Lonchodryinus balticus Olmi & Guglielmino

Olmi, Massimo & Guglielmino, Adalgisa, 2012, A contribution to the knowledge of Dryinidae from Late Eocene Baltic amber (Hymenoptera: Chrysidoidea), with description of new subfamily, Protodryininae subfam. nov., Zootaxa 3351, pp. 15-26 : 16-17

Lonchodryinus balticus Olmi & Guglielmino , sp. nov.

( Fig. 1 )

Diagnosis. Female fully winged; ocelli present; occipital carina complete; epicnemium absent; fore wing provided of three basal cells enclosed by pigmented veins (costal, median and submedian); distal part of stigmal vein longer than proximal part; pterostigma oval, about four times as long as broad; protrochanter short and broad, less than twice as long as broad, not composed of stalk and rounded distal apex; protarsus chelate; chela without rudimentary claw; midleg with one tibial spur. Male unknown.

Material examined. HOLOTYPE, female, in Baltic amber. Type horizon: Late Eocene (40–50 mybp), Baltic amber (Succinite). The locality where the piece was originally found is unknown. The type specimen will be deposited in SNMS, though actually it is in JVC (the buying process is being dealt with).

Description. Female: fully winged; length 4.0 mm. Colour apparently black, except anterior half of face testaceous and antenna and legs brown-testaceous. Antenna 10-segmented, short and slender, covered with dense and short hairs, slightly thickened distally; antennal rhinaria absent; antennal segments in following proportions: 10:6:20:15:10:9:9:8:7:9; antenna much shorter than body, more than three times as long as head: 96:27. Head very slightly convex, shiny, with vertex hairless, punctate, unsculptured among punctures; face hairless, sculptured by many longitudinal striae; clypeus and mandible not visible; occipital carina complete, laterally not reaching eyes; occiput excavated; eye normally bulging, shorter than head (19:27); POL = 2; OL = 2.5; OOL = 10; OPL = 7; TL = 7; greatest breadth of posterior ocellus about as long as POL; frontal line present from anterior ocellus to face testaceous spot, not visible on testaceous spot, where apparently it is replaced by one longitudinal furrow. Pronotum short, flat; anterior transverse impression not visible, because hidden under head; disc flat; pronotum hairless, dull, punctate, unsculptured among punctures, much shorter than head (10:27); pronotal tubercle reaching tegula. Scutum shiny, hairless, punctate, unsculptured among punctures, much longer than pronotum (22:10). Notauli complete, posteriorly separated; minimum distance between notauli about as long as antennal segment 2. Scutellum much shorter than scutum (8:22), hairless, as sculptured as scutum. Metanotum shorter than scutellum (4:8), with sculpture not distinctly visible. Propodeum not visible, because hidden under wings. Mesopleuron and metapleuron only partly visible, with sculpture not distinct. Shape of head, scutum, scutellum, metanotum and propodeum usual for Lonchodryinus . Fore wing hyaline, not darkened, with usual venation of Lonchodryinus . Pterostigma oval, longer than broad (25:6). Marginal cell open. Stigmal vein S-shaped, with distal part much longer than proximal part (26:11), almost reaching margin of fore wing; stigmal vein forming angle between proximal and distal parts; fore wing with three basal cells clearly enclosed by pigmented veins (costal, median and submedian). Hind wing not visible, because hidden under fore wing. Foreleg segments in following proportions: 19 (coxa): 6 (trochanter): 39 (femur): 24 (tibia): 21 (tarsal segment 5). Tarsal segments 2, 3 and 4 not visible. Enlarged claw slightly shorter than tarsal segment 5 (16:21). Protrochanter short, not slender, not curved, without long proximal stalk, less than twice as long as broad (6:4). Rudimentary claw not visible. Arolium much shorter than enlarged claw (3:16). Enlarged claw weakly curved, with inner margin not visible, because of closed chela. Distal apex of enlarged claw apparently pointed. Segment 5 of protarsus with inner side not visible, because of closed chela. Midleg segments in following proportions: 11 (coxa): 7 (trochanter): 31 (femur): 25 (tibia): 18 (tarsal segment 1): 7 (tarsal segment 2): 5 (tarsal segment 3): 3 (tarsal segment 4): 7 (tarsal segment 5). Hind leg segments in following proportions: 22 (coxa):? (trochanter not visible): 42 (femur): 33 (tibia): 21 (tarsal segment 1): 10 (tarsal segment 2): 7 (tarsal segment 3): 5 (tarsal segment 4): 7 (tarsal segment 5). Petiole not visible. Maxillary palpi 6-segmented. Labial palpi not visible. Shape, length and breadth of fore wing usual for Lonchodryinus . Shape and morphology of the body usual for Lonchodryinus . Tibial spurs not visible.

Male. Unknown.

Hosts. Unknown.

Etymology. From Baltic amber.

Remarks. Lonchodryinus balticus sp. nov., is the first fossil Lonchodryinus Kieffer, 1905 and one of the few Anteoninae known as fossils [ Deinodryinus areolatus ( Ponomarenko, 1975) , Deinodryinus velteni Guglielmino & Olmi, 2011 , Janzeniola baltica ( Olmi, 2000) ]. L. balticus sp. nov. is very similar to the only extant species of Lonchodryinus known from the Western Palaearctic region: L. ruficornis ( Dalman, 1818) . Though the holotype of L. balticus is almost completely covered with mould ( Fig. 1 ), so that a complete comparison with L. ruficornis is very difficult, the following differences are easily recognizable: notauli complete in L. balticus (0.30-0.75 length of scutum in L. ruficornis ); distal part of stigmal vein more than twice as long as proximal part in L. balticus (as long as, or slightly shorter, or slightly longer than proximal part in L. ruficornis ).

Figures

FIGURES 1, 2. 1. Lonchodryinus balticus sp. nov., holotype, female, length 4.0 mm; 2. Protodryinus eocenicus gen. et sp. nov., female holotype, length 5.0 mm