Dichotomius (Cephagonus) furtadoi, Nunes & Vaz-de-Mello, 2019

Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), Journal of Natural History 53 (37), pp. 2231-2351 : 2321-2322

3.15.: Dichotomius (Cephagonus) furtadoi new species

( Figure 38 (e-i))

Diagnosis. separated from other species in the quadraticeps group by the following combined characters: larger males pronotum with two central-rounded lobes on the anterior portion of the disc; posterior margin of pronotum having 3 – 4 rows of ocellate puncture anterior portion of metasternum anterior lobe with a row of setigerous punctures; ventral hair longı concealing some ventral ventrites and paramera basal excavation having chagrinated microsculpture.

● HOLOTYPE (male): BRASIL: Mato Grosso. Diamantino. Vale da Solidão. 14³22 ’ Sı 56³07 ” Wı 450 m. xi-2012. E Furtado [at CEMT].

● PARATYPES: same data of Holotype [1 Ƌı 1 ♀ at CEMT]; Diamantino. Vale da Solidão. 14³21 ʹ 52 ” Sı 56³16 ʹ 14 ” Wı 450 m. Cerrado rochoso. DCT Oliveira [2 ƋƋı 3 ♀♀ at CEMT]; same municipality at 14³17 ʹ 05 ” Sı 56 ³16 ʹ 14 ” W. Liver. xi-2013. ME Maldaner [1 ♀ at CEMT]; same municipality but at Alto Rio Arinos. I-2001. E Furtado [1 ♀ at CEMT].

Description (Holotypeı male). BL: 20 mmı PW: 11 mm. Colour: blackı shiny. Head: frontoclypeal and genal surfaces smooth. Inter-ocular space with deeply impressed coarse punctures. Gena borders slightly curved inward. Cephalic carina longer than wideı conical with rounded apex. Pronotum: disc shinyı lacking punctures and having two central lobes on the anterior portion. Region adjacent to eyes with coarse deeply impressed punctures. Posterior margin having 3 – 4 rows of ocellate puncturesı punctures larger near basis. Metasternum: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe having a row of ocellate setigerous punctures. Setigerous punctures equally spaced at sides. Elytra: striae deeply impressedı subtle widened towards basis and having coarse punctures spaced by 3-four times their diameter. First interstriae (near elytral suture) convex in relation to other interstriae which are almost flat. Interstriae disc with dark brown colourı lacking microsculpture with very fine puncture (20 x). Abdomen: sides of each ventrite covered by ocellate punctures and excavated near anterior margin. Ventrites having 1 – 2 setae at each side. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite. Aedeagus ( Figure 38 (g-i)): dorsallyı paramere apexes acuteı parallel; longitudinal excavation present taking the whole paramera length; basal excavation having chagrinated microsculpture. Laterallyı abrupt declivity from basis towards apex; with very developed anterior knob. Ventrallyı membranous portion not reaching basis. Each paramera having a transversal groove; subgenital plate membranous apically and emarginated on basis.

Morphological variation. males BL: 17 – 20 mmı PW: 9 – 11 mm. Larger males have developed cephalic horn (rounded at apex) and pronotum central lobes. Medium and smaller males have the cephalic horn produced in a transversal carina and do not have pronotal lobes. Both types of males and females have similar puncture and hair distribution patternı which is also useful to distinguish this species. Females BL: 17 – 20 mmı PW: 9 – 12 mmı distinguished from males as follows: Head: fronto-clypeal surface striated. Coarse puncture present posterior to cephalic carina near both eyes. Cephalic carina transversal. Pronotum: simply convexı lacking lobes. Elytra: interstriae with chagrinated microsculpture. Abdomen: sixth ventrite three times larger than fifth and bearing a central pair of tubercles.

Distribution. municipality of Diamantinoı state of Mato Grossoı Brazil ( Figure 36 ).

Etymology. named after the entomologist Eurides Furtadoı from Diamantinoı Mato Grossoı Brasil.

Figures

Figure 36. Distribution map of D. ricardosilvaiı D. marcoscarvalhoiı D. furtadoiı D. barbaraeı D. tissianiaeı D. eması D. frizzasae and D. candangoi.

Figure 38. D. feeri: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. furtadoi: (e) larger male; (f) smaller male; (g) paramera lateral view (black arrow indicating the gibbosity on the anterior portion of the paramera); (h) paramera dorsal view; (i) paramera dorsal.