Aglaotilla submetallescens ( Turner, 1916 ) Brothers, 2018

Brothers, Denis J., 2018, Aglaotilla, a new genus of Australian Mutillidae (Hymenoptera) with metallic coloration, Zootaxa 4415 (2), pp. 357-368 : 361-367

Aglaotilla submetallescens ( Turner, 1916) , comb. nov.

( FIgS 1–26 )

Ephutomorpha submetallescens Turner, 1916: 116 –117; ♀ (holotype: Brisbane, NHMUK). “ Ephutomorpha ” submetallescens ; Brothers 1984: 9; ♀, ♂.

Diagnosis. FEMALE: INTEgUMENT dARk METALLIc bLUE ON HEAd ANd MESOSOMA , dARk METALLIc PURPLE WITH gOLdEN SHEEN ON METASOMA; T1 WITH TRANSVERSE WHITE APIcAL fRINgE; T2 MOSTLY bLAck PUbEScENT EXcEPT fOR ScATTEREd WHITE EREcT SETAE fORMINg A dIAgONAL ELONgATE PATcH ANTEROLATERALLY, ANd APIcAL OVAL PATcH Of WHITE dIAgONAL ANd TRANSVERSE SETAE; T3–T5 EAcH WITH SMALL INdEfINITE APIcAL PATcH Of WHITE SETAE; HEAd NARROWER THAN MESOSOMA , gENAL cARINA AbSENT, POSTgENAL cARINA ScARcELY INdIcATEd; MESOSOMA AbOUT 1.7 × AS LONg AS WIdE, ITS WIdEST POINT AT SMALL AcUTE TUbERcLE bEARINg METATHORAcIc SPIRAcLE, dORSAL ANd POSTERIOR SURfAcE cOARSELY bUT dENSELY RETIcULATE WITH INTERSPAcES dULL ANd gRANULATE; MESO- ANd METATIbIAE WITHOUT SUbAPIcAL SPINES, INNER METATIbIAL SPUR AbOUT 1.7 × LENgTH Of OUTER SPUR; T2 LONgER THAN WIdE IN dORSAL VIEW, fELT LINE VERY SHORT, AbOUT 0.1 × LATERAL LENgTH Of T2; S2 SPARSELY PUNcTATE ANd SMOOTH, WITHOUT ANY INdIcATION Of LATERAL fELT LINE OR VENTRAL TRANSVERSE dEPRESSION. MALE: INTEgUMENT PREdOMINANTLY METALLIc PURPLE WITH gOLdEN SHEEN, HEAd ANTERIORLY bLUISH-gREEN; T2 WITH bLAck SETAE EXcEPT fOR bROAdLY OVAL INdEfINITE ANTEROLATERAL PATcH Of WHITE SETAE, T3–T7 ENTIRELY WITH WHITE SETAE, HYPOPYgIUM WITH bLAck SETAE; gENAL ANd POSTgENAL cARINAE AbSENT; ScAPE VENTRALLY WITH dENSE EREcT PLUMOSE SETAE AbOUT AS LONg AS ScAPE WIdTH; MANdIbLE WITH SLIgHT INNER fLANgE AT AbOUT MId-LENgTH, dISTINcT VENTRAL bASAL LAMELLATE TOOTH APIcALLY ObLIqUE ANd AbOUT AS LONg AS MANdIbLE bASAL HEIgHT; HUMERAL ANgLE STRONgLY PROdUcEd ANd dENTATE; TEgULA SMOOTH, bROAdLY OVAL WITH EVENLY cONVEX POSTERIOR MARgIN; PARAMERE LONg ANd SUbcYLINdRIcAL WITH ScATTEREd fINE SHORT EREcT SETAE; cUSPIS WITH VENTRAL LObE SHORT ANd TRIANgULAR, SHORTER THAN dIgITUS.

Description. FEMALE: LENgTH 11.9 (10.3–12.9) MM. COLORATION ALMOST ENTIRELY METALLIc, HEAd ANd MESOSOMA gENERALLY dARk bLUE ANd METASOMA gENERALLY PURPLE; HEAd WITH gREENISH gOLd SHEEN IN MALAR AREA, ANTENNAL TUbERcLE ANd cLYPEUS dARk REddISH bROWN, POSTgENAL bRIdgE dARk TO MEdIUM REddISH bROWN; T1 WITH gREENISH SHEEN ON POSTEROdORSAL fAcE; T2 WITH gOLdEN SHEEN ALONg LATERAL MARgIN; HYPOPYgIUM VERY dARk bROWN TO bLAck; ANTENNA dARk TO MEdIUM REddISH bROWN EXcEPT fOR SLIgHT PURPLE SHEEN ON ScAPE; MOUTHPARTS dARk TO MEdIUM REddISH bROWN; LEgS METALLIc PURPLE WITH bLUE ANd gOLdEN SHEEN, EXcEPT TARSI dARk TO MEdIUM REddISH bROWN, TIbIAL SPURS WHITISH. PUbEScENcE MIXEd dARk ANd WHITE bUT MAINLY WHITE ON HEAd, METASOMAL STERNA ANd LEgS, ELSEWHERE MAINLY bLAck PUbEScENT; T1 WITH dENSE WHITE TRANSVERSE bANd ON POSTEROdORSAL fAcE; T2 dARk PUbEScENT EXcEPT WHITE ALONg ANTERIOR ANd LATERAL MARgINS, WHITE MEdIAN OVAL ANd LATERAL SHORT PATcHES ON POSTERIOR MARgIN, ANd INdISTINcT SPARSE WHITE SETAE fORMINg A NARROW dIAgONAL LINE dORSALLY ON EAcH SIdE Of ANTERIOR HALf; T3–T5 EAcH WITH SMALL fAIRLY SPARSE WHITE MEdIAN APIcAL PATcH ANd WHITE PUbEScENcE LATERALLY; T6 ANd S6 ENTIRELY dARk PUbEScENT. Head: NARROWLY TRANSVERSE, 1.25 (1.20–1.30) × AS WIdE AS LONg, 0.88 (0.84–0.92) × AS WIdE AS MESOSOMA , dENSELY PUNcTATE WITH SHINY INTERSPAcES, PUNcTATION bEcOMINg fINER ANTERIORLY, WITH SEMI-dEcUMbENT ANd EREcT bRAcHYPLUMOSE OR SIMPLE SETAE. OccIPITAL cARINA STRONg, PRESENT ONLY dORSALLY. SIdES bEHINd HEAd MOdERATE, ALMOST PARALLEL ANd cONVEX, dISTINcT fROM POSTERIOR MARgIN bUT POSTEROLATERAL ANgLE ROUNdEd; HEAd LENgTH bEHINd EYE 0.68 (0.53–0.78) × EYE LENgTH. EYE VERY STRONgLY PROTRUdINg, fRONS WIdTH 0.57 (0.56–0.60) × HEAd WIdTH. ANTENNAL TUbERcLES SIMPLE, SMOOTH, ScARcELY SEPARATEd bASALLY. CLYPEUS LATERALLY cONcAVE, WITH RAISEd bROAdLY TRIANgULAR PUNcTATE ANd SETOSE REgION AbOVE cOMPLETE SINUATE TRANSVERSE cARINA, ITSELf AbOVE VERY SHORT dEPRESSEd TRANSVERSE gROOVE, VENTRAL MARgIN SLIgHTLY cONVEX. MALAR SPAcE 1.88 (1.88–1.89) × bASAL HEIgHT Of MANdIbLE. GENAL cARINA APPARENTLY AbSENT, PERHAPS REPRESENTEd bY VENTRAL IRREgULAR cARINATE ELEVATION NEAR ORAL fOSSA. POSTgENAL cARINA IRREgULAR ANd SHORT, dISTANT fROM EdgE Of OccIPITAL dEPRESSION ANd fROM ORAL fOSSA. ORAL fOSSA SLIgHTLY LONgER THAN SMOOTH POSTgENAL bRIdgE. ScAPE SIMPLE, fAIRLY SPARSELY ANd fINELY PUNcTATE AbOVE. PEdIcEL 1.38 (1.38–1.38) × AS LONg AS WIdE, 0.53 (0.52–0.55) × AS LONg AS F1; F1 2.30 (2.22–2.33) × AS LONg AS WIdE, 1.51 (1.43–1.62) × AS LONg AS F2. MANdIbLE SLENdER, TAPERINg, VERY WEAkLY ARcUATE, bIdENTATE APIcALLY WITH SUbAPIcAL TOOTH ScARcELY EVIdENT AT APEX Of WEAk INNER LONgITUdINAL cARINA, WITH SLIgHT NARROW VENTRAL bASAL LAMELLA ENdINg APIcALLY SMOOTHLY OR AbRUPTLY; HEIgHT AcROSS APEX Of bASAL LAMELLA 0.75 (0.67–0.88) × MANdIbULAR bASAL HEIgHT, HEIgHT IMMEdIATELY bEYONd bASAL LAMELLA 0.75 (0.67–0.83) × HEIgHT AcROSS APEX Of LAMELLA, bASAL LAMELLA LENgTH 0.81 (0.78–0.82) × MANdIbULAR bASAL HEIgHT. Mesosoma : 1.70 (1.64–1.73) × AS LONg AS WIdE, fAIRLY EVENLY OVATE IN dORSAL VIEW, PRONOTUM LONg WITH SIdES ScARcELY dIVERgINg POSTERAd TO INcISEd SPIRAcLE, MARgIN THEN EVENLY cONVEX TO SLIgHTLY PROJEcTINg PROPOdEAL SPIRAcLE, bROAdEST AT SMALL AcUTE TUbERcLE bEARINg METATHORAcIc SPIRAcLE, MAXIMUM WIdTH 1.24 (1.23–1.24) × WIdTH AcROSS HUMERAL ANgLES, 1.15 (1.13–1.16) × WIdTH AT bASE Of PROPOdEAL SPIRAcLE; PROPOdEUM NARROWEd ANd EVENLY ROUNdEd POSTERIORLY, dISk ANd dEcLIVITY MERgINg. DORSUM cONVEX, dENSELY RETIcULATE, MORE cOARSELY SO ANTEROMEdIALLY, WITH INTERSPAcES cONSPIcUOUSLY SHAgREENEd/gRANULATE ANd dULL, cLOTHEd WITH SHORT SEMIdEcUMbENT SIMPLE SETAE ANd LONg EREcT bRAcHYPLUMOSE OR SIMPLE SETAE. PRONOTAL dORSAL fAcE (EXcLUdINg SMOOTH ANTERIOR cOLLAR) WITH ANTERIOR MARgIN SLIgHTLY cONVEX, AbRUPTLY ROUNdEd IN PROfILE, EPAULET A SMALL TUbERcLE WITH MINUTE TUfT Of SHORT fINE SETAE, HUMERAL ANgLE AbRUPT ANd WEAkLY dENTATE; POSTEROdORSAL MARgIN NOT EVIdENT. MESO- ANd METANOTUM dELIMITEd LATERALLY bY dISTINcT LONgITUdINAL RIdgE. LATERAL fAcE Of PRONOTUM RETIcULATE, LONgITUdINALLY RUgOSE ANTEROVENTRALLY, INTERSPAcES SHINY; PRONOTAL-MESOPLEURAL SUTURE ENTIRELY dISTINcT. MESOPLEURON dENSELY PUNcTATE-RETIcULATE EXcEPT SMOOTH ANTERIORLY, INTERSPAcES SHINY bUT gRANULATE ON dORSAL THIRd; VERTIcAL MESOPLEURAL RIdgE WELL dEVELOPEd ANd cARINATE ON VENTRAL HALf, MERELY WEAkLY cONVEX ANd cOARSELY RETIcULATE TOWARdS METATHORAcIc SPIRAcLE dORSALLY, WITH EREcT bRAcHYPLUMOSE SETAE. METAPLEURON dENSELY ANd cOARSELY PUNcTATE WITH SHINY INTERSPAcES EXcEPT SMOOTH ANd SHININg ON SHORT STRIP ALONg ANTERIOR MARgIN. METAPLEURAL-PROPOdEAL SUTURE A dISTINcT RIdgE ON POSTERIOR HALf, OTHERWISE ObScUREd bY ScULPTURE. LATERAL fAcE Of PROPOdEUM IRREgULARLY RETIcULATE WITH gRANULATE INTERSPAcES, SEPARATEd fROM POSTEROdORSAL fAcE bY dISTINcT RIdgE. METASTERNAL PROcESS TRIANgULAR, APIcALLY AcUTE, AbOUT AS LONg AS METAcOXAL HEIgHT. Legs: FORE LEg WITHOUT TARSAL cOMb. MId ANd HINd TIbIAE WITHOUT dORSAL OR LATERAL SPINES, EAcH WITH 2 OR 3 SMALL APIcAL SPINES ON ScARcELY SWOLLEN TIbIAL APEX; EAcH SUbAPIcALLY WITH VERY INcONSPIcUOUS SEcRETORY PORE NEAR bASE Of INNER APIcAL SPUR. METAcOXA WITH STRONg EVEN LONgITUdINAL cARINA ON INNER MARgIN, ENdINg POSTERIORLY SHORTLY bEfORE SMALL APIcAL LAMELLATE TUbERcLE; INNER METATIbIAL SPUR 1.68 (1.56– 1.73) × AS LONg AS OUTER SPUR, 0.44 (0.43–0.46) × AS LONg AS METAbASITARSUS. Metasoma: 1.18 (1.16–1.19) × AS WIdE AS MESOSOMA . T1 1.73 (1.69–1.80) × AS WIdE AS LONg, 0.51 (0.50–0.52) × WIdTH Of T2; VERY fINELY ANd SPARSELY PUNcTATE ANd MOSTLY SMOOTH ON fLATTENEd ANTERIOR fAcE bUT VERY dENSELY ANd fINELY PUNcTATE ON SHORT POSTEROdORSAL fAcE, MARgIN bETWEEN fAcES dISTINcT ANd cOARSELY RETIcULATE; EVENLY EXPANdEd fROM bASE, POSTEROdORSAL fAcE SLIgHTLY cONSTRIcTEd APIcALLY; WITH EREcT PLUMOSE SETAE bASALLY, bRAcHYPLUMOSE ANd SIMPLE SETAE ELSEWHERE, ANd dENSE POSTERIOR fRINgE Of dEcUMbENT SIMPLE SETAE ORIENTEd dIAgONALLY MESAd ANd EXTENdINg THE ENTIRE WIdTH Of T1; ANTERIOR AURIcLE PROMINENT, VERTIcALLY LAMELLATE bUT APPARENTLY dENTATE fROM AbOVE. T2 1.20 (1.17–1.24) × AS LONg AS WIdE IN dORSAL VIEW, ANTEROMEdIALLY ALMOST IMPERcEPTIbLY dEPRESSEd ON ANTERIOR HALf; fINELY ANd VERY dENSELY PUNcTATE bUT MORE cOARSELY ANd SPARSELY SO ANTEROLATERALLY, INTERSPAcES SHINY; cLOTHEd WITH dEcUMbENT SLENdER SIMPLE OR LATERALLY fLATTENEd SETAE ANd EREcT SIMPLE OR VERY INcONSPIcUOUSLY bRAcHYPLUMOSE SETAE, INcONSPIcUOUS POSTERIOR bANd Of dEcUMbENT SLENdER SIMPLE SETAE ORIENTEd dIAgONALLY MESAd; LATERAL fELT LINE VERY NARROW, SHORT, 0.12 (0.09–0.15) × LATERAL LENgTH Of T2. T3–T5 SHINY ANd VERY fINELY bUT SPARSELY PUNcTATE, cLOTHEd WITH SEMIdEcUMbENT ANd EREcT SIMPLE SETAE. T6 ENTIRELY ANd EVENLY cONVEX, SHINY ANd VERY fINELY bUT SPARSELY PUNcTATE EXcEPT SMOOTH ON NARROW AREA ALONg MIdLINE, WITH EREcT SIMPLE SETAE. S1 WITH WEAk SIMPLE MEdIAN cARINA, ANTERIOR TRIANgULAR TOOTH ANd POSTERIOR TRANSVERSE ARcUATE cARINA. S2 STRONgLY cONVEX ANTEROMEdIALLY ANd VERY WEAkLY cONcAVE POSTERIAd IN PROfILE, SHINY, fINELY ANd VERY SPARSELY PUNcTATE; WITH EREcT SIMPLE SETAE ANd SPARSE APIcAL fRINgE Of SEMI-dEcUMbENT SIMPLE SETAE; WITHOUT fELT LINE. S3–S6 SHINY, VERY fINELY bUT MOdERATELY TO dENSELY PUNcTATE, WITH EREcT SIMPLE SETAE ANd SPARSE APIcAL fRINgES Of SEMI-dEcUMbENT SIMPLE SETAE. HYPOPYgIUM (S6) WITH APEX SHALLOWLY EMARgINATE.

MALE: LENgTH 9.1–9.9 MM. COLORATION ALMOST ENTIRELY METALLIc, MOSTLY PURPLE; HEAd ANTERIORLY bLUISH-gREEN WITH SLIgHT gOLdEN SHEEN ANd POSTERIORLY (INcLUdINg VERTEX) PURPLE WITH gOLdEN ANd gREENISH HIgHLIgHTS; MESOSOMA PURPLE WITH gOLdEN ANd gREENISH HIgHLIgHTS EXcEPT HIgHLIgHTS bLUE ON ScUTELLUM, dORSELLUM ANd PROPOdEAL APEX, TEgULA PURPLE; METASOMA PURPLE, bUT T1 WITH bLUE HIgHLIgHTS ANTEROMEdIALLY, T2 WITH SLIgHT gOLdEN SHEEN, T3–T7 WITH STRONgER gOLdEN SHEEN, S1 WITH bLUE HIgHLIgHTS, S2 EVENLY PURPLE, S3–S6 ANd HYPOPYgIUM bLAckISH WITH bLUE TO gOLdEN TO PURPLE SHEEN; ScAPE gREEN WITH gOLdEN SHEEN TO PURPLISH AT bASE, PEdIcEL ANd fLAgELLUM NONMETALLIc dARk bROWN; MANdIbLE bASILATERALLY gREENISH TO PURPLISH WITH gOLdEN SHEEN, OTHERWISE NONMETALLIc REddISH bLAck; MAXILLA ANd LAbIUM NONMETALLIc dARk bROWN; WINgS WITH MEMbRANE HYALINE ANd ONLY VERY fAINTLY bROWNISH, ANTERIOR VEINS ON fORE WINg dARk bROWN, OTHER VEINS PALER bROWN; LEgS WITH cOXAE PURPLE, TROcHANTERS bROWN, fEMORA WITH bRONzE SHEEN ANd bLUISH dORSALLY ANd PURPLISH VENTRALLY, TIbIAE WITH bRONzE SHEEN ANd PURPLISH dORSALLY ANd bLUISH VENTRALLY, TARSI NONMETALLIc dARk bROWN, TIbIAL SPURS WHITISH. PUbEScENcE MAINLY WHITISH WITH A fEW INTERSPERSEd dARk SETAE; PREdOMINANTLY dARk MEdIALLY ON cLYPEUS, ScUTUM, ScUTELLUM, dORSELLUM, T2 INcLUdINg APIcAL fRINgE (EXcEPT fOR LARgE ANTEROLATERAL PATcH Of WHITE SETAE), S5, S6 ANd HYPOPYgIUM; METASOMAL APIcAL fRINgES SPARSE ANd POORLY dEfINEd. Head: BROAdLY TRANSVERSE, 1.95 (1.92–1.97) × AS WIdE AS LONg, 0.99 (0.96–1.02) × AS WIdE AS MESOSOMA , ENTIRELY fINELY ANd VERY dENSELY PUNcTATE WITH INTERSPAcES NARROW ANd SHININg, WITH EREcT bRAcHYPLUMOSE SETAE, EREcT PLUMOSE SETAE ON cLYPEUS, ANTEROVENTRAL TO EYE, ON POSTgENAL bRIdgE, ANd VENTRALLY ON ScAPE. EYE VERY STRONgLY PROTRUdINg, fRONS WIdTH 0.63 (0.62–0.64) × HEAd WIdTH. SIdES bEHINd HEAd cONVEX ANd ROUNdEd, NARROWLY MERgINg WITH POSTERIOR MARgIN; HEAd LENgTH bEHINd EYE 0.58 (0.55–0.61) × EYE LENgTH. VERTEX cONVEX, OcELLAR TRIANgLE ScARcELY ELEVATEd. OcELLI MOdERATE, MEdIAN OcELLUS 0.08 (0.07–0.08) × WIdTH Of HEAd, bROAdLY SPAcEd, INTEROcELLAR dISTANcE 1.95 (1.91–2.00) × WIdTH Of MEdIAN OcELLUS; INTEROcELLAR dISTANcE 0.62 (0.60– 0.64) × OcELLOcULAR dISTANcE. BLUNT OccIPITAL cARINA dISTINcT dORSALLY ONLY, NO POSTgENAL cARINA. GENAL cARINA AbSENT. ANTENNAL TUbERcLES fINELY PUNcTATE, SLIgHTLY SEPARATEd bASALLY. ANTENNAL ScRObE dEEP, WELL dELIMITEd LATERALLY, NOT REAcHINg EYE MARgIN, WITH SHORT WEAk cURVEd cARINA dORSOMESALLY ANd SEPARATE PROMINENT TUbERcLE AT dORSOLATERAL EXTREMITY Of ScRObAL MARgIN. CLYPEUS fINELY ANd dENSELY PUNcTATE, dEPRESSEd ANd fLATTENEd EXcEPT fOR SHORT dORSOMEdIAN TRIANgLE, VENTRAL MARgIN SINUATE bUT STRAIgHT MEdIALLY. MALAR SPAcE 0.91 (0.90–0.92) × bASAL HEIgHT Of MANdIbLE. ORAL fOSSA SLIgHTLY LONgER THAN SMOOTH POSTgENAL bRIdgE. ScAPE WITH SINgLE WEAk LONgITUdINAL RIdgE ANTEROVENTRALLY; VENTRALLY WITH dENSE EREcT PLUMOSE SETAE AbOUT AS LONg AS ScAPE WIdTH. PEdIcEL 0.98 (0.96–1.00) × AS LONg AS WIdE, 0.50 (0.50–0.50) × AS LONg AS F1; F1 1.75 (1.73–1.78) × AS LONg AS WIdE, 0.87 (0.84–0.90) × AS LONg AS F2. MANdIbLE bIdENTATE APIcALLY, IN dORSAL/ ANTERIOR VIEW bROAdLY ANgLEd, gRAdUALLY TAPERINg bUT AbRUPTLY SO AT AbOUT TWO-THIRdS LENgTH , WITH MOdERATE LONgITUdINAL cARINA dORSALLY ENdINg IN SLIgHT SUbAPIcAL TOOTH, APIcAL TOOTH NARROWLY AcUTE, WITH VENTRAL bASAL LAMELLA fORMINg AN EVEN fLANgE WITH dIAgONAL APEX; HEIgHT AcROSS bASAL LAMELLA 0.96 (0.92–1.00) × MANdIbULAR bASAL HEIgHT, HEIgHT IMMEdIATELY bEYONd bASAL LAMELLA 0.57 (0.55–0.60) × HEIgHT AcROSS LAMELLA, bASAL LAMELLA LENgTH 0.89 (0.88–0.90) × MANdIbULAR bASAL HEIgHT. Mesosoma : 1.50 (1.48–1.52) × AS LONg AS WIdE; dENSELY ANd fINELY PUNcTATE EXcEPT IRREgULARLY ANd SHALLOWLY RUgOSE TO SMOOTH ON ANTEROdORSAL TWO- THIRdS Of METAPLEURON, ANd RETIcULATE ON PROPOdEUM, INTERSPAcES NARROW ANd SHININg ; WITH EREcT bRAcHYPLUMOSE ANd SIMPLE SETAE. PRONOTAL dORSAL fAcE (EXcLUdINg ANTERIOR cOLLAR) WITH ANTERIOR MARgIN VERY SLIgHTLY cONcAVE ANd ROUNdEd IN PROfILE, EPAULET A LARgE OVAL PATcH Of fINE dEcUMbENT SETAE; HUMERAL ANgLE PROdUcEd AS A STRONg AcUTE TOOTH ; LATERAL MARgIN WEAkLY SINUATE, ANTERIORLY cONcAVE; POSTEROdORSAL MARgIN bROAdLY V-SHAPEd. TEgULA 0.45 (0.44–0.46) × AS LONg AS ScUTUM, bROAdLY OVATE , NOT REAcHINg MEdIAN LEVEL Of ScUTO-ScUTELLAR SUTURE, cONVEX bUT VERY WEAkLY REcURVEd POSTERIORLY, SMOOTH ANd gLAbROUS EXcEPT fINELY PUNcTATE ANd SETOSE ALONg ANTERIOR ANd POSTEROMESAL MARgINS, POSTERIOR MARgIN EVENLY cONVEX. MESOScUTUM WITH NOTAULUS dEEP ANd EXTENdINg OVER POSTERIOR HALf Of ScUTUM, PARAPSIdAL LINE VERY SHORT ANd IMMEdIATELY MESAd TO dISTINcT SMOOTH LONgITUdINAL RIdgE TWIcE ITS LENgTH ANd EXTENdINg TO TRANSScUTAL fURROW; POSTEROLATERAL cORNER fORMINg A VERY SLIgHT LObE dIScONTINUOUS WITH AXILLA. ScUTELLUM bROAdLY cONVEX ANd PULVINATE; AXILLA PROdUcEd AS A fLATTENEd HORIzONTAL fLANgE, fORMINg A STRONg AcUTE TOOTH POSTERIORLY . METANOTUM SIMPLE, TRANSVERSE, dORSELLUM WEAkLY dELIMITEd LATERALLY. PROPOdEUM EVENLY cONVEX, dISk ANd dEcLIVITY MERgINg IN LATERAL VIEW bUT MARgIN bETWEEN dORSAL/POSTERIOR ANd LATERAL fAcES AbRUPT. METASTERNAL PROcESS bROAdLY TRIdENTATE WITH APEX AcUTE, SLIgHTLY LONgER THAN METAcOXAL HEIgHT. Wings : HYALINE, MEMbRANE ALMOST cOLOURLESS. FOREWINg WITH MOdERATE ELONgATE ScLEROTIzEd PTEROSTIgMA; MARgINAL cELL 2.47 (2.43–2.50) × AS LONg AS HIgH, TRUNcATE APIcALLY WITH VERY SHORT SPURLIkE VEIN OR SLIgHT ANgULATE SWELLINg POSTERIORLY ; THREE cLOSEd SUbMARgINAL cELLS, THIRd SUbMARgINAL cELL 1.18 (1.17–1.20) × AS LONg AS HIgH; dISTANcE bETWEEN THIRd SUbMARgINAL cELL ANd WINg APEX 1.92 (1.83–2.00) × LENgTH Of THIRd SUbMARgINAL cELL. Legs: MId ANd HINd TIbIAE WITHOUT dORSAL SPINES ; EAcH SUbAPIcALLY WITH MINUTE SEcRETORY PORE NEAR bASE Of INNER APIcAL SPUR; APIcAL SPURS fLATTISH dORSALLY, MARgINS MIcROSERRATE, dENSELY cLOTHEd WITH MIcROSETAE. METAcOXA WITH STRONg REgULAR LONgITUdINAL cARINA ON INNER MARgIN ENdINg WELL bEfORE cOXAL APEX; INNER METATIbIAL SPUR 1.70 (1.67–1.73) × AS LONg AS OUTER SPUR, 0.66 (0.63–0.70) × AS LONg AS METAbASITARSUS. Metasoma: 1.00 (0.98–1.02) × AS WIdE AS MESOSOMA . T1 fINELY ANd dENSELY PUNcTATE bUT SPARSELY SO ANTEROMEdIALLY, 0.56 (0.54–0.59) × AS LONg AS WIdE, 0.44 (0.43–0.44) × WIdTH Of T2 , EVENLY EXPANdEd fROM bASE, fLATTENEd ANTERIOR fAcE dISTINcT fROM SHORT POSTEROdORSAL fAcE, SLIgHTLY cONSTRIcTEd APIcALLY, WITH EREcT SLENdER bRAcHYPLUMOSE ANd SIMPLE SETAE ANd VERY SPARSE POSTERIOR fRINgE Of dEcUMbENT SLENdER SIMPLE SETAE; ANTERIOR AURIcLE PROMINENT, VERTIcALLY LAMELLATE bUT APPARENTLY dENTATE fROM AbOVE. T2 0.88 (0.86–0.90) × AS LONg AS WIdE IN dORSAL VIEW , VERY fINELY ANd dENSELY PUNcTATE, INTERSPAcES dISTINcT ANd SHINY, WITH SEMI-dEcUMbENT SIMPLE SETAE ANTEROLATERALLY ANd dENSER LATERALLY fLATTENEd LANcEOLATE SETAE ELSEWHERE, POSTERIOR fRINgE VERY SPARSE ANd INdEfINITE; fELT LINE VERY NARROW ANd LONg, 0.41 (0.40–0.42) × LATERAL LENgTH Of T2. T3–T6 VERY fINELY ANd dENSELY PUNcTATE, INTERSPAcES SHINY, WITH EREcT SIMPLE SETAE, POSTERIOR fRINgES VERY SPARSE ANd INdEfINITE. T7 MOSTLY fINELY ANd dENSELY PUNcTATE WITH EREcT SIMPLE SETAE, PUNcTATION cOARSER ANd SPARSER POSTEROMEdIALLY, APIcAL MARgIN EVENLY cONVEX ANd REcURVEd. STERNA WITH EREcT TO SEMI-dEcUMbENT SIMPLE SETAE. S1 dENSELY ANd fINELY PUNcTATE, WITH dISTINcT SIMPLE LONgITUdINAL cARINA. S2 IN LATERAL VIEW STRONgLY cONVEX ANTERIORLY ANd WEAkLY cONcAVE POSTERIORLY; PUNcTATION cOARSE ANd SPARSE MEdIALLY bUT fINE ANd dENSE ELSEWHERE, INTERSPAcES SHINY; WITH dISTINcT bUT SHORT LATERAL fELT LINE, 0.47 (0.40–0.54) × AS LONg AS fELT LINE ON T2. S3–S6 VERY fINELY bUT SOMEWHAT SPARSELY PUNcTATE, INTERSPAcES dULL ANd SHAgREENEd. S7 cONcEALEd. HYPOPYgIUM SLIgHTLY SHORTER THAN WIdE, fAIRLY SPARSELY PUNcTATE, WEAkLY cONcAVE, MARgINEd LATERALLY bY dISTINcT RIdgE , POSTERIOR MARgIN cONVEX WITH TWO PROMINENT AcUTE TEETH dIVIdINg MARgIN INTO THIRdS. Genitalia: PARAMERE ELONgATE, SLENdER, ALMOST cYLINdRIcAL, NARROWLY ROUNdEd APEX VERY WEAkLY cURVEd dORSAd, WITH INcONSPIcUOUS ScATTEREd SHORT EREcT SIMPLE SETAE. PARAPENIAL LObE WELL dEVELOPEd, A SIMPLE OVATE LObE IN dORSAL VIEW. CUSPIS VERY WEAkLY cONSTRIcTEd APIcALLY, PARAcUSPIS A SHORT cONVEX LObE, VENTRAL LObE SHORT ANd TRIANgULAR; dIgITUS ELONgATEd ANd SLIgHTLY LATERALLY fLATTENEd, PROMINENT ANd LONgER THAN cUSPIS. PENIS VALVE IN LATERAL VIEW WITH dORSAL MARgIN ALMOST STRAIgHT; ANTEROVENTRAL ARM SLIgHTLY NARROWEd APIcALLY, ANTERIOR APEX AcUTE; LATEROVENTRAL LAMELLA SLIgHT ANd MERgINg WITH APEX Of SUbAPIcAL TOOTH; NARROWLY AcUTE APIcAL TOOTH STRONg ANd PROJEcTINg dIAgONALLY fROM APIcAL MARgIN; SEVERAL TINY SETAE LATERAL TO dORSAL HALf Of APIcAL MARgIN ANd A fEW PROMINENT LONg ANd STRONg SETAE PROJEcTINg fROM dORSAL THIRd Of APIcAL MARgIN; IN dORSAL VIEW ANTEROVENTRAL ARM SUbAPIcALLY SLIgHTLY bROAdENEd.

Specimens examined. AUSTRALIA: QUEENSLANd: BRISbANE, AcAcIA RIdgE, 23.XI.1966 (E.C. DAHMS) (2♀♀ “EX NEST Abispa sp., caged No. 1”, 2♂♂ “attracted to caged ♀♀ No. 1”; ♀ ♂ QM, ♀ ♂ DJBC) ; Peak Downs (“P D, 323”) (♀ ZMUC) . (THE HOLOTYPE ♀ fROM BRISbANE, REAREd bY H. HAckER, 8.II.1912, IS IN NHMUK ANd WAS EXAMINEd ANd PHOTOgRAPHEd bY ME IN 1981.)

Remarks. BROTHERS (1984) MENTIONEd 12 fEMALE ANd 3 MALE SPEcIMENS Of THIS SPEcIES WHIcH HAd bEEN REAREd fROM MUd NESTS Of Abispa SP. ( VESPIdAE , EUMENINAE) IN BRISbANE IN 1912, 1961 ANd 1966, ANd WHIcH HE HAd EXAMINEd; UNfORTUNATELY, MOST Of THOSE SPEcIMENS HAd TO bE RETURNEd TO QM SOME TIME AgO ANd WERE NOT AVAILAbLE fOR RE-EXAMINATION NOW. APART fROM THE REcORdS IN WHIcH MALES ANd fEMALES EMERgEd fROM THE SAME NESTS, THUS INdIcATINg THE SEX ASSOcIATION, THE NEWLY EMERgEd fEMALES EXAMINEd HERE HAd bEEN ENcLOSEd IN MESH cAgES ANd PLAcEd UNdER ObSERVATION IN THE fIELd WHERE THE MALES WERE ATTRAcTEd TO THEM (DAHMS, PERS. cOMM.). THESE ATTRAcTION EXPERIMENTS cONfIRMEd THE ASSOcIATION. THE SPEcIMEN fROM “PEAk DOWNS” IS VERY OLd TO JUdgE fROM THE cONdITION ANd STYLE Of THE LAbELS, bUT UNfORTUNATELY WITHOUT A dATE. SPEcIMENS ARE AMONgST THE LARgEST fOR THE gENUS, WITH THE fEMALES EASILY IdENTIfIAbLE bY THE dIAgONAL ANTEROLATERAL bANdS Of SPARSE WHITE SETAE ON T2; MALES ARE LESS cHARAcTERISTIc, HOWEVER. SINcE MOST SPEcIES Of Aglaotilla APPEAR TO HAVE LIMITEd dISTRIbUTIONS, LOcALITY dATA ARE ALSO USEfUL IN MAkINg dETERMINATIONS.

Figures

FIGURES 1–10. Aglaotilla submetallescens (Turner, 1916), comb. nov. 1–5, ♀; 1. Habitus: dorsal view; 2. Habitus: lateral view; 3. Head: frontal view; 4. Metasoma:dorsal view; 5. Metasoma (posterior region of T4, T5, T6): dorsal view. 6–10, ♂; 6. Habitus: dorsal view; 7. Habitus: lateral view; 8. Metasoma: dorsal view; 9. Head: frontal view; 10. Left wings: dorsal view. Scales = 1 mm.

FIGURES 11–18. Aglaotilla submetallescens (Turner, 1916), comb. nov., sculpture. 11–14, ♀; 11. Head: dorsal view; 12. Mesosoma (antero-mesal region of mesoscutum): dorsal view; 13. Metasoma (lateral regions of T2 and S2): lateral view, showing felt line on T2; 14. Metasoma (postero-mesal region of S2): ventral view. 15–18, ♂; 15. Head: dorsal view; 16. Mesosoma (posterior region of mesoscutum, anterior region of mesoscutellum, and right axilla): dorsal view; 17. Mesosoma (mesal region of propodeum): postero-dorsal view; 18. Metasoma (posterior region of T5, T6, T7): dorsal view. Scale = 0.5 mm.

FIGURES 19–26. Aglaotilla submetallescens (Turner, 1916), comb. nov., ♂, genitalia (drawings omit setae). 19–20. Genitalia: dorsal views (19 = left half only); 21–22. Genitalia: ventral views (22 = left half only); 23–24. Genitalia, left half minus penis valve: sagittal view; 25–26.Left penis valve:lateral view. Scales = 0.5 mm.