Helina naivashensis Emden, 1951 ,,

Couri, Márcia S. & Pont, Adrian C., 2018, The male terminalia of some African species of Helina Robineau-Desvoidy, 1830 (Diptera, Muscidae), Zootaxa 4399 (2), pp. 233-247 : 240

Helina naivashensis Emden, 1951, stat. nov.

( Figs 32–34 )

Examined type material: Holotype male seen; right wing damaged at tip. One paratype from Kenya (Naivasha) dissected and illustrated.

Diagnosis. Fore tarsus with claws and pulvilli very long in male, and apical tarsomere dilated at apex; hind femur with numerous very long, setulose anteroventral and posteroventral hairs; hind tibia with long anteroventral and ventral hairs, one long posterior, and long erect posterior to posteroventral hairs on apical third.

Male terminalia. Sternite 5 longer than wide, with medium-sized and long setae on disc and longer setae at tips of lobes ( Fig. 32 ); cercal plate long, with 3 pairs of short stout setae at apex and two pairs a little above them; surstylus longer than cercal plate, sinuous in apical portion ( Fig. 34 ); aedeagal complex with phallapodeme long; distiphallus short and membranous ( Fig. 33 ).

Notes. This species was originally described as a subspecies of Helina quadruplex ( Stein, 1913). The terminalia of the two subspecies were dissected and, although their external morphology is similar, they differ in the arrangement of setae on the cercal plate, the enlargement of the hypandrium in lateral view, and the length of the distiphallus. According to Emden (1951: 597), the females of both taxa are indistinguishable, but the males differ considerably in the chaetotaxy of the hind leg ( Emden 1951: 597). The status of Helina quadruplex naivashensis is therefore revised and the subspecies is elevated to species rank (stat. nov.).

Figures

FIGURES 28–34. Helina mollis (Stein, 1906), male: (28) sternite 5, dorsal view; (29) cercal plate and surstylus, dorsal view; (30) cercal plate and surstylus, lateral view; (31) aedeagal complex, lateral view. Helina naivashensis Emden, 1951, stat. nov., paratype male:(32) sternite 5, dorsalview; (33) cercal plate, surstylus and aedeagal complex, lateral view; (34) cercal plate and surstylus, dorsal view; Scale bar 0.1 mm.