Labiobaetis moriharai ( Mueller-Liebenau , 1984)

Kaltenbach, Thomas & Gattolliat, Jean-Luc, 2020, Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae), ZooKeys 914, pp. 43-79 : 43

Labiobaetis moriharai ( Mueller-Liebenau, 1984) Figures 9 , 11c , 15a

Diagnosis.

Larva. Following combination of characters: A) dorsal surface of labrum with submarginal arc of 1 + 8-10 simple setae, the first three after central seta longer than others and decreasing in length; B) labial palp segment II with a large, lobed distomedial protuberance, segment III conical, apically slightly truncate; C) fore femur rather broad, length 3.4 × maximum width, dorsal margin with a row of ca. 10 curved, spine-like setae; D) six pairs of gills; E) hindwing pads present, minute; F) scape with well-developed distolateral process ( Müller-Liebenau 1984a: fig. 10f); G) paraproct distally not expanded, with ca. 12 stout marginal spines ( Müller-Liebenau 1984a: fig. 10l).

Distribution.

Malaysia: Selangor, Sabah; Vietnam; Brunei (Fig. 15a ).

Biological aspects.

The specimens were collected at altitudes from 100 m to 300 m, partly on bottom gravel, rock surface or vegetation in stream run or riffles.

Examined material.

Paratype. 1 larva (on slide, no. 41), W. Malaysia, Trib. of Gombak River, 16 ½ miles N of Kuala Lumpur, 14.XI.[19]68, Coll. Bishop. Other material. 1 larva (on slide, GBIFCH 00658106), Brunei, Temburong District, Ulu Temburong National Park, Belalong River (near field station), 04°32.82'N, 115°09.50'E, 100 m, V. 2014, K. Baker leg.; 1 larva (on slide, GBIFCH 00592243), Brunei, Temburong District, Ulu Temborong National Park, Belalong River tributary, 04°32.63'N, 115°08.85'E, 170 m, V. 2014, K. Baker leg.; 5 larvae (2 on slides, GBIFCH 00592241, GBIFCH 00658112, 3 in alcohol, GBIFCH 00515325), Malaysia, Sabah, Tawau River, primary forest, 04°24.23'N, 117°53.58'E, 280 m, 12.III.2008, Mendoza leg. All material deposited in MZL, except paratype in Zoologische Staatssammlung München (ZSM).

Figures

Figure 9. Labiobaetis moriharai, larva morphology: a Labrum b Left mandible c Maxilla d Labium e Foreleg f Metanotum.

Figure 11. Habitus, larvae, dorsal view: a Labiobaetis dayakorum sp. nov. b Labiobaetis paraoperosus c Labiobaetis moriharai.

Figure 15. Distribution of Labiobaetis in Borneo: a Labiobaetis moriharai b Labiobaetis borneoensis c Labiobaetis bakerae sp. nov. and Labiobaetis dayakorum sp. nov. d Labiobaetis penan sp. nov. T: type locality.