Coryphomodes budda, Yin, 2022

Yin, Zi-Wei, 2022, The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae), Zootaxa 5111 (1), pp. 1-211 : 100-102

publication ID	https://doi.org/ 10.11646/zootaxa.5111.1.1
publication LSID	lsid:zoobank.org:pub:836B0F69-037C-4D0F-80DB-94FE454F48E3
DOI	https://doi.org/10.5281/zenodo.6964321
persistent identifier	https://treatment.plazi.org/id/C822F213-FFA8-2412-CB88-A57C773225B2
treatment provided by	Plazi
scientific name	Coryphomodes budda
status	sp. nov.

Treatment

Coryphomodes budda View in CoL sp. nov.

( Figs 43 View FIGURE 43 , 81C View FIGURE 81 , 99F View FIGURE 99 , 100A View FIGURE 100 )

Chinese common name: ṞĽỀȁà甲

Type material ( 2 exx.). HOLOTYPE: CHINA: ♂, ‘China: Tibet (Mêdog County), pass nr. Renqingbeng Temple (29°18’16”N, 95°21’13”E), alt. 1700 m; 17.viii.2016, Zhi-Lin Chen leg. [ƱẪẸṘ仁ů崩寺]’ ( SNUC). GoogleMaps PARATYPE: CHINA: 1 ♂, ‘China: Mêdog County , 80K (ca. 29°39’22”N, 95°29’20”E), 2110 m, 11.viii.2016, Z.-L. Chen leg. [ƱẪẸṘ县 80K]’ ( SNUC). GoogleMaps

Diagnosis. Male. Body length 1.75 mm. Head rounded triangular, as broad as pronotum, tempus short, vertex and frons modified, vertex with lateral impressed areas, frons with median tubercle; antenna lacking modifications. Pronotum with laterally carinate median sulcus and lateral longitudinal sulci, with large antebasal spines and small marginal spines. Elytron with inner two basal foveae close, discal stria shallow and short, extending posteriorly to basal 1/4 of elytral length; disc with sparse, large punctures at posterior 3/4. Fore and middle legs simple, metatrochanter with large ventral projection. Aedeagus strongly asymmetric, stout; ventral stalk protruding apically, dorsal lobe plate-like, parameres reduced.

Description. Male. Body ( Fig. 43A View FIGURE 43 ) length 1.75 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.

Head ( Fig. 43B View FIGURE 43 ) rounded triangular, wider than long, length 0.34–0.35 mm, width across eyes 0.44 mm; vertex smooth, lacking foveae, mediobasal area strongly concave, laterally with round impressed areas, anteriorly preceded by short, transverse ridge; frons anteriorly confluent with clypeus, with antero-lateral ridges, with strongly raised tubercle at middle, sides of tubercle with short setae, posterior margin with tuft of thickened setae; clypeus with smooth surface, its entire anterior margin carinate and raised; ocular-mandibular carina complete, distinct. Venter with granulate surface, mediobasal area broadly impressed; small gular foveae (posterior tentorial pits) originating from shared transverse opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 ommatidia. Antenna lacking modification, length 0.78 mm, distinct club formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each moniliform, 2 as long as wide, 3–7 short, each more or less transverse, 8 smallest, 9 much larger than 8, transverse, 10 broader but shorter than 9, 11 largest, longer than 9 and 10 combined, subconical.

Pronotum ( Fig. 43B View FIGURE 43 ) wider than long, length 0.39–0.40 mm, width 0.44 mm, widest posterior middle; lateral margins rounded at apical 1/2, basally subparallel; disc slightly convex, with sparse but distinct punctures, median longitudinal sulcus with carinate sides, with pair of shallow lateral longitudinal sulci, antebasal tubercles large, lacking discal spines; with small marginal spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thin hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.

Elytra wider than long, length 0.52–0.54 mm, width 0.67–0.68 mm; each elytron ( Fig. 43C View FIGURE 43 ) with three large, asetose basal foveae, inner two close; discal stria shallow and short, extending posteriorly from outer basal fovea to basal 1/4 of elytral length; humerus weakly and broadly prominent, lacking subhumeral fovea, carinate marginal stria extending from basal 2/5 to posterior margin of elytron.

Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated in shared oval opening, large lateral mesoventral foveae forked internally, with short mesoventral process. Metaventrite prominent admesally, moderately impressed at middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Marginal carina complete, extending from anterior margin of mesoventrite to posterior margin of metaventrite.

Legs more or less robust; trochanters each with exceptionally long seta on ventral margin. Mesotrochanter with tiny tubercle on ventral margin; metatrochanter ( Fig. 43D View FIGURE 43 ) with large ventral projection curved at apex.

Abdomen widest at lateral margins of tergite 1 (IV), length 0.46–0.50 mm, width 0.62–0.64 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking mediobasal foveae or discal carinae, basal sulcus thin, with large basolateral foveae at lateral margins of sulcus, inner marginal carina slightly oblique, complete, outer one thicker, complete; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of small mediobasal and two pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) ( Fig. 43E View FIGURE 43 ) membranous, with weakly sclerotized apex.

Aedeagus ( Fig. 43F, G View FIGURE 43 ) 0.24 mm long, strongly asymmetric, stout; median lobe with broad basal capsule and slightly oval foramen, basoventral projection short, ventral stalk with one long, thickened seta at middle, protruding and subtruncately rounded at apex; non-articulated dorsal plate short and flattened, roundly rectangular; parameres reduced.

Female. Unknown.

Comparative notes. The new species is morphologically similar to C. cephalicus and C. chenzhilini , and can be readily separated from both species by the smaller body size, a stouter habitus, and the characteristic cephalic modifications and form of the aedeagus.

Distribution. Mêdog County, Tibet, SW China ( Figs 81C View FIGURE 81 , 99F View FIGURE 99 , 100A View FIGURE 100 ).

Etymology. The species name refers to the tantric Buddhist Guru Rimpoche (Äǩffik士), who is enshrined and worshiped at Renqingbeng Temple.