Lepus Linnaeus 1758

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Lagomorpha, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press, pp. 185-211 : 194

Lepus Linnaeus 1758

Lepus Linnaeus 1758 , Syst. Nat., 10th ed., Vol. 1: 57 .

Type Species: Lepus timidus Linnaeus 1758

Synonyms: Allolagus Ognev 1929 ; Boreolagus Barrett-Hamilton 1911 ; Chionobates Kaup 1829 ; Eulepus Acloque 1899 ; Lagos Palmer 1904 .

Species and subspecies: 32 species with 108 subspecies in 9 subgenera:

Subgenus Lepus (Lepus) Linnaeus 1758

Subgenus Lepus (Macrotolagus) Mearns 1895

Subgenus Lepus (Poecilolagus) Lyon 1904

Subgenus Lepus (Proeulagus) Gureev 1964

Subgenus Lepus (Eulagos) Gray 1867

Subgenus Lepus (Sabanalagus) Averianov 1998

Subgenus Lepus (Indolagus) Gureev 1953

Subgenus Lepus (Sinolagus) Averianov 1998

Subgenus Lepus (Tarimolagus) Gureev 1947

Species Lepus (Macrotolagus) alleni Mearns 1890

Subspecies Lepus (Macrotolagus) alleni subsp. alleni Mearns 1890

Subspecies Lepus (Macrotolagus) alleni subsp. tiburonensis Townsend 1912

Species Lepus (Poecilolagus) americanus Erxleben 1777

Subspecies Lepus (Poecilolagus) americanus subsp. americanus Erxleben 1777

Subspecies Lepus (Poecilolagus) americanus subsp. bairdii Hayden 1869

Subspecies Lepus (Poecilolagus) americanus subsp. cascadensis Nelson 1907

Subspecies Lepus (Poecilolagus) americanus subsp. dalli Merriam 1900

Subspecies Lepus (Poecilolagus) americanus subsp. struthopus Bangs 1898

Subspecies Lepus (Poecilolagus) americanus subsp. virginianus Harlan 1825

Species Lepus (Lepus) arcticus Ross 1819

Subspecies Lepus (Lepus) arcticus subsp. arcticus Ross 1819

Subspecies Lepus (Lepus) arcticus subsp. bangsii Rhoads 1896

Subspecies Lepus (Lepus) arcticus subsp. groenlandicus Rhoads 1896

Subspecies Lepus (Lepus) arcticus subsp. monstrabilis Nelson 1934

Species Lepus brachyurus Temminck 1845

Subspecies Lepus brachyurus subsp. brachyurus Temminck 1845

Subspecies Lepus brachyurus subsp. angustidens Hollister 1912

Subspecies Lepus brachyurus subsp. lyoni Kishida 1937

Subspecies Lepus brachyurus subsp. okiensis Thomas 1906

Species Lepus californicus Gray 1837

Subspecies Lepus californicus subsp. californicus Gray 1837

Subspecies Lepus californicus subsp. deserticola Mearns 1896

Subspecies Lepus californicus subsp. insularis Bryant 1891

Subspecies Lepus californicus subsp. magdalenae Nelson 1907

Subspecies Lepus californicus subsp. melanotis Mearns 1890

Subspecies Lepus californicus subsp. texianus Waterhouse 1848

Species Lepus callotis Wagler 1830

Subspecies Lepus callotis subsp. callotis Wagler 1830

Subspecies Lepus callotis subsp. gaillardi Mearns 1896

Species Lepus capensis Linnaeus 1758

Subspecies Lepus capensis subsp. capensis Linnaeus 1758

Subspecies Lepus capensis subsp. aquilo Thomas and Wroughton 1907

Subspecies Lepus capensis subsp. carpi Lundholm 1955

Subspecies Lepus capensis subsp. granti Thomas and Schwann 1904

Subspecies Lepus capensis subsp. aegyptius Desmarest 1822

Subspecies Lepus capensis subsp. hawkeri Thomas 1901

Subspecies Lepus capensis subsp. isabellinus Cretzschmar 1826

Subspecies Lepus capensis subsp. sinaiticus Ehrenberg 1833

Subspecies Lepus capensis subsp. arabicus Ehrenberg 1833

Subspecies Lepus capensis subsp. schlumbergeri Remy-St. Loup 1894

Subspecies Lepus capensis subsp. atlanticus de Winton 1898

Subspecies Lepus capensis subsp. whitakeri Thomas 1902

Species Lepus (Eulagos) castroviejoi Palacios 1976

Species Lepus (Eulagos) comus Allen 1927

Species Lepus (Eulagos) coreanus Thomas 1892

Species Lepus corsicanus de Winton 1898

Species Lepus (Eulagos) europaeus Pallas 1778

Subspecies Lepus (Eulagos) europaeus subsp. europaeus Pallas 1778

Subspecies Lepus (Eulagos) europaeus subsp. caspicus Hemprich and Erhenberg 1832

Subspecies Lepus (Eulagos) europaeus subsp. connori Robinson 1918

Subspecies Lepus (Eulagos) europaeus subsp. creticus Barrett-Hamilton 1903

Subspecies Lepus (Eulagos) europaeus subsp. cyprius Barrett-Hamilton 1903

Subspecies Lepus (Eulagos) europaeus subsp. cyrensis Satunin 1905

Subspecies Lepus (Eulagos) europaeus subsp. hybridus Desmarest 1822

Subspecies Lepus (Eulagos) europaeus subsp. judeae Gray 1867

Subspecies Lepus (Eulagos) europaeus subsp. karpathorum Hilzheimer 1906

Subspecies Lepus (Eulagos) europaeus subsp. medius Nilsson 1820

Subspecies Lepus (Eulagos) europaeus subsp. occidentalis de Winton 1898

Subspecies Lepus (Eulagos) europaeus subsp. parnassius Miller 1903

Subspecies Lepus (Eulagos) europaeus subsp. ponticus Ognev 1929

Subspecies Lepus (Eulagos) europaeus subsp. rhodius Festa 1914

Subspecies Lepus (Eulagos) europaeus subsp. syriacus Hemprich and Ehrenberg 1832

Subspecies Lepus (Eulagos) europaeus subsp. transsylvanicus Matschie 1901

Species Lepus (Sabanalagus) fagani Thomas 1902

Species Lepus flavigularis Wagner 1844

Species Lepus granatensis Rosenhauer 1856

Subspecies Lepus granatensis subsp. granatensis Rosenhauer 1856

Subspecies Lepus granatensis subsp. gallaecius Miller 1907

Subspecies Lepus granatensis subsp. solisi Palacios and Fernández 1992

Species Lepus habessinicus Hemprich and Ehrenberg 1832

Species Lepus (Indolagus) hainanus Swinhoe 1870

Species Lepus (Proeulagus) insularis W. Bryant 1891

Species Lepus (Eulagos) mandshuricus Radde 1861

Species Lepus (Sabanalagus) microtis Heuglin 1865

Subspecies Lepus microtis subsp. microtis Heuglin 1865

Subspecies Lepus microtis subsp. angolensis Thomas 1904

Subspecies Lepus microtis subsp. senegalensis Rochebrune 1883

Subspecies Lepus microtis subsp. whytei Thomas 1894

Species Lepus (Indolagus) nigricollis F. Cuvier 1823

Subspecies Lepus (Indolagus) nigricollis subsp. nigricollis F. Cuvier 1823

Subspecies Lepus (Indolagus) nigricollis subsp. aryabertensis Hodgson 1844

Subspecies Lepus (Indolagus) nigricollis subsp. dayanus Blanford 1874

Subspecies Lepus (Indolagus) nigricollis subsp. ruficaudatus Geoffroy 1826

Subspecies Lepus (Indolagus) nigricollis subsp. sadiya Kloss 1918

Subspecies Lepus (Indolagus) nigricollis subsp. simcoxi Wroughton 1912

Subspecies Lepus (Indolagus) nigricollis subsp. singhala Wroughton 1915

Species Lepus oiostolus Hodgson 1840

Subspecies Lepus oiostolus subsp. oiostolus Hodgson 1840

Subspecies Lepus oiostolus subsp. hypsibius Blanford 1875

Subspecies Lepus oiostolus subsp. pallipes Hodgson 1842

Subspecies Lepus oiostolus subsp. przewalskii Satunin 1907

Species Lepus (Lepus) othus Merriam 1900

Subspecies Lepus (Lepus) othus subsp. othus Merriam 1900

Species Lepus (Indolagus) peguensis Blyth 1855

Subspecies Lepus (Indolagus) peguensis subsp. peguensis Blyth 1855

Subspecies Lepus (Indolagus) peguensis subsp. vassali Thomas 1906

Species Lepus saxatilis F. Cuvier 1823

Subspecies Lepus saxatilis subsp. saxatilis F. Cuvier 1823

Subspecies Lepus saxatilis subsp. subrufus Roberts 1913

Species Lepus (Sinolagus) sinensis Gray 1832

Subspecies Lepus (Sinolagus) sinensis subsp. sinensis Gray 1832

Subspecies Lepus (Sinolagus) sinensis subsp. formosus Thomas 1908

Subspecies Lepus (Sinolagus) sinensis subsp. yuenshanensis Shih 1930

Species Lepus (Eulagos) starcki Petter 1963

Species Lepus (Proeulagus) tibetanus Waterhouse 1841

Subspecies Lepus (Proeulagus) tibetanus subsp. tibetanus Waterhouse 1841

Subspecies Lepus (Proeulagus) tibetanus subsp. centrasiaticus Satunin 1907

Subspecies Lepus (Proeulagus) tibetanus subsp. craspedotis Blanford 1875

Subspecies Lepus (Proeulagus) tibetanus subsp. pamirensis Günther 1875

Subspecies Lepus (Proeulagus) tibetanus subsp. stoliczkanus Blanford 1875

Species Lepus (Lepus) timidus Linnaeus 1758

Subspecies Lepus (Lepus) timidus subsp. timidus Linnaeus 1758

Subspecies Lepus (Lepus) timidus subsp. ainu Barrett-Hamilton 1900

Subspecies Lepus (Lepus) timidus subsp. begitschevi Koljuschev 1936

Subspecies Lepus (Lepus) timidus subsp. gichiganus J. Allen 1903

Subspecies Lepus (Lepus) timidus subsp. hibernicus Bell 1837

Subspecies Lepus (Lepus) timidus subsp. kamtschaticus Dybowski 1922

Subspecies Lepus (Lepus) timidus subsp. kolymensis Ognev 1923

Subspecies Lepus (Lepus) timidus subsp. kozhevnikovi Ognev 1929

Subspecies Lepus (Lepus) timidus subsp. lugubris Kastschenko 1899

Subspecies Lepus (Lepus) timidus subsp. mordeni Goodwin 1933

Subspecies Lepus (Lepus) timidus subsp. orii Kuroda 1928

Subspecies Lepus (Lepus) timidus subsp. scoticus Hilzheimer 1906

Subspecies Lepus (Lepus) timidus subsp. sibiricorum Johanssen 1923

Subspecies Lepus (Lepus) timidus subsp. transbaicalicus Ognev 1929

Subspecies Lepus (Lepus) timidus subsp. varronis Miller 1901

Species Lepus (Proeulagus) tolai Pallas 1778

Subspecies Lepus (Proeulagus) tolai subsp. tolai Pallas 1778

Subspecies Lepus (Proeulagus) tolai subsp. aurigineus Hollister 1912

Subspecies Lepus (Proeulagus) tolai subsp. buchariensis Ognev 1922

Subspecies Lepus (Proeulagus) tolai subsp. cheybani Baloutch 1978

Subspecies Lepus (Proeulagus) tolai subsp. cinnamomeus Shamel 1940

Subspecies Lepus (Proeulagus) tolai subsp. filchneri Matschie 1907

Subspecies Lepus (Proeulagus) tolai subsp. lehmanni Severtsov 1873

Subspecies Lepus (Proeulagus) tolai subsp. swinhoei Thomas 1894

Species Lepus townsendii Bachman 1839

Subspecies Lepus townsendii subsp. townsendii Bachman 1839

Subspecies Lepus townsendii subsp. campanius Hollister 1915

Species Lepus yarkandensis Günther 1875

Discussion: Formerly included Bunolagus ; see Petter (1972 b); and originally all other genera ( Brachylagus , Caprolagus , Macrotolagus , Poelagus , Pronolagus , Romerolagus , Sylvilagus ) in Leporidae except Pentalagus . The taxonomy of this genus remains controversial. L. crawshayi (including whytei), peguensis , ruficaudatus , and siamensis have been variously treated as separate species or have been included in nigricollis . L. europaeus , corsicanus , granatensis , mediterraneus , tolai , and tibetanus have been placed in capensis or treated as distinct species; see comments therein. For synonyms see ( Pavlinov et al., 1995b).