Dichotomius (Cephagonus) asenjoi, Nunes & Vaz-de-Mello, 2019

Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), Journal of Natural History 53 (37), pp. 2231-2351 : 2313-2314

3.8.: Dichotomius (Cephagonus) asenjoi new species

( Figure 33 (e-h))

Diagnosis. separated from other in the group by the following combined characters: males pronotum with narrow emargination and declivity on anterior portion and having a central pair of knobs on pronotal disc; males cephalic carina inserted both eyesı transversal with acute apex (similar to D. aielloae ); elytral striae and puncture deeply impressed; males paramera apexes convergent (as D. marcoscarvalhoi ) and males paramera subgenital plate taking less than half of ventral surface.

● HOLOTYPE (male): BRASIL: Mato Grosso. Querência. Fazenda São Luiz. 12³41 ’ Sı 52³30 ʹ W. Pitfall . ii-2009. R Andrade [at CEMT ].

● PARATYPES same data of Holotype [5 ♀♀ ar CEMT]; Pará. Alter do Chão. RESEX Arapiuns/Tapajós. FIT. 22-xii-2008 [Ƌ at CEMT]; Juriti. 2³46 ’ Sı 55 ³40 ʹ W. xii-2008. FZ Vaz-de-Mello [1 ♀ at CEMT]; Rondônia: Porto Velho. Santo Antônio (Rio Madeira). Luz. Xii-2004 [1 ♀ at CEMT]. PERU: Madre de Dios. Jorge Chavez. La Viuda Camp (Rio Patuyacú). 12³52 ’ Sı 68 ³56 ʹ W.ı 400 m. Fezes humanas. iii-1999. T Larsen. [1 ♀ at CEMT]; same municipality but Limon Camp (Rio Palma Real Grande). 12³32 ’ Sı 68³51 ʹ Wı 220 m. FIT. x-1999. T Larsen [1 ♀ at CEMT]; idem at Oculto Camp (Rio Patayaçú). 12³39 ʹ 00 ʹ ’ Sı 68³55 ʹ 33 ʹ ’ Wı 400 m. FIT. ii-1999. T Larsen [1 Ƌ at CEMT]; idem at Rio Tambopata. 12³38 ʹ 59 ʹ ’ Sı 69³06 ʹ 24 ” Wı 230 m. FIT. x-1999. T Larsen [1 Ƌı 1 ♀ at CEMT].

Description (Holotypeı male). BL: 17 mmı PW: 8 mm. colour: blackı shiny. Head: frontoclypealı genal and inter-ocular surfaces with fine punctures. clypeo-genal angle roundedı sub-straight (100³). Cephalic carina triangular in frontal view and inserted between both eyes. Pronotum: pronotal disc shinyı smoothı lacking puncturesı with weakly small central emargination sided by two weak knobs. Row of ocellate punctures along the posterior marginı punctures getting larger and elliptical towards posterior angles. Metasternum: sides covered by ocellate setigerous punctures equally spacedı setae with half-length of mesepimeron setae; anterior lobe glabrous only bearing ocellate and coarse puncturesı eventually an isolated short setae may appear along margin. Anterior portion of anterior lobe with ocellate setigerous punctures. Elytra: striae deeply impressed having coarse punctures spaced by twice their diameter. Interstriae convexı disc lacking microsculpture with very fine punctures (20 x). Abdomen: sides of ventrites with ocellate punctures near anterior margin and fine punctures on the disc. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite. Aedeagus ( Figure 33 (f-h)): dorsallyı paramere apexes roundedı strongly flattenedı convergent; longitudinal excavation absentı basal elliptical excavation present. Laterallyı abrupt declivity from basis towards apex; lacking excavation; Ventrallyı sclerotised portion of subgenital plate taking less than half of ventral paramera surface; basis and apex acute.

Morphological variation. males BL: 15 to 17 mmı PW: 8 – 10 mm. Females BL: 14 – 15 mmı PW: 8 – 9 mm. Some males may have opaque surface. Female is distinguished from males as follows: Head: fronto-clypeal surface striated. Cephalic horn absent. Pronotum: simply convexı lacking lobes. Elytra: striae weakly impressedı punctures separated by their diameter. Blue sheen present near basis. Abdomen: sixth ventrite with a central protuberance but not advancing beneath pygidium apex.

Distribution. Brazil (MTı PA and RO states) and Peru (lower areas of Madre de Dios) ( Figure 34 ).

Etymology. named after Dr. Angelico Fortunato Asenjo Flores (Universidade Federal de Mato Grossoı Brasil)ı peruvian entomologist and who helped with many concepts developed in this and other thesis works.

Figures

Figure 33. D. arnaudi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view (grey arrow indicates the position of longitudinal excavation on lateral view). D. asenjoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. barbarae (i) male dorsal habitus; (j) paramera dorsal view (black arrow indicating slight medial constriction); (k) paramera ventral view; (l) paramera lateral view (black arrow indicates the surface of longitudinal excavation, which is smooth with fine punctures (see D. paschoali) and the differentiation of both species).

Figure 34. Distribution map of D. gardneriı D. aielloaeı D. asenjoiı D. arnaudiı D. Clypeatus and D. edmondsi.