Tosapusia neudorfensis ( Schaffer, 1898 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 57-58

Tosapusia neudorfensis ( Schaffer, 1898) nov. comb.

Figs 19A 1 –A 2 , B 1 –B 2 , C 1 –C 2 , D 1 –D 2 , 18A

* Mitra Neudorfensis nov. spec. — Schaffer 1898: 542, fig. 5.

V [exillum]. (U [romitra].) neudorfense (Schff.) — Sieber 1958a: 153.

Vexillum svagrovskyi sp. nov. — Biskupič 2020: 77, figs. 2–5.

Type material. The syntypes have been deposited in the collection of Natural History Museum Vienna ( Schaffer, 1898) but cannot be located .

Additional material. SNM-PM Z 40040, SL: 29.2 mm, MD: 9.6 mm, Konopiská clay pit at Rohožník, Slovakia, middle Miocene (Badenian), Natural History Museum of Slovak National Museum, Bratislava, Slovakia, illustrated in Biskupič (2020, figs 2A–B), holotype of Vexillum svagrovskyi Biskupič, 2020 , figs 19A 1 –A 2; SNM-PM Z 40045, SL: 40.2 mm, MD: 12.5 mm, paratype of Vexillum svagrovskyi Biskupič, 2020 , illustrated in Biskupič (2020, figs 3C–D), figs 19B 1 –B 2; SNM-PM Z 40046, SL: 30.3 mm, MD: 10.4 mm, paratype of Vexillum svagrovskyi Biskupič, 2020 , illustrated in Biskupič (2020, figs 3E–F), figs 19C 1 –C 2; SNM-PM Z 40053, illustrated in Biskupič (2020, figs 2G–H), figs 19D 1 –D 2. 7 specimens, SNM-PM Z 40041–Z 40047, all from Konopiská clay pit at Rohožník, Slovakia, middle Miocene (Badenian), Natural History Museum of Slovak National Museum, Bratislava, Slovakia, paratypes of Vexillum svagrovskyi Biskupič, 2020 .

Revised description. Shell large, moderately slender fusiform, solid, with weakly gradate spire. Protoconch unknown. Teleoconch of 11 whorls; spire whorls subcylindrical, with impressed suture. Axial sculpture of early teleoconch whorls of densely set opisthocline axial ribs, separated by narrower interspaces. Axial ribs weakening on seventh to ninth teleoconch whorl, becoming slightly more widely spaced and blurred on penultimate and last whorls. Spiral cords most prominent on early teleoconch whorls in axial interspaces, separated by narrow grooves. Broad subsutural band formed by deep spiral groove cutting axial ribs. Spiral band solid, or split into three to five densely spaced spiral cords. All spiral sculpture becoming subobsolete on penultimate and last whorls, rarely subsutural band persisting as weak spiral cord. Last whorl moderately convex, slowly contracting at base, covered by weakly sigmoidal growth lines. Several broad, flat spiral cords with delicate axial threads on weak fasciole. Aperture narrow with indistinct anal sinus; columellar callus indistinct, poorly delimited. Columella with four to five folds, weakening rapidly abapically. Outer lip thin without lirae or denticles. Siphonal canal moderately long and wide, straight, with shallow siphonal notch. [Note that this description is based on the holotype and paratypes of Vexillum svagrovskyi Biskupič, 2020 , except for the fifth columellar fold, which is only known from the syntypes of V. neudorfense ].

Shell measurements and ratios. SL: 28.5–40.2 mm, MD: 9.6–12.5 mm; AA: 30–34°, SL/MD: 3.0–3.2, AL/ AW: 4.3–5.1, AH/S: 1.6–2.2.

Discussion. This species is characterised by the presence of a distinct spiral groove a short distance below the suture that bisects the ribs, resulting in a broad subsutural band.

The syntypes of Tosapusia neudorfensis seem to be lost, but the illustration in Schaffer (1889) is rather precise. Based on that illustration, it is most likely that Vexillum svagrovskyi Biskupič, 2020 is a junior synonym of T. neudorfensis . Both taxa are only known from the upper Badenian Studienka Formation in Slovakia. Both species are identical in shape and sculpture and the only difference is the weak fifth columellar fold of Tosapusia neudorfensis . We assume that the presence of a fifth columellar fold is rather a matter of intraspecific variability.

Palaeoenvironment. Only known from the upper Badenian Studienka Formation, which represents middle to outer neritic depositional environments ( Biskupič 2020).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Devínska Nová Ves, Rohožník–Konopiská ( Slovakia) ( Schaffer 1898; Biskupič 2020).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 18 A. Tosapusia neudorfensis (Schaffer, 1898), reproduction of illustration of syntype by Schaffer (1898, fig. 5). B.

FIGURE 19 A1–A2. Tosapusia neudorfensis (Schaffer, 1898), Rohožník (Slovakia), SNM-PM Z 40040. B1–B2. Tosapusia neudorfensis (Schaffer, 1898), Rohožník (Slovakia), SNM-PM Z 40045. C1–C2. Tosapusia neudorfensis (Schaffer, 1898), Rohožník (Slovakia), SNM-PM Z 40046. D1–D2. Tosapusia neudorfensis (Schaffer, 1898), Rohožník (Slovakia), SNM-PM Z 40053. E1–E2. Tosapusia pseudocupressina (Bałuk, 1997), Möllersdorf (Austria), NHMW 2020/0091/0001. F1–F2. Tosapusia pseudocupressina (Bałuk, 1997), Baden (Austria), NHMW 2020/0090/0001. G1–G2. Tosapusia pseudocupressina (Bałuk, 1997), holotype, Baden (Austria), NHMW 1846/0037/0103. H1–H2. Tosapusia pseudocupressina (Bałuk, 1997), Lăpugiu de Sus (Romania), NHMW 1870/0033/0037. I1–I2. Vexillum transalpinum nov. sp., holotype, Steinebrunn (Austria), NHMW 1846/0037/0106. J1–J2. Vexillum transalpinum nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0094/0001. K1–K2. Vexillum transalpinum nov. sp., paratype, Niederleis (Austria), NHMW 2020/0096/0001. L1–L2. Vexillum transalpinum nov. sp., paratype, Steinebrunn (Austria), NHMW 2020/0095/0001.