Catapaguroides, A. MILNE-EDWARDS & BOUVIER, 1892

Komai, Tomoyuki & Rahayu, Dwi Listyo, 2013, The Hermit Crab Genus Catapaguroides A. Milne-Edwards & Bouvier, 1892 (Crustacea: Decapoda: Anomura: Paguridae) From The Bohol Sea, Philippines, With Descriptions Of Eight New Species, Raffles Bulletin of Zoology 61 (1), pp. 143-188 : 186-187

KEY TO SPECIES OF CATAPAGUROIDES

1. Palm of right cheliped with prominent dorsal protuberance at articulation with dactylus....................... C. tuber , new species

– Palm of right cheliped without prominent dorsal protuberance at articulation with dactylus....................................................2

2. Ocular peduncles with corneas reduced, conical in shape.....3

– Ocular peduncles with corneas not appreciably reduced, nor conical in shape, rounded.......................................................6

3. Right cheliped entirely unarmed............................................... ..................... C. microps A. Milne-Edwards & Bouvier, 1892

– Right cheliped armed with spines on chela and carpus.........4

4. Cornea pointed distally; left chela unarmed on dorsal surface.................................................................................................5

– Cornea not distinctly pointed distally; left chela armed with few spines on dorsomesial margin ........................................... ................................ C. hooveri McLaughlin & Pittman, 2002

5. Antennal peduncle and acicle distinctly overreaching distal corneal margin; right male sexual tube relatively short, directed to left across ventral body surface, at most slightly overreaching lateral margin of left coxa .......... C. foresti McLaughlin, 2002

– Antennal peduncle and acicle not overreaching distal corneal margin; right male sexual tube very long, directed to left across ventral body surface and then strongly curved anteriorly, extending to base of left cheliped...... C. conicus , new species

6. Antennal peduncles overreaching distal corneal margins......7

– Antennal peduncles not overreaching distal corneal margins.................................................................................................17

7. Carpus and/or chela of right cheliped with at least 1 or 2 distinct spines..........................................................................8

– Carpus and chela of right cheliped lacking distinct spines ..... ............................................................. C. iris ( Bouvier, 1922)

8. Palm of right cheliped armed with row of 4 or more spines, at least on dorsomesial margin...............................................9

– Palm of right cheliped unarmed or with only 1 or 2 small tubercles or spinules, on or near dorsomesial margin or dorsomesial distal angle........................................................14

9. Spines on palm of right cheliped almost exclusively restricted to dorsomesial margin...........................................................10

– Spines on palm of right cheliped scattered or forming irregular rows on dorsal surface..........................................................13

10. Carpus of right cheliped distinctly shorter than palm ............. ............................................. C. melini de Saint Laurent, 1968

– Carpus of right cheliped as long as or distinctly longer than palm ....................................................................................... 11

11. Carpus of left cheliped with 2 or more spines.....................12

– Carpus of left cheliped unarmed............................................... ................... C. megalops A. Milne-Edwards & Bouvier, 1892

12. Dactylus of right cheliped with dorsomedian ridge or crest, at least in proximal half ............... C. declivis McLaughlin, 1997

– Dactylus of right cheliped without dorsomedian ridge or crest....................................... C. pectinipes ( Lewinsohn, 1969)

13. Palm of right cheliped with numerous small spines at least on distal half of dorsal surface....................................................... ........................................ C. japonicus de Saint Laurent, 1968

– Palm of right cheliped with 3 irregular longitudinal rows of fine spines or spinules on dorsal surface.................................. .................................. C. spinulimanus de Saint Laurent, 1968

14. Antennular peduncles overreaching distal corneal margins by nearly half length of penultimate segment...........................15

– Antennular peduncles overreaching distal corneal margins by entire length of ultimate segment.........................................16

15. Antennal acicle not reaching midlength of fifth segment of antennal peduncle; carpus of right cheliped unarmed at dorsomesial distal angle .... C. inermis de Saint Laurent, 1968

– Antennal acicle elongate, reaching distal margin of fifth segment of antennal peduncle; carpus of right cheliped with prominent spine at dorsomesial distal angle ............................ ...................... C. longior Komai, Yamada & Shirakawa, 2010

16. Ocular peduncle widened proximally with non-dilated cornea; carpus of right cheliped with spines on lateral and mesial angles of dorsodistal margin ..... C. brevidactylus , new species

– Ocular peduncle widened distally with dilated cornea; carpus of right cheliped unarmed on dorsodistal margin .................... ..................................... C. mortenseni de Saint Laurent, 1968

17. Carpus of left cheliped with 1 or more spines.....................18

– Carpus of left cheliped unarmed............................................... ...................................................... C. olfaciens ( Alcock, 1905)

18. Right chela operculiform, dactylus with sharply carinate mesial margin, palm with sharply carinate lateral margin..............19

– Right chela non-operculiform, lateral margin of palm rounded or at most slightly carinate...................................................20

19. Right palm with proximal spines or tubercles on dorsal surface; carpus of right cheliped only with longitudinally aligned 2 spines on dorsal surface; carpus of left cheliped unarmed on dorsolateral margin (except for dorsolateral distal spine)........ .................................................. C. karubar McLaughlin, 1997

– Right palm without proximal spines or tubercles on dorsal surface; carpus of right cheliped with longitudinal row of 5 or 6 spines on dorsal surface; carpus of left cheliped with 2 or 3 spines on dorsolateral margin (except for dorsolateral distal spin e) .......................... C. cristimanus de Saint Laurent, 1968

20. Palm of left chela entirely unarmed.....................................21

– Palm of left chela armed with at least 1 dorsomesial spine...................................................................................................26

21. Palm of right cheliped with dorsomesial spines..................22

– Palm of right cheliped without dorsomesial spines.............23

22. Cornea slightly dilated; dactylus of right cheliped usually with prominent spine on mesial surface proximally; carpus of right cheliped with lateral spine on dorsodistal margin; dactyli of ambulatory legs each with 4–7 corneous spines on ventral margin ........................... C. iejimensis Osawa & Takeda, 2004

– Cornea not dilated; dactylus of right cheliped always unarmed; carpus of right cheliped unarmed on dorsodistal margin; dactyli of ambulatory legs each with 1–3 corneous spines on ventral margin .................................................. C. umbra Komai, 2009

23. Merus of left cheliped with subdistal spine on dorsal margin; merus of left third pereopod with 1 spine on dorsal margin proximal to midlength and 2 spines on ventral margin in distal 0.3 ....................................................... C. pusillus , new species

– Merus of left cheliped unarmed on dorsal margin; merus of left third pereopod unarmed or armed only with ventrolateral distal spine.............................................................................24

24. Palm of right cheliped with prominent proximal protuberance on dorsal surface; ambulatory dactyli terminating in very long corneous claws ............................. C. tenuiclavus , new species

– Palm of right cheliped without protuberance on dorsal surface; ambulatory dactyli terminating in moderately long corneous claws......................................................................................25

25. Right cheliped elongate in male, with carpus subcylindrical and more than 3 times longer than distal width; dactyli of ambulatory legs unarmed or armed with only 1 corneous spine on ventral margins............................. C. hirsutus , new species

– Right cheliped not distinctly elongate in male, with carpus distinctly widened distally and less than 2 times longer than distal width; dactyli of ambulatory legs armed with 5–7 corneous spines on ventral margins.......................................... .......................................................... C. levigatus , new species

26. Palm of right cheliped with dorsomesial row of spines, but without scattered tubercles on dorsal surface.......................27

– Palm of right cheliped without dorsomesial row of spines, but with scattered tiny tubercles on mesial half of dorsal surface .................................................... C. crassimanus , new species

27. Mesial margin of dactylus of right cheliped with thin dorsomesial carina. Dactylus of left cheliped 1.2 times as long as palm, fingers arched ventrally .............................................. ................................................. C. setosus ( Edmondson, 1951)

– Mesial margin of dactylus of right cheliped rounded dorsomesially. Dactylus of left cheliped appreciably shorter than palm, fingers not arched ventrally .................................... .......................................................... C. fragilis ( Melin, 1939)