Pusia? confunda, 1840

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 38-39

Pusia? confunda nov. sp.

Figs 4I , 13H 1 –H 2

Mitra (Costellaria) pseudorecticosta n. nom.— Boettger 1906 (pars): 10 [non Boettger, 1906].

Type material. Holotype: SMF 360341 (= XII.2211 a), Senckenberg Museum , Frankfurt / Main, Germany, SL: 5.75 mm, MD: 2.8 mm, figs 4I, 13H 1 –H 2.

Type locality. Coşteiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, Badenian (Langhian).

Etymology. From confundere (Latin: to mix), referring to the confusion with Pusia pseudorecticosta from the Vienna Basin.

Diagnosis. Pusia species of medium size, stout fusiform profile, with slightly cyrtoconoid spire, and prominent, wide spaced, opisthocline axial ribs.

Description. Shell medium-sized, broad fusiform with slightly cyrtoconoid spire. Protoconch (fig. 4I) multispiral, tall conical of 3.5 smooth convex (height: 580 μm, diameter: 530 μm). Teleoconch of five whorls; first teleoconch whorl low, convex, with periphery close to abapical suture, later whorls increasing in height, moderately convex, with periphery mid-whorl. Suture distinctly incised, weakly undulating along axials ribs. Axial sculpture of very prominent, opisthocline, convex ribs, separated by slightly wider, concave interspaces (14 axial ribs on penultimate whorl). Last whorl convex along periphery, weakly convex above, rapidly contracting below; axial ribs on last whorl weakly sigmoidal, persisting on base. Aperture wide, ovoid with indistinct columellar callus. Columella with four prominent folds, weakening rapidly abapically. Outer lip edge damaged, with seven prominent lirae within. Fasciole with four strong, rounded spiral cords. Siphonal canal short, narrow, straight, with shallow siphonal notch.

Shell measurements and ratios. SL: 5.75 mm, MD: 2.8 mm; AA: 46°, AA: 45–52°, SL/MD: 2.4, AL/AW: 3.3, AH/S: 3.2.

Discussion. This species is reminiscent of Pusia falsitranssylvanica nov. nom. in sculpture, but differs clearly in its much broader profile, its slightly cyrtoconoid and lower spire, and the coarse lirae. Boettger (1906) confused this species with Pusia pseudorecticosta ( Boettger, 1906) from Steinebrunn ( Austria) (which was established by him in the same paragraph). These species, however, have little in common and the small size, stout shape, cyrtoconoid spire, blunt axial ribs of Pusia confunda are in sharp contrast with Pusia pseudorecticosta .

Palaeoenvironment. The assemblages from Coşteiu de Sus suggest middle neritic palaeoenvironments with soft bottom (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1906).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes & Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes & Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037.

FIGURE 13 A1–A2. Pusia avellanella (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360343. B1–B2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0002. C1–C2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. D1–D2. Pusia avellanella (Boettger, 1906), Coşteiu de Sus (Romania), SMF 360344. E1–E2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0004. F1–F2. Pusia avellanella (Boettger, 1906), Steinebrunn (Austria), NHMW 1847/0037/0048. G1–G2. Pusia falsitranssylvanica nov. nom., holotype, Coşteiu de Sus (Roma- nia), SMF 360346. H1–H2. Pusia confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. I1–I2. Pusia moravica (Hoernes & Auinger, 1880), lectotype, Rudice (Czech Republic), NHMW 1862/0029/0006. J. Pusia moravica (Hoernes & Auinger, 1880), paralectotype, Rudice (Czech Republic), NHMW 2020/0097/0001.