Misgolas yorkmainae, Wishart & Rowell, 2008

Wishart, Graham & Rowell, David M., 2008, Trapdoor Spiders of the Genus Misgolas (Mygalomorphae: Idiopidae) from Eastern New South Wales, With Notes on Genetic Variation, Records of the Australian Museum 60 (1), pp. 45-86 : 57-58

Misgolas yorkmainae n.sp.

Figs 7A–J , 24A

Type material. HOLOTYPE?, AM KS38616 , Kerewong State Forest , nr Lorne, (31°36'S 152°34'E), 15 July 1979, pit fall trap field No. 1600, D. Milledge . PARATYPES ??: AM KS1553 , Kerewong SF, nr Lorne, site 108, Kendall Forest Management Area, subtropical rainforest (31°36'S 152°34'E), 28 May 1978, pit fall trap field No. 1167, D. Milledge . AM KS5419 , Kerewong SF, nr Lorne, site 108(2), (31°36'S 152°34'E), 15 July 1979, pit fall trap field No. 1597, D. Milledge . AM KS50020 , Lorne SF, nr Lorne, site 86(3), 31°35'S 152°37'E, 15 July 1979, pit fall trap field No. 1593, D. Milledge . AM KS86230 , all details as for paratype AM KS5419 . All type specimens brittle & easily damaged, possibly first preserved in formalin .

Selection of other material examined. ??, AM KS38615 , Bulga State Forest , Bobbin Fire Trail, 100 m from Padmans Road (31°37'S 152°10'E), 620 m, 4 Feb.–9 Apr. 1993, pit fall trap site 57BM, M. Gray, G. Cassis . AM KS48653 , data as for AM KS38615 . AM KS38657 , Mt. Boss State Forest , 17 1040, North Plateau Rd. at Plateau Beech Picnic Area. Small gully near picnic area end of Plateau Rd. (31°10'S

152°19'E), 4 Feb.–9 Apr. 1993, pit fall trap site 32BG, M. Gray, G. Cassis. AM KS38661 , Mt. Boss State Forest , 17 1120, North Plateau Rd. about 1.5 km by track from Plateau Beech Picnic Area (31°10'S 152°19'E), 4 Feb.–9 Apr. 1993, pit fall trap site 32BM, M. Gray, G. Cassis .

Diagnosis. In male: large dark tan spiders, carapace length c. 8.3–9.1; retrodorsal surface of metatarsi IV with at least 2 spines ( Fig. 7I ); venter entirely pallid ( Fig. 7J ). Bilateral rows of small spines adjacent to scopula on tarsi I and II; bilateral small clusters of spines near distal edges metatarsi I and II ( Fig. 7E–H ). Palpal bulb ( Fig. 7B,C ) embolic flange with c. 10 minute folds; embolus with prolateral bend near tip, apophysis absent. Retrolateral tibial apophysis abruptly swollen midway, distal tibial apophysis massive, cymbium spines strong, attenuate, sub erect ( Fig. 7A,D ). Female not known.

Description

Male holotype ( Fig. 7A–J ). Size. Carapace length 8.82, width 7.35. Abdomen length 7.64, width 5.00. Colour. In alcohol limbs and carapace dark tan; dark brown smudges absent from lateral limb surfaces. Abdomen dorsum dark brown with 6 pallid transverse chevrons. Sparse fine pallid hairs most concentrated on carapace near perimeter & extending onto coxae.Venter entirely pallid. Carapace. Edge fringed with many long black bristles which extend onto a raised post foveal surface. Line of c. 9 (some rubbed off) median bristles on caput arch. Group of 6 bristles between PME, 7 bristles on clypeus. Area below clypeus on pleuron membrane not chitinous, without setae. Fovea width 1.43, deep, slightly procurved, edges recurved; posterior wall centrally divided by two pitted intrusions. Eyes. Raised on mound. Area immediately adjacent to eyes black; anterior width 1.56, posterior width 1.48, length 0.96, width/length ratio 1.63. Line joining posterior edges of ALE transects anterior third of AME. Posterior row recurved in front and behind. Chelicerae. Rastellum row of 6(6) strong spines, few smaller spines retreat along pd edges. Intercheliceral tumescence not examined. Fang groove with 9(8) promarginal teeth & 12(12) smaller retromarginal/intermediate row teeth. Labium. Bulbous, raised, length 0.87, width 1.05. Labio-sternal suture broad, medially divided. Maxillae c. 4–5 (4–5) anteroental basally thickened setae in lieu of developed cuspules. Sternum. Length 4.97, width 3.75. Sigilla, anterior pairs small, round, posterior pair larger, ovate, all their own diameter from margin. Legs. Tibia I with distal bifid apophysis; both processes “stump-like” with 2(2) short blunt spines.

palp I II III IV femur 4.78 8.53 8.04 6.88 8.63 patella 2.21 4.41 4.11 3.32 4.11 tibia 4.35 6.276.14 4.35 7.84 metatarsus — 6.96 6.32 5.89 8.13 tarsus 2.03 3.52 3.44 3.44 4.05 total 13.37 29.69 28.05 23.88 32.76

Palp ( Fig. 7A,D ). Cymbium with 37(33) strong, attenuate, sub-erect spines covering anterior half of d surface. RTA abruptly swollen midway, d and rd surfaces covered with short pointed spines; rv edge of tibial excavation and straight massive DTA suspends brush of longer spines. TEM prominent, pallid, adjacent to RTA; TET not seen. Bulb. ( Fig. 7B,C ). Embolic rl flange with c. 10 minute folds, margin distally sharply convex curved under embolus as if twisted; embolus with pl bend near tip, apophysis absent. Scopula. Complete on tarsi I and II; small remnants on metatarsi I and II; absent on legs III and IV. Trichobothria. Palp: tarsi 11, tibia pd 6, rd 6. Leg I: tarsi 13, metatarsi 12, tibia pd 6, rd 6. Leg II: tarsi 12, metatarsi 11, tibia pd 7, rd 7. Leg III: tarsi 12, metatarsi 12, tibia pd 7, rd 7. Leg IV: tarsi 12, metatarsi 13, tibia pd 9, rd 8. Leg spination. Leg I ( Fig. 7E,F ): tarsi bilateral rows small spines adjacent to scopula pv 12, rv 16; metatarsi apical pv cluster of 12, rv band of 18; tibia pv 011223. Leg II ( Fig. 7G,H ): tarsi bilateral rows small spines adjacent to scopula pv 16, rv 23; metatarsi pv 011cluster of 9, rv band of 11 including apical cluster of 8, pd 010; tibia v 0111112. Leg III: tarsi v 62; metatarsi v 21, pd 011110, rd 010110; tibia pl 0110, rl 0110; patella pd 2. Leg IV ( Fig. 7I ): tarsi v 48; metatarsi v 25, rd 001010; tibia v 8, rl 01110. Abdomen. Dorsum covered with long brown bristles and underlying fine pallid hairs. Venter covered by short brown bristles.

Etymology. The species is named in recognition of Dr Barbara York Main, doyenne of Australian trapdoor spiders.

Distribution and natural history. ( Fig. 24A ) This species is known only from the type locality, the Kerewong State Forest, near the town of Lorne, N.S.W. The burrow is unknown.

Figures

Fig. 7. Misgolas yorkmainae n.sp. A–J?, holotype AM KS38616. (A), right palp retrolateral. (B,C), right bulb: B, dorsal; C, prolateral. (D), left cymbium dorsal. (E,F), right tarsus and metatarsus I: E, prolateral; F, retrolateral. (G,H), right tarsus and metatarsus II: G, prolateral; H, retrolateral. (I), right tarsus and metatarsus IV retrolateral. (J), venter.

Fig. 24. A–D. Distribution of Misgolas species in the New South Wales Central Coast (eastern Australia) based on material examined (Inset [Fig. 24A] with catchment area).