Clivina antoni, Balkenohl, 2018

Balkenohl, Michael, 2018, Notes on Oriental Clivinini and the description of two new species (Coleoptera: Carabidae: Clivinini), Linzer biologische Beiträge 50 (1), pp. 197-215 : 199-202

Clivina antoni nov.sp. ( Figs 1, 2 )

T y p e m a t e r i a l: Holotype: ♂, with labels and data: white, printed "N-Laos, Khan River, at light 5km E Luang Prabang leg. K.W. Anton, 21.IV.1999 " (CBB). Paratypes: 2♂♂: same data as holotype (CBB).

E t y m o l o g y: The species is dedicated to Klaus Werner Anton (Emmendingen near Freiburg, Germany), well known specialist in Bruchidae , and is expressed as Latinised adjective.

D i a g n o s i s: A medium sized brown Clivina species with conspicuous subcylindrical habitus. Distinguished from the most similar species C. rugosofemoralis BALKENOHL by the convex frons with rough irregular transverse rugae and the raised clypeal field separated from frons by a transverse furrow. Moreover, the antennae are elongated and reaching over the base of the pronotum. The pronotum exhibits a straight reflexed lateral margin. On the elytron the intervals three and four are wider at base and raised with a knob-like carina at interval three. Interval three shows five setigerous punctures and interval eight is carinate in its whole length. In addition both parameres possess two setae at apex.

D e s c r i p t i o n:

Measurements: Length 5.11 / 5.10 / 5.18 mm; width 1.29 / 1.28 / 1.33 mm; ratio length/width of pronotum 1.11 / 1.13 / 1.12; ratio length/width of elytra 2.14 / 2.22 / 2.16.

Colour: Dorsal and ventral surface glossy; fuscus; apical half of mandibles fuscous, other mouthparts and tarsalia leoninus, antennae and front legs fuscous, intermediate, and hind legs fuscus.

Head ( Fig. 1 ): One fourth smaller than pronotum. Clypeus relatively wide, nearly straight thought slightly emarginated anteriorly, with indistinct tooth at each side, wings of moderate size, separated from and not as far projecting as middle part of clypeus; clypeus, wings, and supraantennal plates indistinctly margined. Clypeal field slightly elevated, separated from frons by transverse furrow. Supraantennal plates vaulted, not as far projecting laterally as eyes, nearly smooth, with obtuse rounded angle anteriorly, separated from wings by distinct notches, with two supraorbital setae at each side situated at mid-eye and posterior eye-level. Supraorbital furrows deep, wide, diverging anterior clypeal foveae in which the clypeal setae are situated, diverging posteriorly and running beyond hind-genae level. Supraorbital carinae small, tubercle-like. Neck constriction indistinct, formed by short longitudinal flat rugae. Frons flattened, covered with rough irregular to transverse rugae. Eyes projecting laterally. Genae distinct. Antenna elongated, reaching over base of pronotum, segments four to ten somewhat flattened, scapus with longitudinal reticulation, with the seta situated dorsofrontally at apical quarter. Labrum not as wide as middle part of clypeus, amargined, straight, seven-setose, with transverse reticulation. Mandible slightly fractuate basally, lateral margin bent dorsally towards base, with few fine longitudinal carinae in basal half. The two apical segments of maxillary palpus securiform, tip elongated, bent laterally; segment two of labial palpus bisetose, apical segment guttuliform. Ligula wide, ovate, the two apical setae separated.

Pronotum ( Fig. 1 ): Slightly elongated, indistinctly convex on disc (lateral view), distinctly convex in frontal view. Anterior angles indistinctly pointed, rounded-off. Posterior angles small, traceable in dorsal view at 80 times, indistinctly visible as rounded knob in lateral view. Reflexed lateral margin straight and indistinctly converging in anterior half, convex directly before anterior angles, slightly convex to posterior setigerous puncture.

Marginal channel wide between setigerous punctures, with row of moderately sized foveae. Posterior setigerous puncture situated in distinct fovea, removed from lateral channel by diameter of pore. Reflexed margin thin in whole length, running from posterior angles to base as slightly sigmoid line. Pleura somewhat swollen, just not visible in dorsal view. Anterior transverse line deep, reaching anterior margin, not joining lateral margin, interrupted at middle. Median line deep, complete, surpassing anterior transverse line without fusing, narrow towards base. Surface smooth on disc, basal impression formed by longitudinal group of deep foveae, with some short broad transverse rugae at base and laterally.

Elytra ( Fig. 1 ): Subcylindric, flattened in lateral view, distinctly convex in frontal view. Lateral margin nearly straight, contracted to humerus and in apical third. Humerus rounded obtuse-angularly. Base concave at declivity, intervals at declivity somewhat overhanging over pedunculus, distinctly margined from humerus to peduncle. Setigerous tubercle at base of first stria. Scutellar stria absent. Striae punctuate-striate, one to three free at base, four to seven joining at base, one and seven, three and four, five and sex joining apically. Intervals convex, eights distinctly carinate in whole length, seventh with short carina at humerus and apex, third and fourth at base wider and distinctly raised, third at base with knob-like carina. Marginal channel with uninterrupted series of setigerous punctures, another series of setigerous punctures situated mesially. Third interval with five setigerous punctures, all approaching third stria. Base with small band of isodiametric reticulation. Reflexed lateral margin crenulate, more distinct in basal half.

Hind wings: Fully developed.

Ventral surface: Proepisternum nearly smooth at middle, laterally covered with distinct isodiametric reticulation and some transverse wrinkles, submarginal furrow complete. Epipleuron with distinct isodiametric reticulation. Abdomen laterally with distinct isodiametric reticulation, smooth at middle but with irregular situated punctures at all tergites, punctures of terminal segment bigger, the two apical setigerous punctures widely separated.

Legs: Profemora with few and slight longitudinal rugae ventrally, protibia with distinct and complete carina and sulcus dorsally, terminal spine wide, ensiform, turned distinctly ventrally and laterally; movable spur as long as spine, sharp, turned slightly ventrally, three praeapical lateral denticles wide, ensiform, setae at all denticles fractuate. First segment of protarsus as long as segments two to four together. Intermediate tibia with distinct spur apically, furnished with arcuate seta; all terminal tarsomeres with long claws.

Male genitalia ( Fig. 2 ): Median lobe slender, slightly arcuate at middle, distinctly arcuate apically. Endophallus strongly folded, with some minute knobs laterally. Both parameres slender, arcuate, distorted, with two setae of different length at apex.

Female genitalia: Unknown.

Variation: Variation was observed in the structure of the rough transverse rugae on the frons of the head. The transverse furrow, separating the clypeal field from the frons is more or less interrupted at middle.

D i s t r i b u t i o n: Known from the type locality 5 km East of Luang Prabang.

Habits: According to the collector, the light trap was positioned near the river with partially open view to the muddy bank. N 19°53’03’’, E 102°11’13’’, altitude 300 m. The specimens were collected together with Clivina mekongensis LESNE.

R e l a t i o n s h i p s: Possible relationships have been already pointed out in the description of Clivina rugosofemoralis BALKENOHL ( BALKENOHL 1999). In that contribution it was emphasized the species might represent a member of a separate new group among the genus Clivina not jet characterised especially. C. antoni nov.sp. is another example for this hypothesis.

Figures

Figs 1-2: Clivina antoni nov.sp.: (1) Holotype, habitus; (2) holotype, aedeagus and parameres, dorsal view.