Naviauxella varians Wiesner and Constant, 2019

Wiesner, Jürgen & Constant, Jérôme, 2019, Records of tiger beetles (Coleoptera: Cicindelidae) collected in Cambodia, with description of a new species. 149. Contribution towards the knowledge of the Cicindelidae, Insecta Mundi 700 (700), pp. 1-12 : 6

Naviauxella varians Wiesner and Constant , new species

( Fig. 13–21 )

Type depository. Holotype male, four paratype males and five paratype females in RBINS, one para- type male and one paratype female in RUPP-CEI, three paratype males and two paratype females in coll. Wiesner.

Type status. Holotype male, type labels: “Coll. I. R.Sc.N.B. / Cambodia, Kampong Speu prov. / Phnom Aural W.S., nr Srae Ken vill. / 10-14.v.2018, 200– 400m / 11°59′N 104°08′E / GTI Project, leg. J. Constant / & P. Limbourg, I.G.: 33.732 [printed, yellow]», « HOLOTYPE / Naviauxella / varians / Wiesner & Constant ded. 2019 [printed, red]”.

Paratypes: 8 males and 8 females with the same label and “ PARATYPE / Naviauxella / varians / Wiesner & Constant ded. 2019 [printed, red]”.

Distribution. Cambodia (Kampong Speu).

Etymology. The species name varians refers to the high variability of elytral maculation.

Diagnosis. Using the key of Matalin (2018b: 297), N. varians n. sp. falls with the Thai N. clivitatis Naviaux, 1991 , from which it is distinguished by the large apical dot of the elytral maculation and the shape of apical tip of the aedeagus, which is bent ventrally and not evenly rounded.

Description. Size: Total length (without labrum) males 6.4 – 6.9 mm (mean = 6.7mm, n = 9), females 7.0 – 7.7 mm (mean = 7.3 mm, n = 8). Head: Dark coppery above, clypeus and genae bluish green; surface smooth, glabrous, with two setigerous punctures next to each eye in front and at centre; strong longitudinal striae on vertex and orbital plates, striae becoming slightly concentric on vertex and irregular, transversely waved, behind on neck; genae roughly longitudinally striated. Eyes pale yellow. Ratio of width of head to elytra, mean = 1.0. Labrum of males ( Fig. 15 ) light brown with darker lateral margin, distinctly wider than long, ratio of width to length, mean = 1.8; with four marginal setae and three apical teeth, middle tooth reduced and obtuse-angled, outer teeth protruding and acute-angled; with a wide, low, central carina. Labrum of females ( Fig. 16 ) dark brown, slightly wider than long, ratio of width to length, mean = 1.1; with four marginal setae and three protruding, acute-angled apical teeth, middle tooth longer than outer teeth; with a wide, low, central carina. Mandibles yellowish, darkened on apical and inner teeth in males and completely darkened except at base in females. Labial and maxillary palpi light brown, last palpomere apically metallic dark greenish in males, completely dark greenish in females; basal labial palpomere with long, erect white setae. Antennae slender, long, reaching one-third of elytral length in males, a little shorter in females; antennomeres one to four brownish in males, darker in females, with metallic luster; scape with one long apical seta, following three antennomeres almost glabrous; antennomeres five to eleven dark blackish brown, dull, finely and evenly pubescent. Thorax: Pronotum as wide as long (ratio of width to length, mean = 1.0), sides somewhat rounded between transverse constrictions; glabrous, transversely waved before and behind transverse constrictions and at lateral margin, with irregular wrinkles on disc, v-shaped beside shallow middle furrow; color dark brown, transverse constrictions and lateral margins bluish green. Sterna and episterna metallic bluish black, pro- and mesothoracic coxae with some white setae, several white setae on lateral margin of metathoracic coxae and on lateral and basal margins of metasternum, scattered white setae on metepisternum, otherwise glabrous. Mesepisternum of females with a longitudinal furrow in posterior half. Elytra: ( Fig. 17–21 ) twice as long as wide, ratio of width to length, mean = 1.9, parallel-sided, somewhat rounded in middle; color dark brown on disc, dark blue at lateral margin. Elytral testaceous maculation consists of a medium-sized roundish humeral dot in males only, in one case with a thin subhumeral extension, basal and apical dots connected as a middle band, or isolated, with an apical dot of same size or smaller to nearly absent, and a large subapical dot, either comma-shaped or not. Surface with fine microsculpture on disc, some short transverse furrows near suture in basal third, and much larger, evenly spaced punctures laterally; preapical impression strong, apical microserration fine; apical border distinctly rounded and restricted towards suture, with a tiny sutural spine. Epipleura light brown. Ventral aspect: Venter glabrous, black with metallic green blue luster; trochanters, femora, tibiae and tarsi light brown, distal tips of segments darker; legs covered with some white setae. Aedeagus: ( Fig. 14 ) in left lateral view fusiform, straight, slightly enlarged at middle, with a long and slender tapering apex, slightly bent ventrally (total length, mean 2.0 mm).

Figures

Figures 7–13. Cambodian tiger beetles.All scale bars = 5 mm. 7) Calochroa interruptofasciata flavolineata (Chaudoir, 1865) from Chambok. 8) Calochroa bramani (Dokhtouroff, 1882) from Chambok. 9) Calomera angulata angulata (Fabricius, 1798) from Phnom Aural. 10) Cosmodela duponti duponti (Dejean, 1826) from Yeak Laom Lake. 11) Cosmodela juxtata (Acciavatti and Pearson, 1989) from Bokor N. P. 12) Lophyra (Spilodia) lineifrons (Chaudoir, 1865) from Phnom Aural.

Figures 13–21. Naviauxella varians, new species. 13) Habitus, paratype male, scale bar = 5 mm. 14) Left lateral view of aedeagus, paratype, scale bar = 1 mm. 15–16. Labrum, scale bar = 1 mm. 15) Paratype male. 16) Paratype female. 17–21. Left elytron, scale bar = 2 mm. 17) Paratype male. 18) Paratype female. 19) Paratype male. 20) Paratype male. 21) Paratype female.