Dichotomius (Cephagonus) clypeatus ( Luederwaldt 1929 )

Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), Journal of Natural History 53 (37), pp. 2231-2351 : 2310-2311

3.5.: Dichotomius (Cephagonus) clypeatus ( Luederwaldt 1929) ( Figure 32 (a-e))

Pinotus (Cephagonus) bicuspis var. clypeatus Luederwaldt 1929 ı p. 129 (description) Pinotus (Cephagonus) clypeatus ı Luederwaldt 1936 ı p. 214

Diagnosis. this species is separated from other in the group by the following combined characters: males pronotal disc with a central pair of tubercles on anterior portion; elytral striae and punctures strongly impressed (as in D. ingens and D. arnaudi ); males cephalic horn or tubercle rounded apically (as D. bicuspis ); genae borders divergentı posterior angle rounded; paramera lacking basal nor longitudinal excavations and sides of elytra subparallel in dorsal view (as in D. arnaudi ).

● HOLOTYPE (male) ( Figure 32 (e)): Oxapampa Peru {whiteı printed}/ Pinotus quadraticeps Felsche {whiteı handwritten}/Muséum Paris 1936 Coll. A. Boucomont {white with black marginsı printed}/ Pinotus bicuspis Har. var. clypeatus . Clypeatus nov. var. Luederwaldt 1931 {whiteı handwritten} [MNHN].

Non-type material. ARGENTINA: Salta: Sierra Tartagal. 35 km N Tartagal. 22³19 ʹ 36 ʹ ’ Sı 63³51 ʹ 00 ʹ ’ Wı 874 m. 14.xi.2006. Berger-Dostal [1 Ƌ at CEMT]; BOLIVIA: Santa Cruz: Florida.

18³08 ’ Sı 63³48 ʹ Wı 1730m. 19-xi-2006. WD Edmonds & P Reyes [1 Ƌı 1 ♀ at CEMT]; Tarija. Entre Riosı Quebrada del Abra. 25-ii-1998. Ayzama & F Guerra [1 Ƌı 1 ♀ at CEMT].

Redescription. colour: blackı shiny. Head: fronto-clypealı genal and inter-ocular surfaces smooth. Gena borders not parallel. Cephalic carina conical with rounded apex. Pronotum: borders smoothı not serrated. Pronotal disc shinyı lacking punctures and delimited anteriorly by two central weak tubercles. Metasternum: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe glabrous. Setigerous punctures evenly spaced at sides. Elytra: sub-parallel. Striae deeply impressed having fine weakly impressed punctures spaced by one to three times their diameter. Interstriae convexı lacking microsculpture with very fine puncture (20 x). Abdomen: sides of each ventrite excavated. 1st ventrite with a single row of hair along anterior margin. Each side of ventrites 2 to 5 with groups of 1 – 5 ocelatte setigerous punctures. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite. Aedeagus ( Figure 32 (b-d)). Dorsallyı paramere apexes roundedı convergent; longitudinal and basal excavations lacking; membranous portion taking more than 2/3 of dorsal surface. Laterallyı subtle declivity from basis towards apex. Ventrallyı subgenital plate widely emarginate at apex and rounded at basis.

Morphological variation. Male BL: 13 – 15 mmı PW: 7 – 9 mm. Female BL: 16 mmı PW: 6 mm and distinguished from males as follows: Head: dorsal surface striated. Clypeo-genal junction very obtuseı rounded. Cephalic carina absent forming at most a central weak knob. Pronotum: simply convex. Elytra: interstriae microsculpture chagrinated. Abdomen: sixth ventrite three times larger than fifthı pygidium swollenı lacking modifications at medial portion.

Distribution. Argentina (Salta)ı Bolivia (Santa Cruzı Tarija) and Peru (Oxapampaı type locality)

Figures

Figure 32. D. clypeatus: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) type labels. D. aielloae: (f) Holotype dorsal habitus; (g) detail of male cephalic carina in frontal view; (h) type labels.