstitial in Rh. micholitzi   ), hind tarsus distinctly longer than hind tibia (almost equal to it in Rh. micholitzi   ), and the first metasomal tergite short (long in Rh. micholitzi   ).


The name of this species refers to the Latin words ‘ albus ’ (white) and ‘ setosis ’ (pubescence), after the white body setosity of this species.

Material examined

Holotype YEMEN • ♀; “Yemen (6117), Al Kowd, 21–28.viii.2001, light trap, A.v. Harten, S. Al Haruri, RMNH’02”; RMNH.  

Paratypes YEMEN • 1 ♀; “ Yemen, Al Kowd, vii.2000, light tr., no 4820, A.v. Harten & S. Al Haruri, RMNH’01”; RMNH   1 ♀; “ Yemen (4054), Al Kowd , viii.1999, light trap, A.v. Harten & S. Al Haruri, RMNH’00”; RMNH   .



MEASUREMENTS. Body length 5.5–6.9 mm; fore wing length 3.7–4.5 mm.

HEAD. Head width 1.3–1.5 times its median length, 1.2–1.3 times width of mesoscutum. Head behind eyes very convex anteriorly, than weakly roundly narrowed; transverse diameter of eye 1.2–1.3 times larger than temple length. Ocelli medium-sized, in triangle with base 1.2 times its sides; POL 1.0–1.4 times Od, 0.45–0.60 times OOL. Eye glabrous, weakly emarginated opposite antennal sockets, 1.2 times as high as broad. Malar space height 0.40–0.45 times height of eye, 0.75–0.80 times basal width of mandible. Malar suture fine but rather distinct, curved. Face width almost equal to height of eye and 1.2–1.3 times height of face and clypeus combined. Upper margin of clypeus situated almost on lower level of eyes. Hypoclypeal depression round, its width 0.75–0.80 times distance from edge of depression to eye, 0.40–0.45 times width of face. Occipital carina not fused with hypostomal carina below being obliterated for rather long distance upper base of mandible; below present several additional transverse carinae. Vertex convex.

ANTENNA. Antenna slender, weakly setiform, 37–42-segmented, about as long as body. Length of scape 1.7–1.9 times its maximum width. First flagellar segment 5.5–6.0 times longer than its apical width, 1.2 times longer than second segment. Penultimate segment 4.5–4.7 times longer than wide, 0.4 times as long as first segment, 0.7–0.8 times as long as apical segment; latter apically distinctly pointed.

MESOSOMA. Length 2.1–2.2 times its height. Pronotum anteriorly weakly convex (dorsal view), posteriorly strongly and widely convex dorsally (lateral view). Pronotal carina rather distinct, widely separated from posterior margin of pronotum, distances from carina to anterior and posterior margins of pronotum subequal. Mesoscutum weakly and roundly elevated above pronotum. Notauli deep, complete and crenulate. Mesoscutum without median longitudinal depression. Prescutellar depression rather short, deep, its posterior sides straight and obliquely directed posterolaterally, depression with distinct median carina, rather densely and distinctly crenulate, about 0.3 times as long as convex scutellum. Precoxal sulcus distinct, densely rugose-crenulate or crenulate, weakly S-shape, long, running along entire lower part of mesopleuron. Prepectal carina rather strong, not widened below, without lobes opposite fore coxae. Metanotum with distinct but not high subpointed tooth (lateral view). Metapleural lobe rather short, narrow and rounded distally.

WINGS. Length of fore wing 3.9–4.0 times its maximum width. Radial (marginal) cell not or only weakly shortened. Metacarpus (1-R1) 1.15–1.20 times longer than pterostigma, almost 7.0 times longer than distance from apex of radial (marginal) cell to apex of wing. Radial vein (r) arising weakly before middle of pterostigma, from basal 0.45. First radial abscissa (r) forming distinct obtuse angle with second abscissa (3-SR). Second radial abscissa (3-SR) 2.6–2.7 times longer than first abscissa (r), 0.5 times as long as third abscissa (SR1), 1.5 times longer than first radiomedial vein (2-SR). Second radiomedial (submarginal) cell not widened distally, its length 3.0–3.3 times maximum width, 0.85–1.00 times length of the wide brachial (subdiscal) cell. First medial abscissa (1-SR+M) distinctly sinuate. Recurrent vein (m-cu) weakly postfurcal. Mediocubital vein (M+CU1) rather distinctly curved to anal vein (1-1A) in distal half. Distance from nervulus (cu-a) to basal vein (1-M) 0.6–0.7 times nervulus (cu- a) length. Brachial (subdiscal) cell rather gently and almost roundly closed almost on level of recurrent vein (m-cu); posterior bulla on brachial vein (CU1b) present; posterior abscissa of anal vein (2-1A) (behind brachial vein (CU1b)) shortly present. Hind wing not narrowed basally, 5.4–5.5 times longer than wide. First costal abscissa (C+SC+R) 0.5–0.6 times as long as second abscissa (1-SC+R). First abscissa of mediocubital vein (M+CU) 0.50–0.55 times as long as second abscissa (1-M). Recurrent vein (m-cu) long and unsclerotised.

LEGS. Hind femur with distinct wide baso-dorsal protuberance, its length 2.7–2.9 times maximum width. Hind tarsus 1.1–1.2 times longer than hind tibia. Hind tibia distally with five–eight outside spines. Hind basitarsus 0.6 times combined length of second-fifth segments. Second tarsal segment 0.4–0.5 times as long as basitarsus, 2.6–3.0 times longer than fourth segment, 1.3–1.4 times longer than fifth segment (without pretarsus).

METASOMA. Metasoma 1.1–1.2 times longer than head and mesosoma combined, with six dorsally visible hard tergites. Posterior width of first tergite 1.8–2.1 times its anterior width; its length 1.05–1.20 times posterior width. Second tergite without anterior area, with distinct, narrow and weakly curved transverse submedian furrow separated lenticular posterior area; this area 0.85–0.90 times as long as remaining part of tergite. Median length of second tergite (with posterior area) 0.6–0.7 times its anterior width, 1.5–2.0 times length of third tergite. Second suture distinct and rather narrow. Sixth tergite large, rounded in posterior margin, with rather deep and more or less wide median emargination, without posteroventral lobes. Sixth tergite 1.5–2.0 times longer than fifth tergite, 1.7–2.3 times longer than fourth tergite. Ovipositor sheath almost as long as metasoma, 1.5 times longer than mesosoma, 0.8 times as long as fore wing.

SCULPTURE AND PUBESCENCE. Vertex distinctly and densely transversely striate, partly with rugulosity between striae; frons densely and coarsely rugose-reticulate; temple rugose-striate, reticulate-coriaceous anteriorly along eye. Face densely rugulose-reticulate, with curved striation in lower lateral parts. Mesoscutum densely to very densely and finely granulate with rugulosity partly, with distinct striation or rugosity along notauli, with two weakly convergent and undulate striae in medio-posterior third. Scutellum densely reticulate-coriaceous. Mesopleuron entirely densely reticulate-rugulose and partly with rugosity, smooth or almost smooth below precoxal sulcus. Propodeum with basolateral areas delineated by distinct carinae, with median carina in anterior 0.5–0.6, areola short or not delineated, if present then pentagonal, rugose; basolateral areas densely coriaceous-granulate to partly smooth entirely or at least in anterior half, rugose-reticulate in posterior half, usually distinctly densely striate along carinae. Hind legs densely and finely coriaceous, femur almost smooth in lower half. First tergite with complete dorsal carinae, entirely with several rather sparse longitudinal striae, densely rugulose-striate between striae. Second to fifth tergites distinctly and densely striate, with distinct and dense rugulosity or striation between striae; sixth tergite striate with very dense reticulation in anterior two–thirds, striae more or less concentrically in posterior third. Second-fifth tergites laterally with rather distinct striae and very dense rugosity-reticulation between it. Vertex with long and rather dense semi-erect white setae mainly directed backwards. Mesoscutum with dense semi-erect white setae widely arranged along notauli and anteriorly and in usually single line laterally. Mesopleuron almost entirely with sparse long and white setae. Metasoma dorsally almost entirely with long and rather dense semi-erect white setae directed backwards. Hind tibia dorsally with almost erect or semi-erect, rather dense white setae; length of these setae 1.0–1.3 times maximum width of hind tibia.

COLOUR. Head reddish brown or dark reddish brown, faintly paler around eyes; mesosoma and metasoma dark reddish brown to almost black partly. Scape reddish brown, pedicel and flagellum in basal two– thirds or half yellowish brown to light reddish brown, flagellum distinctly darkened in apical third or half. Palpi brown to dark brown, sixth maxillary segment light brown or at least paler. All legs reddish brown, fore coxae light reddish brown, all tibiae basally distinctly paler, all tarsi yellowish brown to almost yellow. Ovipositor sheath black. Fore wing faintly and almost evenly infuscate. Pterostigma black to dark brown, whitish yellow or white in basal third or quarter and apically.






Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]


National Museum of Natural History, Naturalis