Pusia schafferi ( Meznerics, 1933 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 44-46

Pusia schafferi ( Meznerics, 1933) nov. comb.

Figs 4K , 15E 1 –E 2 , F 1 –F 2 , G 1 –G 2 , H

[ Mitra ] elegans [Partsch]— Hörnes 1848: 17 [nomen nudum] [non Lea, 1841; non Reeve, 1845].

Mitra recticosta Bell — Hörnes 1852b: 106, pl. 10, figs 31a–c [non Bellardi, 1850].

[ Mitra ] recticosta Bell. — Auinger 1871: 8 [non Bellardi, 1850].

[ Costellaria ] recticosta Bell. —Hoernes 1880: 125 [non Bellardi, 1850].

[ Costellaria ] Borsoni Bell. —Hoernes 1880: 125 [non Bellardi, 1850].

Mitra (Costellaria) Borsoni Bell. — Hoernes & Auinger 1880: 86, pl. 10, fig. 9 [non Bellardi, 1850].

Mitra (Costellaria) borsoni Bell. — Boettger 1906: 10 [non Bellardi, 1850].

* Turricula (Uromitra) schafferi n. sp. — Meznerics 1933: 343, pl. 14, figs 6a–b.

V [exillum]. (U [romitra].) schafferi (Mezn.) — Sieber 1958a: 153.

Vexillum (Uromitra) schafferi (Mezn.) — Sieber 1958b: 149.

Vexillum (Uromitra) schafferi Meznerics , sp. 1933— Glibert 1960: 33.

Vexillum (V.) cimbricum Brocc. —Stancu & Andrescu 1968: 464, pl. 6, fig. 73 [non Brocchi, 1814].

non Vexillum (Uromitra) schafferi ( Meznerics 1933) — Steininger 1973: 432.

Type material. Holotype: NHMW 1887 /0018/0018, illustrated in Meznerics (1933, pl. 14, figs 6a–b). The Meznerics (1933) collection is stored in the NHMW but the specimen could not be found.

Additional topotypic material. NHMW 2020 /0078/0001, SL: 10.9 mm, MD: 3.8 mm, Steinebrunn ( Austria) , 15G 1 –G 2; NHMW 2020 /0078/0002, SL: 9.8 mm, MD: 3.5 mm, Steinebrunn ( Austria) , 15E 1 – E 2, 4 K; NHMW 2020 /0078/0003, SL: 11.8 mm, MD: 4.0 mm, Steinebrunn ( Austria) , 15F 1 –F 2; NHMW 2020 /0079/0001, SL: 11.0 mm, MD: 3.6 mm, Steinebrunn ( Austria) , 15H; NHMW 1884 /0037/2895, 61, specimens, Steinebrunn ( Austria) ; NHMW 1860 /0001/0113, Steinebrunn ( Austria) , illustrated in Hoernes & Auinger (1880 pl. 10, fig. 9); NHMW 1846 /0037/0111, Steinebrunn ( Austria) , illustrated in Hörnes (1852b, pl. 10, figs 31a–c)

Revised description. Shell moderately large, moderately slender fusiform, with high spire. Protoconch (fig. 4K) high conical of 3.5 smooth, weakly convex whorls (height: 900 μm, diameter: 720 μm). Teleoconch of six to seven weakly convex to subcylindrical whorls, with periphery close above abapical suture; suture deeply incised. Sculpture comprising prominent, weakly opisthocyrt axial ribs, separated by interspaces of roughly equal width. Axial ribs often slightly swollen at adapical tip, resulting in weakly gradate spire. Delicate spiral cords restricted to interspaces between axial ribs. Axial ribs slightly broader on last whorl. Last whorl slightly inflated, evenly convex, strongly constricted at base. Aperture wide, ovoid, with indistinct anal sinus. Columellar callus narrow, sharply delimited. Columella with three prominent folds, weakening abapically; very weak fourth fold in some specimens. Outer lip thin, bearing about ten delicate lirae within. About four weak spiral cords on base. Siphonal canal short, bent slightly to left, with shallow siphonal notch.

Shell measurements and ratios. SL: 8.4–12.9 mm, MD: 3.3–4.9 mm; AA: 33–35°, SL/MD: 2.7–3.0, AL/AW: 2.2–3.5, AH/S: 3.0–4.5.

Discussion. This species was originally identified by Hörnes (1852b) as Mitra recticosta Bellardi, 1850 , which lacks spiral sculpture. Consequently, Hoernes & Auinger (1880) identified it as Uromitra borsoni Bellardi, 1850 , based on the rather schematic drawings in Bellardi (1850). Meznerics (1933) doubted this identification and described the specimens from Steinebrunn as new species. This view is supported by the illustration of a syntype of Vexillum borsoni ( Bellardi, 1850) from the Tortonian S. Agata ( Italy) (Ferrero-Mortara et al., 1981, pl. 52, figs 14a–b). It differs from the Austrian species in its sigmoidal axial ribs, the strongly constricted base and the longer siphonal canal.

Pusia schafferi was also mentioned from the Ottnangian (Burdigalian) of Bavaria (Kaltenbachgraben) by Steininger (1973) based on a spire fragment. The specimen has never been illustrated and the identification remains unclear.

Pusia schafferi ( Meznerics, 1933) has a similarly elongated fusiform shape as P. pseudorecticosta ( Boettger, 1906) , but differs in having fine spiral grooves visible in the axial interspaces, whereas in P. pseudorecticosta only the strong spiral cords over the siphonal fasciole are present.

Palaeoenvironment. Steinebrunn represents a silty-sandy inner neritic environment with sea grass meadows (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): North-Alpine-Carpathian Foredeep: Hrušovany nad Jevišovkou, Jerutek at Lysice, Drnovice u Vyškova ( Czech Republic) ( Hoernes & Auinger 1880); Vienna Basin: Gainfarn, Niederleis, Steinebrunn, Vienna /Pötzleinsdorf ( Austria), Sedlec (= Porzteich), Mikulov ( Czech Republic), ( Hoernes & Auinger 1880; Sieber 1958b), Rohožník–Konopiská ( Slovakia) ( Biskupič 2020); Eisenstadt-Sopron Basin: Forchtenau ( Austria) ( Sieber 1956); Danube Basin: Hlohovec (= Bischofswart) (Slovakia) ( Hoernes & Auinger 1880).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes & Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes & Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037.

FIGURE 15 A1–A2. Pusia pseudorecticosta (Boettger, 1906), lectotype, Steinebrunn (Austria), NHMW 2020/0076/0002. B1–B2. Pusia pseudorecticosta (Boettger, 1906), paralectotype, Steinebrunn (Austria), NHMW 2020/0076/0001. C1–C2. Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0004. D. Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0003. E1–E2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. F1–F2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0003. G1–G2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0001. H. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0079/0001. I1–I2. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. J1–J2. Pusia vexans (Boettger, 1902), lectotype, Coşteiu de Sus (Romania), SMF 360339. K1–K2. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2018/0248/0327.