Pupinidae Pfeiffer, 1853

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Family Pupinidae Pfeiffer, 1853

Remarks.

Currently, there are three subfamilies within the family Pupinidae : Pupininae , Liareinae Powell, 1946, and Pupinellinae Kobelt, 1902 ( Bouchet et al. 2017). The subfamily Liareinae was endemic to New Zealand, originally established as a family ( Powell 1946), and this familial assignment was adopted by Egorov (2013). Later, Ponder and Warén (1988) treated this taxon as a subfamily of the Pupinidae ; this classification scheme was adopted by Bouchet et al. (2017) and MolluscaBase (2022).

The subfamily Pupinellinae was originally established as a section under the Pupinidae , and the only diagnostic character that distinguished this subfamily from the Pupininae is the shell surface ( Kobelt 1902). The Pupininae has a shell surface covered by glaze, which is glossy and completely smooth, whereas the shell surface of the Pupinellinae is without glaze, being either striated, matt or silky-shiny ( Kobelt 1902; Egorov 2013). Whether this character is a subfamilial synapomorphy needs further confirmation because at least one Pupina species has a matt surface (e.g., P. arula ) and some Pupinella species have a somewhat glossy surface (e.g., P. mansuyi , P. illustris ).