Speocera trapezialis Li & Li, 2019

Tong, Yanfeng, Li, Fengyuan, Song, Yang, Chen, Haifeng & Li, Shuqiang, 2019, Thirty-two new species of the genus Speocera Berland, 1914 (Araneae: Ochyroceratidae) from China, Madagascar and Southeast Asia, Zoological Systematics 44 (1), pp. 1-75 : 56

Speocera trapezialis Li & Li sp. nov. ( Figs 43–44 , 56I )

Material examined. Holotype ♂ ( IZCAS), Vietnam, Quang Binh Province, Phong Nha-ke Bang National Park, Botanic Garden (17º33.150′N, 106º18.083′E), elev. 117 m, 14.V.2016, Leg. Q. Zhao & Z. Chen. Paratype 1♀ ( IZCAS), same data as holotype .

Etymology. The specific name is derived from ancient Greek and refers to the trapezoidal male palpal bulb; adjective.

Diagnosis. The males of this new species can be distinguished from all the other congeners by the trapezoidal palpal bulb ( Figs 43A–B ) and the females by the very long, tubular spermathecae ( Fig. 44A ).

Description. Male. Total length 0.99; carapace length 0.51; carapace width 0.43; abdomen length 0.53; abdomen width 0.36. Prosoma: carapace pyriform, narrowing gradually anteriorly; light yellow, suffused with purplish pattern ( Fig. 44C ); pars cephalica flat; fovea not visible. Sternum light yellow, suffused with purplish color, as long as wide. Endites light yellow suffused with purplish tinge; with serrula; labium light yellow suffused with purplish tinge. Clypeus nearly vertical; short. Chelicerae light yellow, laterally slightly curved; promargin with a series of six teeth reaching the lamina; retromargin with two very small denticles ( Fig. 56I ). Eyes: six contiguous eyes surrounded by black pigmentation; PME and ALE nearly subequal, PLE largest ( Fig. 44C ). Opisthosoma oval; purplish. Legs light yellow; leg formula 4123; leg lengths I 2.04 (0.59, 0.14, 0.61, 0.41, 0.28); II 1.88 (0.55, 0.16, 0.54, 0.37, 0.26); III 1.63 (0.48. 0.13, 0.43, 0.34, 0.25); IV 2.15 (0.63, 0.14, 0.59, 0.47, 0.32). Genitalia ( Fig. 43 ). Palpal femur long; palpal tibia as long as cymbium ( Figs 43A–B ). Cymbium conical, without prolateral extension. Bulb nearly trapezoidal. Embolus short, dark and basally without distinct associated sclerites; sperm duct initially wide, narrowing distally ( Fig. 43D ).

Female. Total length 1.19; carapace length 0.51; carapace width 0.42; abdomen length 0.50; abdomen width 0.37. Similar to male in color and general features ( Figs 44D–E ) but larger. Chelicerae unmodified. Leg lengths I 1.96 (0.55, 0.16, 0.58, 0.40, 0.28); II missing; III 1.57 (0.44. 0.14, 0.41, 0.34, 0.24); IV 2.11 (0.59, 0.15, 0.61, 0.46, 0.31). Genitalia. Internal genitalia with afferent spermathecae very short, strongly curved posteriorly from the epigastric furrow, not extending past the sides of the abdomen ( Fig. 44B ); spermathecae long, tubular ( Fig. 44A ).

Distribution. Known only from the type locality.

Figures

Figure 43. Speocera trapezialis Li & Li sp. nov., holotype, male. A. Left palp, prolateral view. B. Left palp, retrolateral view. C. Left palp, dorsal view. D. Left palp, retrolateral view, cleared in lactic acid. Scale bars=0.05mm.

Figure 44. Speocera trapezialis Li & Li sp. nov. A, B, D, E. Paratype, female. C. Holotype, male. A. Genitalia, dorsal view. B. Genitalia, ventral view. C. Habitus, dorsal view. D. Habitus, dorsal view. E. Habitus, ventral view. Scale bars: A–B=0.05mm; C– E=0.30mm.

Figure 56. Male chelicerae of Speocera spp. A–E, G–I. Anterior view. F. Lateral view. A. S. cattien Li & Li sp. nov. B. S. gexuejuni Li & Li sp. nov. C. S. heilan Li & Li sp. nov. D. S. lahrak Li & Li sp. nov. E–F. S. manhao Li & Li sp. nov. (arrow shows the small apophysis) G. S. longyan Li & Li sp. nov. H. S. nuichua Li & Li sp. nov. I. S. trapezialis Li & Li sp. nov. Scale bars=0.1mm.