Achilia bifossifrons (Reitter, 1883)

Giorgio Sabella, Sergey A. Kurbatov & Giulio Cuccodoro, 2017, A revision of the Chilean Brachyglutini - Part 2. Revision of Achilia Reitter, 1890: A. crassicornis, A. tumidifrons, A. bifossifrons, and A. lobifera species groups (Coleoptera: Staphylinidae: Pselaphinae), Revue suisse de Zoologie 124 (1), pp. 119-140 : 137

Achilia bifossifrons (Reitter, 1883)

Additional material (1907 ex.): MNHN (coll. Raffray); 3 ♂; Chili. CENTRAL CHILI: Región Aysén:Aysén prov.: MHNG; 1 ♂; 30 km N Puyuhuapi, station 107; 100 m; 29.I.1985; S. & J. Peck. – Región Los Lagos: Chiloé prov.: MNHN; 1 ♀ (sub A. lobifera ); Chepu; 42° 03’S; 15.X.1958; G. Kuschel; forest. – MSNG; 2 ♂; Chiloé Island, Chepu; TC-275; 19.II.1991; T. Cekalovic. – MSNG; 3 ♂ and 6 ♀; same locality; TC-580; 09.II.1999, T. Cekalovic. – MSNG; 1 ♂ and 2 ♀; same locality; TC-610; 20.I.2000; T. Cekalovic. – MSNG; 27 ♂ and 13 ♀; same locality; TC-624; 26.I.2000; T. Cekalovic. – MSNG; 29 ♂ and 30 ♀; same locality; TC-625; 26.I.2000; T. Cekalovic. – FMNH (FMHD #97-22); 8 ♂ and 6 ♀; Chiloé Island, SE edge of Tepuhueico; 42°48.11’S 73°55.36’S; 50 m; 15.I.1997; valdivian rainforest; A. Newton & M. Thayer 992; berlese, leaf & log litter. – MSNG; 1 ♀; Mocopulli, SyTC-223; 18.II.1989; S. & T. Cekalovic. – MHNG; 3 ♂ and 1 ♀; Chiloé Island, Huillinco Lake; 31.I.1983; T. Cekalovic. – DBUC; 1 ♂; same data; T. Cekalovic. – MSNG; 3 ♂ and 2 ♀; same locality; TC-278; 21.II.1991; T. Cekalovic. – MSNG; 5 ♂ and 7 ♀; Chiloé Island, 1 km W of Huillinco Lake; TC-564; 24.I.1998: T. Cekalovic. – FMNH (FMHD #2002-72); 1 ♂; S side of Huillinco lake, road to Bellavista; 1.3 km S road of Cucao; 42°41.81’S 73° 55.88’W; 45 m; 12-22.XII.2002; A. Newton & M. Thayer 1062; valdivian rainforest w/emergent Saxegothea conspicua , flight intercept trap. – FMNH (FMHD #2002-74); 1 ♂; same locality; 12.XII.2002; A. Newton & M.Thayer 1062; valdivian rainforest w/emergent Saxegothea conspicua , berlese, leaf & log litter. – FMNH (FMHD #2002-092); 4 ♂ and 13 same locality; 22.XII.2002; A. Solodovnikov 1062; valdivian rainforest w/emergent Saxegothea conspicua , berlese, leaf & log litter. – FMNH (FMHD #2002-93); same data; A. Solodovnikov 1062; valdivian rainforest w/emergent Saxegothea conspicua , berlese, wet debris at stream. – PHPC; 1 ♂; Chiloé Island, Cucao, WDS-T-209; 42° 35’S 74° 05’W; 02.III.2008; W. D. Shepard; litter sifting. – FMNH (FMHD #97-25); 1 ♂; Miraflores, road to (0.6 km W Hwy 5); 42° 46.73’S 73° 47.71’W; 130 m; 17.I.1997; A. Newton & M. Thayer 994; secondary Valdivian rainforest, berlese, leaf & log litter. – FMNH (FMHD #2002-77); 5 ♂ and 3 ♀; same locality; 12.XII.2002; A. Newton & M. Thayer 1063; secondary valdivian rainforest with few conifers, berlese, leaf & log litter. – FMNH (FMHD #97-24); 27 ♂ and 32 ♀; road to Colonia Yungay (3.6 km W Hwy 5); 42° 59’S 73° 41’W; 90 m; 17.I.1997; A. Newton & M. Thayer 995; grazed secondary valdivian rainforest remnants, berlese, leaf & log litter. – FMNH (FMHD #2002-78); 9 ♂ and 23 ♀; road to Colonia Yungay, ca 4 km NW Hwy 5; 42° 59.12’S 73° 42.02’W; 110-115 m; 13.XII.2002; A. Solodovnikov & M. Thayer 1064; disturbed valdivian rainforest w/ recent selective cutting, berlese, leaf & log litter. – MSNG; 3 ♂ and 6 ♀; Chiloé Island, Puente La Caldera; TC-466; 15.II.1996, T. Cekalovic. – MSNG; 2 ♂; same locality; TC-524; 18.II.1997; T. Cekalovic. – MSNG; 3 ♂ and 12 ♀; Chiloé Island, 1 km N of Puente Notuco; TC- 528; 20.II.1997; T. Cekalovic. – MSNG; 3 ♂ and 11 ♀; Chiloé Island, 5 km SW Chonchi; TC-560; 21.I.1998; T. Cekalovic. – MSNG; 1 ♂ and 2 ♀; same locality; 14.I.1999; T. Cekalovic. – MSNG; 3 ♂ and 11 ♀; same locality; TC-623; 25.I.2000; T. Cekalovic. – MSNG; 1 ♂ and 1 ♀; Chiloé Island, Estero Llicaldad; TC-608; 19.I.2000; T. Cekalovic. – MSNG; 3 ♂ and 3 ♀; Chiloé Island, Puente Milildeo; TC-471; 15. II.1995; T. Cekalovic. – MSNG; 5 ♀; Chiloé Island, Estero Tablin; TC- 609; 19.I.2000, T. Cekalovic. – MSNG; 4 ♂ and 1 ♀; Quinchao Island, Quetro; TC-582; 12.II.1999, T. Cekalovic. – MSNG; 1 ♂; same locality; TC-559; 20.I.1998; T. Cekalovic. – MSNG; 1 ♂ and 3 ♀; Quinchao Island, Laguna Pulul; C-615; 22.I.2000; T. Cekalovic. – Llanquihue prov.: MNHN; 1 ♂ and 1 ♀; Los Riscos; 11.IV.1954; G. Kuschel. – FMNH (FMHD #57-125); 1 ♂ and 2 ♀; Río Maullín; III.1957; L. Peña. – MHNS; 2 ♂ (sub mislabelled paratypes of Achilia globiceps n° 1650 and 1651); Frutillar; 20.IX.1954; G. Kuschel. – MNHN; 6 ♂; Frutillar; 41° 08’S; 20.IX.1954; G. Kuschel. – MHNG; 11 ♂; Frutillar Bajo, Universitad Chile Forest Reserve; 100 m; 22.XII.1984/02.II.1985; S. & J. Peck; FIT ravine mixed forest. – DBUC; 2 ♂; same data; S. & J. Peck. – FMNH (FMHD #85-935, #85-51); 1 ♂; Frutillar Bajo, Universitad Chile Forest Reserve; 100 m; 22.XII.1984; S. & J. Peck; bracket fungi, berlese. – MHNG; 1 ♂; Petrohué; 30.I.1979; A. De Chambrier. – MHNG; 27 ♂ and 62 ♀; Vicente Perez National Park, Salto Petrohué; 150 m; 23.XII.1984; S. & J. Peck; mixed forest litter, berlese. MHNG; 10 ♂ and 21 ♀; same locality; 04.II.1985; S. & J. Peck; mixed forest, sifted litter. – DBUC; 4 ♂; same data; S. & J. Peck. – FMNH (FMHD #85-938, #85-54); 14 ♂ and 17 ♀; same data; S. & J. Peck. – FMNH (FMHD #85-995, #85-112); 5 ♂ and 3 ♀; same data; S. & J. Peck. – UNHC; 1 ♂ and 3 ♀; Saltos Petrohué, 6.4 km SW Petrohué; 140 m; 28.XII.1982; A. Newton & M. Thayer; valdivian rainforest, berlese, leaf & log litter, forest floor. – FMNH (FMHD #97- 8); 4 ♂; Vicente Perez Rosales National Park, 9.2 km NE Ensenada, on road to Petrohué; 41° 10.20’S 72° 27.10’W; 125 m; 02-28.I.1997; A. Newton & M. Thayer 987; valdivian rainforest w/ Nothofagus spp., flight intercept trap. – FMNH (FMHD #97-10); 7 ♂ and 14 ♀; same locality; 02.I.1997; A. Newton & M. Thayer 987; valdivian rainforest w/ Nothofagus spp., berlese, leaf & log litter. – FMNH (FMHD #2002- 085); 4 ♂ and 9 ♀; Ensenada, Las Cascadas road, 0.9 km N of La Burbuja; 41° 11.27’S 72° 32.6’W; 80 m; 16.XII.2002; M. Chani; Nothofagus dombeyi , mixed hardwoods, berlese, litter. – FMNH (FMHD #97-38); 11 ♂ and 17 ♀; Vicente Perez Rosales National Park, SW slope Vn Osorno, km 4 to La Burbuja; 41° 09.95’S 72° 30.80’W; 310 m; 27.I.1997; A. Newton & M. Thayer 1007; secondary valdivian rainforest w/ Nothofagus dombeyi- Eucryphia cordifolia berlese, leaf & log litter. – MHNG; 1 ♀; La Arena, 45 km SE Puerto Montt; 100 m; 25.XII.1984; S. & J. Peck. – FMNH (FMHD #97-16); 6 ♂ and 26 ♀; Lago Chapo, near SE end, km 9.9 on road from Rollizo; 41° 30.63’S 72° 23.98’W; 385 m; 04.I.1997; A. Newton & M. Thayer 989; valdivian rainforest on steep slope, berlese, leaf & log litter. – MSNG; 1 ♂; Cruce Abtao; SyTC-227; 21.II.1989; S. & T. Cekalovic. – MSNG; 4 ♂ and 5 ♀; Abtao, TC-282; 23.II.1991; T. Cekalovic. – Osorno prov.: FMNH (FMHD #96-248); 4 ♂ and 4 ♀; 15.1 km W Puaucho; 40° 34.97’S 73° 37.68’W; 50 m; 30.XII.1996; A. Newton & M. Thayer 984; valdivian rainforest remnant in sm. ravine, w/large ferns, berlese, leaf & log litter. – MHNG; 1 ♂; Puyehue; 05.XI.1979; A. De Chambrier. – MHNG; 5 ♂ and 3 ♀; Puyehue National Park, Anticura Repucura trail; 500 m; 0 6. II.1985; S. & J. Peck; forest litter. – DBUC; 2 ♂ and 1 ♀; same data; S. & J. Peck. – FMNH (FMHD #4, #85-113); 8 ♀; same data; S. & J. Peck. – UNHC; 10 ♂ and 2 ♀; Puyehue National Park, 4.1 km E Anticura, trap site 662; 430 m; 19-26.XII.1982; A. Newton & M. Thayer; valdivian rainforest, screen sweeping. – FMNH; 3 ♂; same data; A. Newton & M. Thayer; valdivian rainforest, screen sweeping at dusk. – FMNH; 30 ♂ and 85 ♀; same data; A. Newton & M. Thayer; valdivian rainforest, voucher associated with larvae. berlese, leaf and log litter. – A. Newton & M. Thayer; valdivian rainforest, FMNH; 8 ♂; same data; A. Newton & M. Thayer; valdivian rainforest, window trap 662. – FMNH (FMHD# 96-250); 166 ♂; Puyehue National Park, 4 km E Anticura; 40° 39.73’S 72° 08.10’W; 460 m; 30.XII.1996/30.I.1997; A. Newton & M. Thayer 985-1; valdivian rainforest w/large, Saxegothea , flight intercept trap. – FMNH (FMHD #97-5); 101 ♂; same locality; 30.I.1997; A. Newton & M. Thayer 985-3; valdivian rainforest w/large, Saxegothea , flight intercept trap. – FMNH (FMHD# 97-4); 14 ♂ and 1 ♀; same data; A. Newton & M. Thayer 985-2; valdivian rainforest w/ large, Saxegothea , flight intercept trap. – FMNH (FMHD# 97-39); 65 ♂ and 108 ♀; same locality; 30.I.1997; A. Newton & M. Thayer 985-3; valdivian rainforest w/large, Saxegothea berlese, leaf & log litter. – FMNH (FMHD# 97-40); 21 ♂ and 65 ♀; same locality; 30.I.1997; A. Newton & M. Thayer 985-2; valdivian rainforest w/large, Saxegothea , berlese, leaf and log litter – FMNH (FMHD# 97-41); 25 ♂ and 18 ♀; same data; A. Newton & M. Thayer 985-1; valdivian rainforest w/large, Saxegothea , berlese, leaf & log litter. – MHNG; 15 ♂ and 9 ♀; Puyehue National Park, Aguas Calientes, Pionero track; 500 m; 28.XII.1984; S. & J. Peck; sifted forest stick litter. – FMNH (FMHD # 85-928, #85-43); 1 ♂ and 1 ♀; same data; S. & J. Peck. – FMNH (FMHD #2002-090); 11 ♂ and 24 ♀; Puyehue National Park, Ruta 215; km 4.5 of Aduana station; 40° 40.23’S 72° 05.21’W; 580 m; 19.XII.2002; A. Newton, M. Thayer, D. J. Clarke & M. Chani 1071; valdivian rainforest, berlese, leaf & log litter. – FMNH; 2 ♂ and 3 ♀; Puyehue National Park, Antillanca road; 470 m; 20-25.XII.1982; A. Newton & M. Thayer; valdivian rainforest, berlese, vouchers associated with larvae, leaf & log litter, forest floor.– MHNG; 3 ♂; Puyehue National Park,Antillanca road; 500-1000 m; 18-20.XII.1984; S. & J. Peck; car netting. – FMNH (FMHD #85-923, #85-38); 1 ♂; same data; S. & J. Peck. – MHNG; 6 ♂ and 25 ♀; Puyehue National Park, Aguas Calientes, station 25a; 400- 500 m; 31.XII.1990/01.I.1991; M. Agosti & D. Burckhardt. – PHPC; 3 ♂ and 5 ♀; Puyehue National Park, near Termes Aguas Calientes, 26.2 km E Entre Lagos; 40° 44.130’S 72° 18.427’W; 460 m; 09-12. III.2008; H. Wood & C. Griswold. – UNHC; 1 ♂; 7.7 km NE Termas de Puyehue, site 664; 200 m; 19-25.XII.1982; A. Newton & M. Thayer; valdivian rainforest, berlese, leaf & log litter, forest floor. – MHNG; 1 ♂; Umg. Osorno; H. Franz. – MHNG; 3 ♂ and 13 ♀; Pucatrihue, 65 km W Osorno, station 21; 40° 28’S 73° 43’W; 150 m; 04.XII.1984; D. Burckhardt; valdivian rainforest, sifting of moss on dead tree trunks, branches and rocks and of vegetable detritus. – MHNG; 32 ♂ and 123 ♀; 3 km S Maicolpué, Bahìa Mansa; 200 m; 03.II.1985; S. & J. Peck; mixed forest litter. – FMNH (FMHD #85-994, #85-111); 1 ♂ and 1 ♀; same data; S. & J. Peck. – FMNH (FMHD # 96-247); 1 ♂ and 1 ♀; Hills S of Maicolpué; 40° 36.57’S 73° 44.91’W; 160 m; 30.XII.1996; A. Newton & M. Thayer 983; disturbed valdivian rainforest, berlese, leaf & log litter. – MSNG; 1 ♂ and 1 ♀; Los Ñilques; TC-553; 17.I.1998, T. Cekalovic. – Región Los Ríos: Ranco prov.: FMNH (FMHD #85-921, #85-36); 1 ♀; 34 km WNW La Unión; 700 m; 17.XII.1984; S. & J. Peck; litter mixed forest, berlese. – UNHC; 3 ♂ and 1 ♀; 4.1 km W Anticura, site 663; 270 m; 19-25.XII.1982; A. Newton & M. Thayer; valdivian rainforest, flight intercept (windows) trap. – FMNH; 1 ♂ and 16 ♀; same data;A. Newton & M. Thayer; valdivian rainforest, voucher associated with larvae. – Valdivia prov:MHNG; 1 ♂; 35 km N Valdivia; 12.V.1979; T. Cekalovic. – DBUC; 1 ♂ and 2 ♀; same data; T. Cekalovic. – FMNH (FMHD #97-42); 1 ♂; Rincón de La Piedra, turnoff, 14.8 km SE Valdivia; 39° 55’ 32’’S 73° 06’ 27’’W; 50 m; 0 2. II.1997; A. Newton & M. Thayer 990; disturbed Valdivian rainforest, with Nothophagus dombeyi and Podocarpus saligna , berlese, litter in ground bromeliad (? Puya sp.). leaf axils (live & dead). – FMNH (FMHD #97-18); 1 ♂; same data; 11.I-01.II.1997; A. Newton & M. Thayer 990; disturbed Valdivian rainforest, with Nothophagus dombeyi and Podocarpus saligna , flight intercept (windows) trap. – FMNH; 1 ♂; Casa de Piedra, Lago Calafquen; 26.I.1995; T. Cekalovic. – Región Araucanía: Cautín prov.: MHNG; 4 ♂; 15 km NE Villarica, Flor del Lago; 300 m; 14.XII.1984/10.II.1985; S. & J. Peck; 2 FIT Nothophagus forest. – FMNH (FMHD #85-999, #85-116); same data; 1 ♂ and 1 ♀; S. & J. Peck. – MHNG; 2 ♂; same locality; 14.XII.1984; S. & J. Peck; sifting litter Nothophagus forest. – MHNG; 1 ♂ and 2 ♀; 15 km NE Villarica, Flor del Lago, station 16; 500 m; 10.II.1985; S. & J. Peck; sifting forest litter. – UNHC; 2 ♂ and 1 ♀; Bellavista, North shore Villarica Lake, site 655; 310 m; 15-30.XII.1982; A. Newton & M. Thayer; valdivian rainforest, flood debris forest stream. – FMNH; 4 ♂ and 3 ♀; same data; A. Newton & M. Thayer; valdivian rainforest, vouchers associated with larvae. – MHNG; 5 ♀; Huerquehue National Park, station 17a; 800 m; 22–25.XII.1980; M. Agosti & D. Burckhardt; forest litter. – DBUC; 2 ♂; same data; M. Agosti & D. Burckhardt. – FMNH; 1 ♂ and 4 ♀; Ñielol National Park, near Temuco, site 652; about 250 m; 14/30.XII.1982; A. Newton & M. Thayer; native forest remnants with Nothophagus, vouchers associated with larvae, leaf & log litter, forest floor. – FMNH (FMHD #85-909, P#85-23); 1 ♀; Cerro Ñielol National Park, Temuco; 300 m; 13.XII.1984; S. & J. Peck; mixed forest litter. – Malleco prov.: MHNG; 6 ♂; 17 km W Angol; 800 m; 08.XII.1984/16.II.1985; S. & J. Peck; FIT mixed Nothophagus. – DBUC; 1 ♂; same data; S. & J. Peck. – MHNG; 1 ♂; Nahuelbuta National Park, 45 km W Angol; 1400 m; 09.XII.1984/16.II.1985; S. & J. Peck; Nothophagus - Araucaria forest, car trap. – MHNG; 1 ♀; Purén, Contulmo Natural Monument; 350 m; 13.II.1985; S. & J. Peck; mixed forest litter. – MHNG; 3 ♂; Purén, Contulmo Natural Monument; 300- 600 m; 11.XII.1984/13.II.1985; S. & J. Peck; FIT mixed evergreen forest & MT Nothophagus forest. – FMNH (FMHD# 2002-64); 1 ♂ and 3 ♀; Contulmo Natural Monument, Sendero Lemu Mau; 38° 00.74’S 73° 11.13’’W; 410 m; 08.XII.2002; A. Newton & A. Solodovnikov 1059; Nothophagus obliqua-Eucryphia cordifolia w/fern & bamboo understory, flood debris, small stream – Región Bío Bío: Arauco prov.: MSNG; 14 ♂ and 24 ♀; Rio Caramávida; TC-588; 21. III.1999; T. Cekalovic. – Concepción prov.: MHNG; 10 ♂ and 16 ♀; Pinares; 18.III.1973; T. Cekalovic. – DBUC; 1 ♂ and 1 ♀; same data; T. Cekalovic. – MHNG; 1 ♂; Pinares; 20.XII.1970; T. Cekalovic. – MHNG; 1 ♂ and 1 ♀; Hualpén; 05.III.1973; T. Cekalovic. – FMNH; 2 ♂ and 2 ♀; Puente Pelun, TC-342; 18.I.1993; T. Cekalovic. – MSNG; 7 ♂ and 4 ♀; Puente Pelun; TC-358; 21.II.1993, T. Cekalovic. – MSNG; 1 ♂ and 3 ♀; Chiguayante; TC-236; 11.IX.1990; T. Cekalovic. – MSNG; 1 ♂; Estero Nonguén; TC-460; 27.I.1996; T. Cekalovic. – MSNG; 10 ♂ and 5 ♀; same locality; TC-462; 06.II.1996, T. Cekalovic. – MSNG; 1 ♂ and 1 ♀; same locality; TC-540; 26.III.1997; T. Cekalovic. – MSNG; 7 ♂ and 3 ♀; same locality; TC-541a; 27.III.1997; T. Cekalovic. – MSNG; 1 ♂; Concepciòn, Lonco; TC-245; 01.XI.1989; T. Cekalovic. – MSNG; 1 ♀; Periquillo; TC-311; 15.IX.1992, T. Cekalovic. – Ñuble prov.: MNHN; 1 ♀ (sub A. frontalis ); Cordillera of Chillán; 36° 54’S; P. Germain.