Amphineurus (Rhamphoneurus) fuscifusus Alexander, 1929a, nomen dubium

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.12. Amphineurus (Rhamphoneurus) fuscifusus Alexander, 1929a, nomen dubium

Amphineurus (Rhamphoneurus) fuscifusus Alexander, 1929a: 190 (description).

Remarks.

Amphineurus (R.) fuscifusus Alexander is the only species of the genus in which the male is unknown. Additionally, the only identified specimen is the female holotype, currently housed in the Natural History Museum (BMNH). The absence of an associated male specimen and its terminalia is problematic because this is an important source of characters and taxonomic data. Lacking a well-defined diagnosis in the description, the erection of the new species was proposed mostly due to subtle differences in coloration. The only considerable difference in coloration in A. (R.) fuscifusus is the pale halter, which is unlike the yellowish brown halter of A. (R.) nothofagetorum Alexander.

Some morphological data described by Alexander (1929a) suggest this specimen shares similar features with clade 33 (Fig. 32 ). The presence of long setae on the antenna, "weak branches" in medial veins, three stripes on the scutum, and a marking near the fork of Rs are features shared between the holotype and this group. The holotype was collected in the same location as the holotype of A. (R.) nothofagetorum Alexander. However, the absence of males added to the fact that there is only a single known specimen makes the synonymization or identity confirmation of the species problematic. For these reasons, we define Amphineurus (Rhamphoneurus) fuscifusus Alexander as nomen dubium.

Figures

Figure 32. Most parsimonious tree (part) yielded by parsimony analysis with implied weighting (K = 3), depicting recovered relationships among outgroup taxa. Numbers at tree branches designate nodes (see 4.2.). - Abbreviations: Aus, Australia; NC, New Caledonia; NG, New Guinea; NZ, New Zealand, SAm, South America; Tas, Tasmania.