Trichotichnus (s. str.) noctuabundus Habu, 1954

Kataev, Boris M., Hongbin, Liang & Wrase, David W., 2022, New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies, Zootaxa 5159 (3), pp. 301-353 : 337-338

Trichotichnus (s. str.) noctuabundus Habu, 1954

( Figs. 76 , 89–93 )

Trichotichnus noctuabundus Habu, 1954: 56

Material examined. China. ZHEJIANG: 1 female, Anji Co., Longwangshan , 1000 m, at light, 20–21.IX.2004, S. Belokobylskij leg. ( ZIN) ; 1 male, 1 female, Hangzhou Pref., Tianmu Shan , 40 km WNW Linan, water reservoir, 300 m, 30°20′56′′N 119°18′42′′E, litter from rock edges, sifted, [39], 17.VI.2007, D.W. Wrase leg. (cWR) ; 1 male, Deqing, Moganshan Station , pitfall traps, 30.60209°N 119.86108°E, 522 m, 17.VI.2004, H.-B. Liang, T. Sota leg. ( IOZ) . FUJIAN: 1 male, Ziyundongshan , NW slopes, 25°46′N 117°20′E, 900–1100 m, 13–14.VII.2007, J. Turna leg. (cWR) . HUBEI: 1 male, 1 female, W Hubei, Daba Shan , river bank 8 km SE Muyuping, 31°23′N 110°24′E, ca 1000 m, slopes with deciduous forest / leaves, 18.VII.2001, D.W. Wrase leg. (cWR) . SICHUAN: 3 males, 3 females, C Sichuan, Wenjiang Dist. , Dujiangyan Co., Qingcheng Shan , 975 m, 56 km NW Chengdu, 30°54′N 103°33′E, field ridge, 18.VI.1999, D.W. Wrase leg. (cWR, ZIN) ; 1 female, Qingcheng Shan , 65 km NW Chengdu, 8 km W Taiping, 103°33′E 30°53′N, 800–1000 m, 18.V.–4.VI.1997, A. Pütz leg. (cPTZ) ; 1 female, Emei Shan, Wannian , 800 m, 21–29.III.1999, P. Jäger leg. ( SMNS) ; 1 male, 1 female, Wenjiang Distr., Guanxian Co., 56 km NW Chengdu, Qingcheng Shan , 975 m, 30°53'84"N 103°32'80"E, 18.VI.1998, leg. A. Pütz leg. (cWR, cPTZ) . GUANGXI: 4 females, Longsheng, Huaping Nature Reserve, Hongtan , light trap, 4.X.2007, night, Liu Chunxiang leg. ( IOZ, ZIN) ; 1 male, Longsheng, Huaping Nature Reserve, Cujiang , light trap, 23.IX.–1.X.2007, Liu Chunxiang leg. ( IOZ) .

Vietnam. CAO BANG: 1 male, 40 km W of Cao Bang Phia Oac Mt. , 22°36′25′′N 105°52′08′′E, h ~ 1650–1700 m, deciduous forest, 3–11.X.2018, D. Fedorenko leg. ( SIEE) ; 3 males, 4 females, vic. Tinh Tuc, Son Do , Nui Pia Oac Nature Res., 22°37′55′′N 105°52′98′′E, 1850– 1300 m, 9–15.V.2014, by light, A. Weigel leg. ( NME, ZIN) . LAO CAI: 1 female, pass 8 km NW Sa Pa , 22°21′13′′N 103°46′01′′E, 2030 m, forest margin, 9.VIII.2013, V. Assing leg. [10+2] ( MFNB) .

Re-description. Habitus as in Fig. 76 . External morphological character states as described for T. schmidti sp. n., but differing as follows.

Body, on average, larger (length 8.1–9.5 mm). Microsculpture on head and pronotum highly obliterated, not forming distinct meshes; left mandible in frontal view moderately narrowed apically, with subtruncate apex (somewhat intermediate in this respect between that of T. schmidti sp. n. and T. subangulatus ); pronotum widest in the end of anterior third, comparatively strongly narrowed basally; its sides markedly rounded along entire length, occasionally more or less straight basally; basal angles obtuse, narrowly rounded at apices, occasionally more distinct.

Median lobe of aedeagus ( Figs. 89–91 ) more robust, with relatively large basal bulb, in lateral view rather evenly arcuate, strongly narrowed apically, with apex slightly protruding dorsally forming there a very small apical capitulum; in dorsal view median lobe comparatively wide, only slightly narrowed apically, with terminal lamella ( Fig. 89 ) wide and short, roundly narrowed posteriorly. Ventral side finely edged laterally in apical part and flattened between margins. Apical orifice in dorsal position, wide apically. Internal sac with a spiny formation apically.

Female genitalia ( Figs. 92 & 93 ): gonocoxite arcuate, comparatively narrow in ventral view, about 0.7 times as long as gonosubcoxite, with wide base.

Proportions (6 males and 5 females measured): HWmax/PWmax 0.71–0.75; HWmin/PWmax 0.54–0.59; HWmax/HWmin 1.23–1.35; PWmax/PL 1.45–1.60; PWmax/PWmin 1.20–1.36; EL/EW 1.54–1.59; EL/PL 2.93–3.22; EW/PWmax 1.26–1.33.

Distribution. Described from Japan ( Habu 1954) and recorded from China ( Taiwan: Habu 1980, Ito 2021b; Sichuan: Hua 2002; Zhejiang, Hubei, Fujian: Kataev & Wrase 2017b), South Korea (Jujudo: Paik 1995, Moon & Paik 2006) and Vietnam ( Ito 2021b). It is recorded here for the Chinese province of Guangxi for the first time; it will probably also be found in Yunnan.

Figures

FIGURES 74–76. Trichotichnus spp., habitus. 74, T. subangulatus Ito, 2021 (NNE Tengchong); 75. T.? subangulatus Ito, 2021 (S Longyuan); 76, T. noctuabundus Habu, 1954 (Vietnam). Scale bar = 1.0 mm.

FIGURES 89–93. Trichotichnus noctuabundus Habu, 1954 (Vietnam). 89, terminal lamella of aedeagus; 90 & 91, median lobe of aedeagus; 92, gonocoxite; 93, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (89 & 91), lateral (90 & 92) and ventral (93) views. Scale bars: A = 0.5 mm (89, 92 & 93), B = 1.0 mm (90 & 91).